408 lines
72 KiB
XML
408 lines
72 KiB
XML
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<mods:title id="64251A113E56D8BFD78EE142C682EAF0">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="BCD778025860568C9DD1F7EB3AB8A2D7">Suksai, Benjamart</mods:namePart>
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<mods:nameIdentifier id="233CB79B924800D53C5A7A4268CE2751" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:namePart id="105035AB793A61E414E4584D57974F17">Gustafson, Grey T.</mods:namePart>
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<mods:affiliation id="0EA61DF215837485BDBB69CA1647B050">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. & Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="1A3CA7D0123AD7B17EDC7AE277011EA5">Sites, Robert W.</mods:namePart>
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<mods:nameIdentifier id="71CD42A338507AB3F89F418B969A7205" type="ORCID">0000-0002-3895-813X</mods:nameIdentifier>
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<mods:affiliation id="692270FB8D4AE234F6D9A049DF9C0419">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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<mods:namePart id="B0D2AA48CCBCBD50D1684FADEC683CFF">Sangpradub, Narumon</mods:namePart>
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<mods:affiliation id="C48D2082FF67BE92E32BD3100926B913">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand & Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
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<mods:nameIdentifier id="7E34B2FF2D04DFD9F6E68DF0A5820507" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:title id="AE922034DB1064BFCC9EF9BF9C3926E6">Zootaxa</mods:title>
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<mods:date id="2236E341B5F0E36F6B8718082E78D37E">2021</mods:date>
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<mods:number id="DC31D9C401E64B3176CCECA0812E9F41">2021-06-25</mods:number>
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<taxonomicName id="4C154D54FFFA225B0CC07921FE815066" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[151,350,769,796]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
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<emphasis id="B961EAC5FFFA225B0CC07921FE815066" bold="true" box="[151,350,769,796]" italics="true" pageId="16" pageNumber="577">Patrus nanensis</emphasis>
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(
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,
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,
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<figureCitation id="132E2A52FFFA225B0D3C7906FEA45042" box="[363,379,805,831]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">9</figureCitation>
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)
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<elevation id="0038D1E4FFFA225B0F0979DEFC2D576A" box="[862,1010,1021,1048]" metricMagnitude="3" metricUnit="m" metricValue="1.474" pageId="16" pageNumber="577" unit="m" value="1474.0">
|
||
<quantity id="4CED9B32FFFA225B0F0979DEFC6B5765" box="[862,948,1021,1048]" metricMagnitude="3" metricUnit="m" metricValue="1.474" pageId="16" pageNumber="577" unit="m" value="1474.0">1474 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFAB1017FFFA225B0FA979DEFBA9576A" box="[1022,1142,1021,1047]" pageId="16" pageNumber="577" value="2017-05-30">
|
||
<collectingDate id="EFEFE9FFFFFA225B0FA979DEFBA9576A" box="[1022,1142,1021,1047]" pageId="16" pageNumber="577" value="2017-05-30">30.V.2017</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26E05301FFFA225B08E579DDFA99576A" box="[1202,1350,1021,1048]" pageId="16" pageNumber="577">P. Sapewisut</collectorName>
|
||
,
|
||
<specimenCount id="9D13FD5EFFFA225B090779DEFA43576A" box="[1360,1436,1021,1047]" pageId="16" pageNumber="577" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED04AE12FFFA225B0CC87E02FF3D5746" box="[159,226,1057,1083]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="16" pageNumber="577">KKU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B7D3C8AFFFA225B0CAE7E02FD9B5746" box="[249,580,1057,1083]" collectingDate="2017-05-30" collectionCode="THNHM, UMC" collectorName="P. Sapewisut" country="Thailand" county="Ban Khun Sathan" elevation="1474" isTailEnumeration="true" latitude="18.273367" location="Na Noi District" longLatPrecision="1" longitude="100.48633" municipality="San Ta Subdistrict" pageId="16" pageNumber="577" specimenCount="2" specimenCount-female="2" stateProvince="Nan" typeStatus="paratype">
|
||
<specimenCount id="9D13FD5EFFFA225B0CAE7E02FEB55746" box="[249,362,1057,1083]" pageId="16" pageNumber="577" type="female">2 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED04AE12FFFA225B0D2B7E02FE365746" box="[380,489,1057,1083]" pageId="16" pageNumber="577">THNHM</collectionCode>
|
||
,
|
||
<collectionCode id="ED04AE12FFFA225B0DA17E02FDE35746" box="[502,572,1057,1083]" pageId="16" pageNumber="577">UMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7D3C8AFFFA225B0E047E01FAC05722" collectingDate="2017-05-30" collectionCode="UMC" collectorName="P. Sapewisut" country="Thailand" county="Ban Khun Sathan" elevation="1356" isTailEnumeration="true" latitude="18.278349" location="Na Noi District" longLatPrecision="1" longitude="100.507484" municipality="San Ta Subdistrict" pageId="16" pageNumber="577" specimenCount="1" specimenCount-male="1" stateProvince="Nan" typeStatus="paratype">
|
||
same locality except for, water supply weir,
|
||
<collectingMunicipality id="6BCEACADFFFA225B08087E02FAE65746" box="[1119,1337,1057,1084]" pageId="16" pageNumber="577">San Ta Subdistrict</collectingMunicipality>
|
||
,
|
||
<location id="8ECA600CFFFA225B09107E01FF2F5722" LSID="urn:lsid:plazi:treatment:03BC87C1FFFA225A0CC07921FDAF54A0:8ECA600CFFFA225B09107E01FF2F5722" country="Thailand" latitude="18.278349" longLatPrecision="1" longitude="100.507484" municipality="San Ta Subdistrict" name="Na Noi District" pageId="16" pageNumber="577" stateProvince="Nan">Na Noi District</location>
|
||
,
|
||
<geoCoordinate id="EE215010FFFA225B0CAB7E66FE735722" box="[252,428,1093,1119]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="16" pageNumber="577" precision="1" seconds="42.06" value="18.278349">18°16’42.06”N</geoCoordinate>
|
||
<geoCoordinate id="EE215010FFFA225B0DE47E66FDAE571D" box="[435,625,1093,1120]" degrees="100" direction="east" minutes="30" orientation="longitude" pageId="16" pageNumber="577" precision="1" seconds="26.95" value="100.507484">100°30’26.95”E</geoCoordinate>
|
||
,
|
||
<elevation id="0038D1E4FFFA225B0E2A7E66FCCD5722" box="[637,786,1093,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.3559999999999999" pageId="16" pageNumber="577" unit="m" value="1356.0">
|
||
<quantity id="4CED9B32FFFA225B0E2A7E66FD0C571D" box="[637,723,1093,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.3559999999999999" pageId="16" pageNumber="577" unit="m" value="1356.0">1356 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFAB1017FFFA225B0F487E66FC495722" box="[799,918,1093,1119]" pageId="16" pageNumber="577" value="2017-05-30">
|
||
<collectingDate id="EFEFE9FFFFFA225B0F487E66FC495722" box="[799,918,1093,1119]" pageId="16" pageNumber="577" value="2017-05-30">30.V.2017</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26E05301FFFA225B0F837E65FBB75722" box="[980,1128,1093,1120]" pageId="16" pageNumber="577">P. Sapewisut</collectorName>
|
||
,
|
||
<specimenCount id="9D13FD5EFFFA225B08247E66FB1E5722" box="[1139,1217,1093,1119]" pageId="16" pageNumber="577" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED04AE12FFFA225B08877E66FAC85722" box="[1232,1303,1093,1119]" pageId="16" pageNumber="577">UMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFFA225B0C907E4AFD2E5696" pageId="16" pageNumber="577" type="diagnosis">
|
||
<paragraph id="8BAA36D7FFFA225B0C907E4AFD2E5696" blockId="16.[151,1437,877,2020]" pageId="16" pageNumber="577">
|
||
<emphasis id="B961EAC5FFFA225B0C907E4AFE9B57FE" bold="true" box="[199,324,1129,1155]" pageId="16" pageNumber="577">Diagnosis.</emphasis>
|
||
Body form in lateral view weakly dorsoventrally convex; elytral glabrous region of male (
|
||
<figureCitation id="132E2A52FFFA225B09607E4AFA4857FE" box="[1335,1431,1129,1155]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="16" pageNumber="577">Figs. 2E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFA225B0CC07EAEFF6457DA" box="[151,187,1165,1191]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">9A</figureCitation>
|
||
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to considerably wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; female elytral glabrous region (
|
||
<figureCitation id="132E2A52FFFA225B0D097EF6FE1D5792" box="[350,450,1237,1263]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="16" pageNumber="577">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFA225B0D877EF6FE2E5792" box="[464,497,1237,1263]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">9E</figureCitation>
|
||
) dissimilar to male, broad in anterior 4/5 occupying 9/11 elytral width, abruptly acuminate medially in posterior 1/3, posterolateral corners produced into acute points; elytral apex of both sexes similar, obliquely truncate without apicolateral sinuation, epipleural angle distinct, weakly rounded; male protibia (
|
||
<figureCitation id="132E2A52FFFA225B0CF77F62FF265626" box="[160,249,1345,1371]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="16" pageNumber="577">Fig. 4E</figureCitation>
|
||
) wedge-shaped, protibial apex truncate without prominent distolateral angle; male protarsus (
|
||
<figureCitation id="132E2A52FFFA225B096B7F62FA4B5626" box="[1340,1428,1345,1371]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="16" pageNumber="577">Fig. 4E</figureCitation>
|
||
) ca. 1/2 length of protibia, protarsomeres width gradually decreasing apically; mesoventrite without medial pit; aedeagus (
|
||
<figureCitation id="132E2A52FFFA225B0D447FAAFE4E56DE" box="[275,401,1417,1443]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">Fig. 9B–D</figureCitation>
|
||
) with median lobe tapered, apex strongly constricted to a small needle-like point; apex not constricted in lateral view (
|
||
<figureCitation id="132E2A52FFFA225B0DA97F8EFD8656BA" box="[510,601,1453,1479]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">Fig. 9C</figureCitation>
|
||
); female reproductive tract (
|
||
<figureCitation id="132E2A52FFFA225B0FC87F8EFC2656BA" box="[927,1017,1453,1479]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">Fig. 9G</figureCitation>
|
||
) without additional sclerite between gonocoxae (
|
||
<figureCitation id="132E2A52FFFA225B0D747FF2FEA65696" box="[291,377,1489,1515]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">Fig. 9F</figureCitation>
|
||
), fertilization duct as in
|
||
<figureCitation id="132E2A52FFFA225B0EC57FF2FD335696" box="[658,748,1489,1515]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="16" pageNumber="577">Fig. 9H</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFFA225A0C907FD6FCD05628" lastPageId="17" lastPageNumber="578" pageId="16" pageNumber="577" type="description">
|
||
<paragraph id="8BAA36D7FFFA225B0C907FD6FEE05549" blockId="16.[151,1437,877,2020]" pageId="16" pageNumber="577">
|
||
<emphasis id="B961EAC5FFFA225B0C907FD6FE835572" bold="true" box="[199,348,1525,1551]" pageId="16" pageNumber="577">Description.</emphasis>
|
||
<emphasis id="B961EAC5FFFA225B0D357FD6FE635572" box="[354,444,1525,1551]" italics="true" pageId="16" pageNumber="577">Habitus</emphasis>
|
||
. Size. Male: TL =
|
||
<quantity id="4CED9B32FFFA225B0EC57FD6FCE5556D" box="[658,826,1525,1552]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.0699999999999998" metricValueMin="1.04" pageId="16" pageNumber="577" unit="mm" value="10.55" valueMax="10.7" valueMin="10.4">10.4–10.7 mm</quantity>
|
||
, TW = 6.0–
|
||
<quantity id="4CED9B32FFFA225B0FE87FD6FBC6556D" box="[959,1049,1525,1552]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="16" pageNumber="577" unit="mm" value="6.2">6.2 mm</quantity>
|
||
; female: TL =
|
||
<quantity id="4CED9B32FFFA225B08E97FD6FAB8556D" box="[1214,1383,1525,1552]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.0699999999999998" metricValueMin="1.06" pageId="16" pageNumber="577" unit="mm" value="10.649999999999999" valueMax="10.7" valueMin="10.6">10.6–10.7 mm</quantity>
|
||
, TW =
|
||
<quantity id="4CED9B32FFFA225B0CF87C3AFEE35549" box="[175,316,1561,1588]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.7" pageId="16" pageNumber="577" unit="mm" value="5.75" valueMax="5.8" valueMin="5.7">5.7–5.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFA225B0C907C1EFD5255E2" blockId="16.[151,1437,877,2020]" pageId="16" pageNumber="577">Body form elongate oval, evenly attenuated anteriorly and posteriorly, maximum width just posterior of humeral angle; in lateral view dorsoventrally weakly convex, maximum convexity posterior to scutellar region, evenly depressed anteriorly and posteriorly.</paragraph>
|
||
<paragraph id="8BAA36D7FFFA225B0C907C8AFCD15452" blockId="16.[151,1437,877,2020]" pageId="16" pageNumber="577">
|
||
<emphasis id="B961EAC5FFFA225B0C907C8AFE9B55BE" box="[199,324,1705,1731]" italics="true" pageId="16" pageNumber="577">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="132E2A52FFFA225B0D027C8AFE6755BE" box="[341,440,1705,1731]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="16" pageNumber="577">Figs. 2E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFA225B0D927C8AFE3755BE" box="[453,488,1705,1731]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="16" pageNumber="577">3E</figureCitation>
|
||
). Dorsally head, pronotum, and elytra margined with golden pubescence; glabrous regions of head pronotum and elytra black with reticulation creating brassy sheen. Pronotal and elytral lateral margins dark. Venter dark, reddish brown to black. Middle and hind legs from trochanter to tarsus and ultimate abdominal ventrite lighter in color, red to orangish red.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFA225A0C907D1AFE955384" blockId="16.[151,1437,877,2020]" lastBlockId="17.[151,1437,151,2013]" lastPageId="17" lastPageNumber="578" pageId="16" pageNumber="577">
|
||
<emphasis id="B961EAC5FFFA225B0C907D1AFEDB542E" box="[199,260,1849,1875]" italics="true" pageId="16" pageNumber="577">Head</emphasis>
|
||
. Head capsule broad, convex. Labrum short, ca. 3.5× wider than long; setose medially with setae situated in large deep punctures; apex broadly rounded. Clypeus narrow, ca. 1/2 width of labrum; strongly reticulate and densely punctured, reticulation hatched in appearance with numerous short, well-impressed transverse lines. Frons similarly with strong hatched reticulation and dense punctation; pseudofrontal ridge present; furrow beneath pseudofrontal ridge without setae. Vertex also with strongly impressed hatch reticulation and dense punctation between dorsal eyes, reticulation becoming effaced in narrow band on occipital region. Dorsal eye delimited by strong orbital furrow; distance between dorsal eyes ca. 2.75× diameter of an eye; interorbital area strongly setose.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFB225A0C907B20FC5A5068" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C907B20FEFE5260" box="[199,289,259,285]" italics="true" pageId="17" pageNumber="578">Thorax.</emphasis>
|
||
Pronotum (
|
||
<figureCitation id="132E2A52FFFB225A0DF57B20FDDD5260" box="[418,514,259,285]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="17" pageNumber="578">Figs.2E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFB225A0E5C7B20FDF05260" box="[523,559,259,285]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">9A</figureCitation>
|
||
) with narrow lateral margins; pronotal transverse impressed line absent; lateral pubescence anteriorly extending onto pronotal disc as far as 2/3 width of dorsal eye; pronotal glabrous region densely and uniformly punctate with small weakly impressed punctures, strongly reticulate with uniform wellimpressed hatched reticulation similar to frons and vertex. Scutellar shield visible. Protrochanter with setose line along ventral (anterior) face. Profemur with setigerous punctures along anterior margin of ventral face, posterior margin of ventral face with row of setae along basal 2/3 ending with patch of longer denser setae. Protibia (
|
||
<figureCitation id="132E2A52FFFB225A09397B94FF665288" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="17" pageNumber="578">Fig. 4E</figureCitation>
|
||
) wedge shaped in males, not constricted basally, ventral (inner) margin nearly straight; distal apex truncate, distolateral angle without prominence. Male protarsus (
|
||
<figureCitation id="132E2A52FFFB225A0F487BDCFCA65164" box="[799,889,511,537]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="17" pageNumber="578">Fig. 4E</figureCitation>
|
||
) ca. 1/2 length of protibia, protarsomeres with distolateral angle round, not overlapping following tarsomere, lateral margins weakly rounded, fifth tarsomere 2× length of fourth, with lateral margins weakly rounded. Elytra (
|
||
<figureCitation id="132E2A52FFFB225A0FFE7864FBD0511C" box="[937,1039,583,609]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="17" pageNumber="578">Figs. 2E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFB225A08487864FB9D511C" box="[1055,1090,583,609]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">9A</figureCitation>
|
||
) with narrow lateral margins gradually widened apically; glabrous region cordiform, elytral glabrous region along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to just wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; reticulation and punctation similar to pronotal glabrous region; elytral apex obliquely truncate, without apicolateral sinuation, epipleural angle distinct, weakly rounded, sutural angle straight forming near 90° angle. Mesoventrite without medial pit.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFB225A0C90793CFD39576C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C90793CFE7B5044" box="[199,420,799,825]" italics="true" pageId="17" pageNumber="578">Sexual dimorphism</emphasis>
|
||
. Female smaller in size with narrow maximum width. Glabrous region of female elytra different (
|
||
<figureCitation id="132E2A52FFFB225A0D5C7960FEAF5020" box="[267,368,835,861]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="17" pageNumber="578">Figs. 3E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFB225A0D297960FE7E5020" box="[382,417,835,861]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">9E</figureCitation>
|
||
), extending ca. 7/9 length of elytra, broad in anterior 4/5 occupying 9/11 elytral width, lateral margins straight creating parallel-sided appearance, abruptly acuminate medially in posterior 1/3 with posterolateral corners produced into acute points, posterior margin unevenly convex towards suture with subtruncate appearance; in lateral view width of pubescent margin at humeral region nearly equal to that extending posterior. Protibia of female narrower with more strongly prominent distolateral angle. Female protarsus unexpanded, narrow and parallel-sided in appearance.
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFB225A0C907E38FCEB57DC" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
Male genitalia. Aedeagus (
|
||
<figureCitation id="132E2A52FFFB225A0DA97E38FD875748" box="[510,600,1051,1077]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9B</figureCitation>
|
||
) with median lobe just shorter than parameres, median lobe tapered for 3/5 length, parallel-sided in apical 1/3 before acumination, acumination short, 1/5 length of median lobe, apex strongly constricted to a small needle-like point; apex not constricted in lateral view (
|
||
<figureCitation id="132E2A52FFFB225A08237E40FB115700" box="[1140,1230,1123,1149]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9C</figureCitation>
|
||
), operculum as in (
|
||
<figureCitation id="132E2A52FFFB225A0CF77EA4FF2257DC" box="[160,253,1159,1185]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9D</figureCitation>
|
||
); parameres narrowly rounded apically (
|
||
<figureCitation id="132E2A52FFFB225A0E9B7EA4FCF857DC" box="[716,807,1159,1185]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFB225A0C907E88FA805670" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
Female genitalia. Reproductive tract with large sac-like spermatheca (
|
||
<figureCitation id="132E2A52FFFB225A0FB37E88FB9F57B8" box="[996,1088,1195,1221]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9G</figureCitation>
|
||
); fertilization duct sclerotized, highly convoluted (
|
||
<figureCitation id="132E2A52FFFB225A0DD47EECFE3A5794" box="[387,485,1231,1257]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9H</figureCitation>
|
||
); small sclerite present posterior to oviduct; additional round sclerite between gonocoxae absent; gonocoxae narrow, apically narrowly rounded, lateral margins weakly curved (
|
||
<figureCitation id="132E2A52FFFB225A08AD7ED0FA8D5670" box="[1274,1362,1267,1293]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAA36D7FFFB225A0C907F35FCD05628" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C907F35FE9E564D" bold="true" box="[199,321,1302,1328]" pageId="17" pageNumber="578">Variation.</emphasis>
|
||
There is considerable variation in size and width among the specimens examined. Large males weakly broader in total body width than smaller males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFFB225A0C907F7DFDB05430" pageId="17" pageNumber="578" type="diagnosis">
|
||
<paragraph id="8BAA36D7FFFB225A0C907F7DFDB05430" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C907F7DFE145605" bold="true" box="[199,459,1374,1400]" pageId="17" pageNumber="578">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C154D54FFFB225A0D837F7CFD595605" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[468,646,1375,1400]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFFB225A0D837F7CFD595605" box="[468,646,1375,1400]" italics="true" pageId="17" pageNumber="578">Patrus nanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFFB225A0EC77F7CFD355604" bold="true" box="[656,746,1375,1401]" pageId="17" pageNumber="578">
|
||
<taxonomicNameLabel id="A25257BEFFFB225A0EC77F7CFD355604" box="[656,746,1375,1401]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="4C154D54FFFB225A0F957F7CFBE85604" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[962,1079,1375,1401]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFFB225A0F957F7CFBE85604" box="[962,1079,1375,1401]" italics="true" pageId="17" pageNumber="578">P. apicalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C154D54FFFB225A08267F7CFAD55604" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[1137,1290,1375,1401]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="subapicalis">
|
||
<emphasis id="B961EAC5FFFB225A08267F7CFAD55604" box="[1137,1290,1375,1401]" italics="true" pageId="17" pageNumber="578">P. subapicalis</emphasis>
|
||
</taxonomicName>
|
||
. Males of
|
||
<taxonomicName id="4C154D54FFFB225A09DF7F7CFF2856E1" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFFB225A09DF7F7CFF2856E1" italics="true" pageId="17" pageNumber="578">P. nanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFFB225A0CA87FA0FE8756E0" bold="true" box="[255,344,1411,1437]" pageId="17" pageNumber="578">
|
||
<taxonomicNameLabel id="A25257BEFFFB225A0CA87FA0FE8756E0" box="[255,344,1411,1437]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be readily distinguished by the shorter protarsi, which are about 1/2 the length of the protibia, and the width of the protarsomeres gradually decreases apically (
|
||
<figureCitation id="132E2A52FFFB225A0F317F84FC6356BC" box="[870,956,1447,1473]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="17" pageNumber="578">Fig. 4E</figureCitation>
|
||
), compared to the other two species, which have longer male protarsi that are about 3/4 the length of the protibia, and the width of protarsomeres 1–4 are mostly subequal throughout (
|
||
<figureCitation id="132E2A52FFFB225A0DDF7FCCFDDD5574" box="[392,514,1519,1545]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="17" pageNumber="578">Fig. 4A, H</figureCitation>
|
||
). The aedeagus of
|
||
<taxonomicName id="4C154D54FFFB225A0E857FCCFC5C5575" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[722,899,1519,1544]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFFB225A0E857FCCFC5C5575" box="[722,899,1519,1544]" italics="true" pageId="17" pageNumber="578">Patrus nanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFFB225A0FDD7FCCFC3D5574" bold="true" box="[906,994,1519,1545]" pageId="17" pageNumber="578">
|
||
<taxonomicNameLabel id="A25257BEFFFB225A0FDD7FCCFC3D5574" box="[906,994,1519,1545]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132E2A52FFFB225A0FA57FCCFB995574" box="[1010,1094,1519,1545]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9B</figureCitation>
|
||
) though similar in form to that of
|
||
<taxonomicName id="4C154D54FFFB225A0CE17C30FEF55550" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[182,298,1555,1581]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFFB225A0CE17C30FEF55550" box="[182,298,1555,1581]" italics="true" pageId="17" pageNumber="578">P. apicalis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132E2A52FFFB225A0D6D7C30FE505550" box="[314,399,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="17" pageNumber="578">Fig. 5B</figureCitation>
|
||
) can be distinguished by the narrower and more needle-like apex of the median lobe as well as by the more narrowly rounded parameres. The aedeagus of
|
||
<taxonomicName id="4C154D54FFFB225A0F127C14FC02552C" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[837,989,1591,1617]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="subapicalis">
|
||
<emphasis id="B961EAC5FFFB225A0F127C14FC02552C" box="[837,989,1591,1617]" italics="true" pageId="17" pageNumber="578">P. subapicalis</emphasis>
|
||
</taxonomicName>
|
||
is considerably different (
|
||
<figureCitation id="132E2A52FFFB225A08AA7C14FABF552C" box="[1277,1376,1591,1617]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="17" pageNumber="578">Fig. 12B</figureCitation>
|
||
) as it lacks a needle-like apex to the median lobe. Females of
|
||
<taxonomicName id="4C154D54FFFB225A0EAB7C78FC835509" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[764,860,1627,1652]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFFB225A0EAB7C78FC835509" box="[764,860,1627,1652]" italics="true" pageId="17" pageNumber="578">nanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFFB225A0F347C78FC655508" bold="true" box="[867,954,1627,1653]" pageId="17" pageNumber="578">
|
||
<taxonomicNameLabel id="A25257BEFFFB225A0F347C78FC655508" box="[867,954,1627,1653]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the elytral glabrous region which is relatively narrower, occupying 9/11 of the elytral width (
|
||
<figureCitation id="132E2A52FFFB225A0FED7C5CFBD255E4" box="[954,1037,1663,1689]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="17" pageNumber="578">Fig. 9E</figureCitation>
|
||
) compared to that of
|
||
<taxonomicName id="4C154D54FFFB225A08AC7C5CFAB255E4" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1275,1389,1663,1689]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFFB225A08AC7C5CFAB255E4" box="[1275,1389,1663,1689]" italics="true" pageId="17" pageNumber="578">P. apicalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C154D54FFFB225A0CC07C80FEF155C0" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[151,302,1699,1725]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="subapicalis">
|
||
<emphasis id="B961EAC5FFFB225A0CC07C80FEF155C0" box="[151,302,1699,1725]" italics="true" pageId="17" pageNumber="578">P. subapicalis</emphasis>
|
||
</taxonomicName>
|
||
, in which it occupies 10/11 of the elytral width (
|
||
<figureCitation id="132E2A52FFFB225A0F117C80FC7A55C0" box="[838,933,1698,1725]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="17" pageNumber="578">Figs. 5E</figureCitation>
|
||
,
|
||
<figureCitation id="132E2A52FFFB225A0FE67C80FC3F55C0" box="[945,992,1699,1725]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="17" pageNumber="578">12E</figureCitation>
|
||
). Further, they have a differently shaped posterior margin of the elytra, being unevenly convex toward the suture with some extent of the convexity appearing subtruncate in the former, compared to evenly convex and rounded towards the suture in the latter two species. Finally, a well-impressed medial pit is always present on the mesoventrite of males of
|
||
<taxonomicName id="4C154D54FFFB225A08467D2CFB5B5454" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1041,1156,1807,1833]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B961EAC5FFFB225A08467D2CFB5B5454" box="[1041,1156,1807,1833]" italics="true" pageId="17" pageNumber="578">P. apicalis</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is frequently absent in both sexes of
|
||
<taxonomicName id="4C154D54FFFB225A0DC07D10FDCE5431" authority="Suksai & Gustafson & Sites & Sangpradub, 2021" authorityName="Suksai & Gustafson & Sites & Sangpradub" authorityYear="2021" box="[407,529,1843,1868]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
|
||
<emphasis id="B961EAC5FFFB225A0DC07D10FDCE5431" box="[407,529,1843,1868]" italics="true" pageId="17" pageNumber="578">P. nanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B961EAC5FFFB225A0E4F7D10FDB05430" bold="true" box="[536,623,1843,1869]" pageId="17" pageNumber="578">
|
||
<taxonomicNameLabel id="A25257BEFFFB225A0E4F7D10FDB05430" box="[536,623,1843,1869]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFFB225A0C907D74FF3D54E8" pageId="17" pageNumber="578" type="etymology">
|
||
<paragraph id="8BAA36D7FFFB225A0C907D74FF3D54E8" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C907D74FE8F540C" bold="true" box="[199,336,1879,1905]" pageId="17" pageNumber="578">Etymology.</emphasis>
|
||
The specific epithet is an adjective in reference to the province where this new species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F655CFFFB225A0C907DBDFDAF54A0" pageId="17" pageNumber="578" type="distribution">
|
||
<paragraph id="8BAA36D7FFFB225A0C907DBDFDAF54A0" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="578">
|
||
<emphasis id="B961EAC5FFFB225A0C907DBDFEBB54C5" bold="true" box="[199,356,1950,1976]" pageId="17" pageNumber="578">Distribution.</emphasis>
|
||
This species is currently known from only one high-elevation stream in Khun Sathan National Park,
|
||
<collectingRegion id="49D1F835FFFB225A0C8D7DE0FEA554A0" box="[218,378,1987,2013]" country="Thailand" name="Nan" pageId="17" pageNumber="578">Nan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3027647FFFB225A0DD27DE0FE3454A0" box="[389,491,1987,2013]" name="Thailand" pageId="17" pageNumber="578">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="132E2A52FFFB225A0DAD7DE0FDBC54A0" box="[506,611,1987,2013]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="17" pageNumber="578">Fig. 14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |