196 lines
20 KiB
XML
196 lines
20 KiB
XML
<document ID-DOI="10.11646/zootaxa.5231.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7575308" ID-ZooBank="EBD53535-397E-4894-B0A4-9AC3CC38DF3C" checkinTime="1674811458118" checkinUser="plazi" docAuthor="Lelej, Arkady S. & Williams, Kevin A." docDate="2023" docId="03CF87A93124B506FF17FF00F67B379F" docLanguage="en" docName="zootaxa.5231.2.1.pdf" docOrigin="Zootaxa 5231 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocephalotilla Bischoff 1920" docType="treatment" docVersion="3" lastPageNumber="125" masterDocId="FFF6FFD13133B51EFF80FF97F345374A" masterDocTitle="Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)" masterLastPageNumber="140" masterPageNumber="101" pageNumber="124" updateTime="1674840100621" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lelej, Arkady S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-7501-0981</mods:nameIdentifier>
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<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
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<mods:nameIdentifier type="email">lelej@biosoil.ru</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Williams, Kevin A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
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<mods:affiliation>Plant Pest Diagnostic Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
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<mods:nameIdentifier type="email">illiams@cdfa.ca.gov</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5231</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5231.2.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5231.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7575308</mods:identifier>
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<subSubSection box="[151,609,151,178]" pageId="23" pageNumber="124" type="nomenclature">
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<paragraph blockId="23.[151,609,151,213]" box="[151,609,151,178]" pageId="23" pageNumber="124">
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<heading bold="true" box="[151,609,151,178]" fontSize="11" level="1" pageId="23" pageNumber="124" reason="1">
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<emphasis bold="true" box="[151,609,151,178]" pageId="23" pageNumber="124">
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20.
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<taxonomicName authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[195,609,151,178]" class="Insecta" family="Mutillidae" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[195,425,151,178]" italics="true" pageId="23" pageNumber="124">Pseudocephalotilla</emphasis>
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<bibRefCitation author="Bischoff, H. & Monographie der Mutilliden Afrikas" box="[432,609,151,177]" pageId="23" pageNumber="124" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) & 86 A (4 - 5), 1 - 480 (1920) & 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,295,187,213]" pageId="23" pageNumber="124" type="description">
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<paragraph blockId="23.[151,609,151,213]" box="[151,295,187,213]" pageId="23" pageNumber="124">
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(
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<figureCitation box="[159,288,187,213]" captionStart="FIGURES 61–66" captionStartId="23.[152,268,1938,1962]" captionTargetBox="[160,1427,838,1914]" captionTargetId="figure-241@23.[160,1427,838,1915]" captionTargetPageId="23" captionText="FIGURES 61–66. 61–62. Psammotherma flabellata (Fabricius, 1804), ♁: 61. Lateral habitus, 62. Dorsal habitus; 63–65. Pseudocephalotillabeira (Péringuey,1914):63.♀,dorsalhabitus,64.♀,lateralhabitus,65.♁, lateralhabitus;66.Pseudocephalotilla tettensis (Gerstaecker in Peters, 1858), ♀, dorsal habitus. (Photographs by K. Williams, and D. Brothers (62–65))." figureDoi="http://doi.org/10.5281/zenodo.7575326" httpUri="https://zenodo.org/record/7575326/files/figure.png" pageId="23" pageNumber="124">Figs 63–66</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="124" type="reference_group">
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<paragraph blockId="23.[151,1436,255,313]" pageId="23" pageNumber="124">
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<treatmentCitationGroup pageId="23" pageNumber="124">
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<treatmentCitation author="Bischoff, H. & Monographie der Mutilliden Afrikas" box="[151,539,256,280]" page="24" pageId="23" pageNumber="124" year="1920">
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<taxonomicName authority="Bischoff, 1920: 24" authorityName="Bischoff" authorityPageNumber="24" authorityYear="1920" box="[151,539,256,280]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,347,256,280]" italics="true" pageId="23" pageNumber="124">Pseudocephalotilla</emphasis>
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<bibRefCitation author="Bischoff, H. & Monographie der Mutilliden Afrikas" box="[352,539,256,280]" pageId="23" pageNumber="124" refId="ref25275" refString="Bischoff, H. (1920 - 1921) Monographie der Mutilliden Afrikas. Archiv f ¸ r Naturgeschichte, 86 A (1 - 3) & 86 A (4 - 5), 1 - 480 (1920) & 481 - 830 (1921)." type="journal volume" year="1920">Bischoff, 1920: 24</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(key), 1921: 514 (description), ♁;
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<treatmentCitation author="Nonveiller, G." box="[889,1093,255,280]" page="15" pageId="23" pageNumber="124" year="1979">
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<bibRefCitation author="Nonveiller, G." box="[889,1093,255,280]" pageId="23" pageNumber="124" pagination="1 - 184" refId="ref27953" refString="Nonveiller, G. (1979 [" 1978 "]) Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Beograd, 13, 1 - 184." type="journal article" year="1979">Nonveiller 1979: 15</bibRefCitation>
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</treatmentCitation>
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, ♁,
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<specimenCount box="[1134,1154,256,279]" pageId="23" pageNumber="124" type="female">♀</specimenCount>
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;
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<treatmentCitation author="Lelej, A. S. & Brothers, D. J." box="[1165,1432,256,280]" page="49" pageId="23" pageNumber="124" year="2008">
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<bibRefCitation author="Lelej, A. S. & Brothers, D. J." box="[1165,1432,256,280]" pageId="23" pageNumber="124" pagination="1 - 79" refId="ref27054" refString="Lelej, A. S. & Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej & Brothers 2008: 49</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Brothers, D. J. & Lelej, A. S." box="[199,469,286,310]" page="96" pageId="23" pageNumber="124" year="2017">
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<bibRefCitation author="Brothers, D. J. & Lelej, A. S." box="[199,469,286,310]" pageId="23" pageNumber="124" pagination="1 - 97" refId="ref25501" refString="Brothers, D. J. & Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers & Lelej 2017: 96</bibRefCitation>
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</treatmentCitation>
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, ♁,
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<specimenCount box="[511,531,287,310]" pageId="23" pageNumber="124" type="female">♀</specimenCount>
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;
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<treatmentCitation author="Pagliano, G. & Brothers, D. J. & Cambra, R. & Lelej, A. S. & Lo Cascio, P. & Matteini Palmerini, M. & Scaramozzino, P. L. & Williams, K. A. & Romano, M." box="[543,800,286,310]" page="185" pageId="23" pageNumber="124" year="2020">
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<bibRefCitation author="Pagliano, G. & Brothers, D. J. & Cambra, R. & Lelej, A. S. & Lo Cascio, P. & Matteini Palmerini, M. & Scaramozzino, P. L. & Williams, K. A. & Romano, M." box="[543,800,286,310]" pageId="23" pageNumber="124" pagination="5 - 425" refId="ref28648" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. & Romano, M. (2020 [" 2018 "]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torin o, 36 (1 - 2), 5 - 425." type="journal article" year="2020">
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Pagliano
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<emphasis box="[638,690,286,310]" italics="true" pageId="23" pageNumber="124">et al.</emphasis>
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2020: 185
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||
</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,1401,356,385]" pageId="23" pageNumber="124" type="type_taxon">
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<paragraph blockId="23.[151,1437,356,815]" box="[151,1401,356,385]" pageId="23" pageNumber="124">
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<emphasis bold="true" box="[151,306,356,383]" pageId="23" pageNumber="124">
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||
<typeStatus box="[151,210,357,383]" pageId="23" pageNumber="124">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName authority="Bischoff, 1921" authorityName="Bischoff" authorityYear="1921" box="[313,798,356,383]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="beirana">
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<emphasis box="[313,623,356,382]" italics="true" pageId="23" pageNumber="124">Pseudocephalotilla beirana</emphasis>
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Bischoff, 1921
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</taxonomicName>
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||
, ♁, by subsequent designation of Bischoff 1921: 514.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="23" pageNumber="124" type="diagnosis">
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||
<paragraph blockId="23.[151,1437,356,815]" pageId="23" pageNumber="124">
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||
<emphasis bold="true" box="[199,314,392,418]" pageId="23" pageNumber="124">Diagnosis</emphasis>
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||
. MALE. Flagellomeres not overlapped. Mandible not widened apically, beneath with large subbasal tooth, first preapical inner tooth distinctly smaller, second preapical tooth larger than apical one. Pronotum anteriorly pulled forward with weak carina. T2 with long lateral felt lines, S2 with short lateral felt lines. Volsella without basal external lobe, with wide acuminated cuspis; basivolsella without short setae. Penis valve shortened, swollen apex curved down and oriented outside. FEMALE. Head not widened posterad, posteriorly rouded. Minimal distance between eyes 1.6–1.8 × longitudinal eye diameter. Pronotum without protruding humeral part, at most scarcely wider than propodeum. Scutellar scale more or less developed. T2 with one or three spots of pale setae disposed in central horizontal line. T1 posterodorsally with small medial spot of pale setae. Pygidial plate elongated, widened basally, longitudinally striated.
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</paragraph>
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<paragraph blockId="23.[151,1437,356,815]" pageId="23" pageNumber="124">
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<emphasis bold="true" box="[206,512,716,743]" pageId="23" pageNumber="124">Diversity and distribution</emphasis>
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. Four species [two of them:
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<taxonomicName authority="(Smith, 1855)" baseAuthorityName="Smith" baseAuthorityYear="1855" box="[830,1102,716,743]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="atropos">
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<emphasis box="[830,941,717,742]" italics="true" pageId="23" pageNumber="124">P. atropos</emphasis>
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||
(
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<bibRefCitation author="Smith, F." box="[955,1094,716,743]" pageId="23" pageNumber="124" pagination="1 - 5" refId="ref29668" refString="Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Trustees of the British Museum, London, [4] + 206 pp., pl. A, pls. 1 - 5." type="book chapter" year="1855">Smith, 1855</bibRefCitation>
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)
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</taxonomicName>
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with eleven subspecies and
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<taxonomicName class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="tettensis">
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<emphasis italics="true" pageId="23" pageNumber="124">P. tettensis</emphasis>
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</taxonomicName>
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(Gerstaecker in
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<bibRefCitation author="Peters, W. C. H." box="[425,564,752,779]" pageId="23" pageNumber="124" pagination="509 - 513" refId="ref29231" refString="Peters, W. C. H. (1858 [" 1857 "]) Ubersicht der von ihm in Mossambique aufgefundenen und von Hrn. Dr. Gerstacker bearbeiteten Hymenopteren aus der Familien der Crabronites, Sphegidae, Pompilidae und Heterogyna. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1857, 509 - 513." type="journal article" year="1858">Peters, 1858</bibRefCitation>
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) with eight subspecies] are recognized from the Afrotropical
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<collectingRegion box="[1243,1323,752,778]" country="Iraq" pageId="23" pageNumber="124">Region</collectingRegion>
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||
(
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<bibRefCitation author="Pagliano, G. & Brothers, D. J. & Cambra, R. & Lelej, A. S. & Lo Cascio, P. & Matteini Palmerini, M. & Scaramozzino, P. L. & Williams, K. A. & Romano, M." pageId="23" pageNumber="124" pagination="5 - 425" refId="ref28648" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. & Romano, M. (2020 [" 2018 "]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torin o, 36 (1 - 2), 5 - 425." type="journal article" year="2020">
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||
Pagliano
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||
<emphasis box="[151,207,788,814]" italics="true" pageId="23" pageNumber="124">et al.</emphasis>
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||
2020
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</bibRefCitation>
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||
).
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||
</paragraph>
|
||
</subSubSection>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7575326" ID-Zenodo-Dep="7575326" httpUri="https://zenodo.org/record/7575326/files/figure.png" pageId="23" pageNumber="124" startId="23.[152,268,1938,1962]" targetBox="[160,1427,838,1914]" targetPageId="23">
|
||
<paragraph blockId="23.[151,1437,1938,2035]" pageId="23" pageNumber="124">
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||
<emphasis bold="true" box="[152,348,1938,1963]" pageId="23" pageNumber="124">FIGURES 61–66.</emphasis>
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61–62.
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<taxonomicName authority="(Fabricius, 1804)" baseAuthorityName="Fabricius" baseAuthorityYear="1804" box="[439,891,1939,1963]" class="Insecta" family="Mutillidae" genus="Psammotherma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="flabellata">
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<emphasis box="[439,701,1939,1963]" italics="true" pageId="23" pageNumber="124">Psammotherma flabellata</emphasis>
|
||
(
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||
<bibRefCitation author="Fabricius, J. C." box="[720,884,1939,1963]" pageId="23" pageNumber="124" pagination="1 - 30" refId="ref25818" refString="Fabricius, J. C. (1804) s. n. In: Systema Piezatorum secundum Ordines, Genera, Species adjectis Synonymis, Locis, Observationibus, Descriptionibus. Carolum Reichard, Brunsvigae [Braunschweig], pp. i - xiv + 15 - 440 + 1 - 30. https: // doi. org / 10.5962 / bhl. title. 10490" type="book chapter" year="1804">Fabricius, 1804</bibRefCitation>
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)
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</taxonomicName>
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||
, ♁: 61. Lateral habitus, 62. Dorsal habitus; 63–65.
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<taxonomicName authority="(Peringuey, 1914)" baseAuthorityName="Peringuey" baseAuthorityYear="1914" box="[151,579,1975,1999]" class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="beira">
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||
<emphasis box="[151,398,1975,1999]" italics="true" pageId="23" pageNumber="124">Pseudocephalotilla beira</emphasis>
|
||
(
|
||
<bibRefCitation author="Peringuey, L." box="[408,572,1975,1999]" pageId="23" pageNumber="124" pagination="323 - 358" refId="ref29190" refString="Peringuey, L. (1914) Notes on South African Mutillidae (Hymenoptera), with descriptions of new or little known species. Annals of the South African Museum, 10 (10), 323 - 358." type="journal article" year="1914">Péringuey, 1914</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
:63.♀, dorsal habitus, 64.♀, lateral habitus,65. ♁, lateral habitus;66.
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<taxonomicName class="Insecta" genus="Pseudocephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="tettensis">
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<emphasis italics="true" pageId="23" pageNumber="124">Pseudocephalotilla tettensis</emphasis>
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</taxonomicName>
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(Gerstaecker in
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<bibRefCitation author="Peters, W. C. H." box="[397,523,2011,2035]" pageId="23" pageNumber="124" pagination="509 - 513" refId="ref29231" refString="Peters, W. C. H. (1858 [" 1857 "]) Ubersicht der von ihm in Mossambique aufgefundenen und von Hrn. Dr. Gerstacker bearbeiteten Hymenopteren aus der Familien der Crabronites, Sphegidae, Pompilidae und Heterogyna. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1857, 509 - 513." type="journal article" year="1858">Peters, 1858</bibRefCitation>
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), ♀, dorsal habitus. (Photographs by K. Williams, and D. Brothers (62–65)).
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</paragraph>
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||
</caption>
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||
<subSubSection pageId="24" pageNumber="125" type="discussion">
|
||
<paragraph blockId="24.[151,1436,151,213]" pageId="24" pageNumber="125">
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||
<emphasis bold="true" box="[199,308,151,177]" pageId="24" pageNumber="125">Remarks</emphasis>
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. Based on a recent molecular phylogenetic analysis (Waldren
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<emphasis box="[1007,1064,151,177]" italics="true" pageId="24" pageNumber="125">et al.</emphasis>
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in press), this genus belongs in a new tribe. Most of the species known in this genus are included in Bischoff’s (1921) key to
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<taxonomicName authorityName="Thomson" authorityYear="1870" box="[1180,1334,187,213]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="125" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1180,1334,187,213]" italics="true" pageId="24" pageNumber="125">Smicromyrme</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |