treatments-xml/data/03/BC/87/03BC87A6B34E5A1BFE4E3E47DF1FFAD9.xml

<document id="9C64B8A90FAC68932620EE2159EA825D" ID-CLB-Dataset="7807" ID-DOI="10.5281/zenodo.4503564" ID-GBIF-Dataset="24d1e5b6-6c26-4a68-8a63-43271c42a588" ID-ISSN="0374-1036" ID-Zenodo-Dep="4503564" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612461437530" checkinUser="felipe" docAuthor="Štys, Pavel &amp; Baňař, Petr" docDate="2007" docId="03BC87A6B34E5A1BFE4E3E47DF1FFAD9" docLanguage="en" docName="ActaEntMusNatPra.47.3-15.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 47" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Systelloderes loebli Štys &amp; Baňař 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FF85FFDEB34F5A13FF8F3D38DE5BFFC4" masterDocTitle="The first species of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from New Caledonia" masterLastPageNumber="15" masterPageNumber="3" pageNumber="4" updateTime="1698927106903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EC96A8BE88E971A6A6BFCBB1593EC45B">The first species of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from New Caledonia</mods:title>
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<mods:namePart id="E0DD155EA4EAAAA14C034AA8FAA4F9E7">Štys, Pavel</mods:namePart>
<mods:affiliation id="B59D38E0FA89940D1480621891034E63">) &amp;) Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: pavelstys @ gmail. com</mods:affiliation>
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<mods:namePart id="5B14966AFFA4362CE08A00EED7DE0A68">Baňař, Petr</mods:namePart>
<mods:affiliation id="993231C995047C94D931310C416D4577">) &amp;) Department of Forest Protection, Forestry and Game Management Research Institute, Jíloviště - Strnady, CZ- 156 04 Praha 5 - Zbraslav, Czech Republic; e-mail: petrbanar @ seznam. cz</mods:affiliation>
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<mods:title id="94C4E19E042F9372C1C39943538E486B">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="30B9FA0F6FF20A9279253F19F2CF1ADD">2007</mods:date>
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<paragraph id="8BAA36B0B34E5A12FE4E3E47DCB4FC5C" blockId="1.[449,751,895,920]" box="[449,751,895,920]" pageId="1" pageNumber="4">
<heading id="D0E281DCB34E5A12FE4E3E47DCB4FC5C" bold="true" box="[449,751,895,920]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="4" reason="3">
<emphasis id="B961EAA2B34E5A12FE4E3E47DCB4FC5C" bold="true" box="[449,751,895,920]" pageId="1" pageNumber="4">
<taxonomicName id="4C154D33B34E5A12FE4E3E47DCC8FC5C" ID-CoL="ffd239c9-f42b-412c-9f37-5da3b8b1f98b" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[449,659,895,920]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="4" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B34E5A12FE4E3E47DCC8FC5C" bold="true" box="[449,659,895,920]" italics="true" pageId="1" pageNumber="4">Systelloderes loebli</emphasis>
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<taxonomicNameLabel id="A25257D9B34E5A12FD163EB8DCB4FC5C" box="[665,751,896,920]" pageId="1" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30F653BB34E5A12FDAA3EA6DCD1FC75" box="[549,650,926,945]" pageId="1" pageNumber="4" type="description">
<paragraph id="8BAA36B0B34E5A12FDAA3EA6DCD1FC75" blockId="1.[549,650,926,945]" box="[549,650,926,945]" pageId="1" pageNumber="4">
(
<figureCitation id="132E2A35B34E5A12FDA43EA6DCD8FC75" box="[555,643,926,945]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="4.[151,190,1155,1174]" captionStartId-1="5.[113,152,896,915]" captionStartId-2="7.[113,152,1295,1314]" captionStartId-3="8.[152,191,640,659]" captionTargetBox-0="[152,1126,215,1129]" captionTargetBox-1="[133,1073,218,864]" captionTargetBox-2="[166,1053,217,1271]" captionTargetBox-3="[155,1122,221,613]" captionTargetId-0="figure-10@4.[152,1129,201,1132]" captionTargetId-1="figure-190@5.[104,1079,198,868]" captionTargetId-2="figure-89@7.[142,1060,195,1271]" captionTargetId-3="figure-294@8.[76,1134,210,619]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." captionText-1="Figs. 4-7. Systelloderes loebli sp. nov., holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: aprSCx – anterior prosupracoxale; ar – coxo-trochanteral articulation; atNPtl – articulatory tubercles of neopatella; bst – probasisternum; C – collum; Cx1 – fore coxa; F1 – fore femur and its articulatory processes; ltC – lateral tubercle of collum; NPtl – central part of neopatella; pc – precollum; pprSCx – (edge of) posterior prosupracoxale; s – coxal scale; se – stridulatory edge of basisternum; stll –prosternellum; Ti – fore tibia with a subdivided apex." captionText-2="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." captionText-3="Figs. 15-16. Systelloderes loebli sp. nov., holotype, female, right fore leg (anterior view); vestiture omitted. 15 – apicitibial armature; 16 – tarsal armature." figureDoi-0="http://doi.org/10.5281/zenodo.4503568" figureDoi-1="http://doi.org/10.5281/zenodo.4503570" figureDoi-2="http://doi.org/10.5281/zenodo.4503574" figureDoi-3="http://doi.org/10.5281/zenodo.4503576" httpUri-0="https://zenodo.org/record/4503568/files/figure.png" httpUri-1="https://zenodo.org/record/4503570/files/figure.png" httpUri-2="https://zenodo.org/record/4503574/files/figure.png" httpUri-3="https://zenodo.org/record/4503576/files/figure.png" pageId="1" pageNumber="4">Figs. 1-16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F653BB34E5A12FFFF3EEBDCF3FBF2" pageId="1" pageNumber="4" type="materials_examined">
<paragraph id="8BAA36B0B34E5A12FFFF3EEBDCDEFC22" blockId="1.[112,1088,978,1078]" box="[112,645,978,999]" pageId="1" pageNumber="4">
<emphasis id="B961EAA2B34E5A12FFFF3EEBDEB2FC22" bold="true" box="[112,233,978,998]" pageId="1" pageNumber="4">
<typeStatus id="54AE8812B34E5A12FFFF3EEBDEC6FC22" box="[112,157,979,998]" pageId="1" pageNumber="4">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3027620B34E5A12FF613EECDF2FFC22" box="[238,372,979,999]" name="New Caledonia" pageId="1" pageNumber="4">New Caledonia</collectingCountry>
, Mount Koghi, 
<quantity id="4CED9B55B34E5A12FE743EEBDC03FC23" box="[507,600,979,999]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="4" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400-500 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BAA36B0B34E5A12FFFF3ED6DCF3FBF2" blockId="1.[112,1088,978,1078]" pageId="1" pageNumber="4">
<emphasis id="B961EAA2B34E5A12FFFF3ED6DEAEFBC5" bold="true" box="[112,245,1005,1025]" pageId="1" pageNumber="4">Type material.</emphasis>
<typeStatus id="54AE8812B34E5A12FF753ED6DF16FBC4" box="[250,333,1006,1025]" pageId="1" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
: 
<specimenCount id="9D13FD39B34E5A12FED63ED3DF32FBC6" box="[345,361,1003,1026]" pageId="1" pageNumber="4" type="female">♀</specimenCount>
, ‘ 
<collectingCountry id="F3027620B34E5A12FEF83ED6DFA4FBC5" box="[375,511,1006,1025]" name="New Caledonia" pageId="1" pageNumber="4">New Caledonia</collectingCountry>
/ Mt. Koghi, prim. for. / 
<quantity id="4CED9B55B34E5A12FD5B3ED6DD75FBC5" box="[724,814,1006,1025]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="4" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400-500m</quantity>
, 
<date id="FFAB1070B34E5A12FCB83ED6DDE7FBC5" box="[823,956,1006,1025]" pageId="1" pageNumber="4" value="1998-10-18" valueMax="1998-10-19" valueMin="1998-10-18">18.-19.x / 1998</date>
, I. Löbl, litter’. The specimen bears the following red label: ‘ 
<typeStatus id="54AE8812B34E5A12FE673931DC06FBD8" box="[488,605,1033,1052]" pageId="1" pageNumber="4" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C154D33B34E5A12FDE43931DC80FBDF" authorityName="Blanchard" authorityYear="1852" box="[619,731,1033,1051]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B961EAA2B34E5A12FDE43931DC80FBDF" box="[619,731,1033,1051]" italics="true" pageId="1" pageNumber="4">Systelloderes</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C154D33B34E5A12FD653931DD40FBDF" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[746,795,1033,1051]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="4" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B34E5A12FD653931DD40FBDF" box="[746,795,1033,1051]" italics="true" pageId="1" pageNumber="4">loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B34E5A12FC903931DD3AFBD8" box="[799,865,1033,1052]" pageId="1" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
/ Štys &amp; Baňař det. 2007’; collection of Muséum d’Histoire Naturelle, 
<collectingRegion id="49D1F852B34E5A12FE69391BDC7CFBF2" box="[486,551,1059,1078]" country="Switzerland" name="Geneve" pageId="1" pageNumber="4">Genève</collectingRegion>
(
<collectingCountry id="F3027620B34E5A12FDBC391BDCC5FBF2" box="[563,670,1059,1078]" name="Switzerland" pageId="1" pageNumber="4">Switzerland</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F653BB34E5A12FFFF3976DF87FB23" pageId="1" pageNumber="4" type="diagnosis">
<paragraph id="8BAA36B0B34E5A12FFFF3976DF87FB23" blockId="1.[112,1087,1102,1255]" pageId="1" pageNumber="4">
<emphasis id="B961EAA2B34E5A12FFFF3976DEBEFBA2" bold="true" box="[112,229,1102,1126]" pageId="1" pageNumber="4">Diagnosis.</emphasis>
Large species, over 
<quantity id="4CED9B55B34E5A12FE453976DC52FBA2" box="[458,521,1102,1126]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
long, macropterous, brownish. Antennal segment 2 longer than segment 3, segments 2-4 isomorphic, terete. Fore leg: coxa with a long, narrow, free, erect scale; femur with a prominent basidorsal extension rising high above the level of tarsus; femoro-tibial membrane with dorsal neopatellar sclerites; tibial process absent; tibial and tarsal armatures as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="C30F653BB34E5A1BFFFF39C5DD11FB8A" lastPageId="8" lastPageNumber="11" pageId="1" pageNumber="4" type="description">
<paragraph id="8BAA36B0B34E5A12FFFF39C5DFABFAF2" blockId="1.[112,1088,1277,1590]" pageId="1" pageNumber="4">
<emphasis id="B961EAA2B34E5A12FFFF39C5DEA0FAD1" bold="true" box="[112,251,1277,1301]" pageId="1" pageNumber="4">Description.</emphasis>
Body elongate, moderately robust, extremities short, coloration and vestiture uniform and inconspicuous (
<figureCitation id="132E2A35B34E5A12FE2E3826DFBBFAF2" box="[417,480,1310,1334]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="1" pageNumber="4">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BAA36B0B34E5A11FF013806DF9CFEB0" blockId="1.[112,1088,1277,1590]" lastBlockId="2.[151,1127,219,1588]" lastPageId="2" lastPageNumber="5" pageId="1" pageNumber="4">
<emphasis id="B961EAA2B34E5A12FF013806DFA1FA92" bold="true" box="[142,506,1341,1366]" pageId="1" pageNumber="4">
Measurements. 
<emphasis id="B961EAA2B34E5A12FECF3805DFA1FA92" bold="true" box="[320,506,1341,1366]" italics="true" pageId="1" pageNumber="4">Total body length</emphasis>
</emphasis>
– 5.75-5.85 (abdomen deformed). 
<emphasis id="B961EAA2B34E5A12FCE93805DDFAFA92" bold="true" box="[870,929,1341,1366]" italics="true" pageId="1" pageNumber="4">Head</emphasis>
. Anterior lobe, L – 0.71; posterior lobe, L – 0.31, W – 0.38; distance of eye to apex of antennifer – 0.49; diatone (max W across eyes) – 0.36; min interocular distance, dorsal – 0.24; min interocular distance, ventral – 0.18; eye, L – 0.15. 
<emphasis id="B961EAA2B34E5A12FD9738A5DC35FA72" bold="true" box="[536,622,1437,1462]" italics="true" pageId="1" pageNumber="4">Labium</emphasis>
. Total, L – 0.95; segment 1, L – 0.12; segment 2, L – 0.21; segment 3, L – 0.48; segment 4, L – 0.16. 
<emphasis id="B961EAA2B34E5A12FD7C3886DD09FA12" bold="true" box="[755,850,1470,1494]" italics="true" pageId="1" pageNumber="4">Antenna</emphasis>
. Segment 1, L – 0.24; segment 2, L – 0.62; segment 3, L – 0.56; segment 4, L – 0.42. 
<emphasis id="B961EAA2B34E5A12FCA438E6DDC1FA31" bold="true" box="[811,922,1502,1525]" italics="true" pageId="1" pageNumber="4">Pronotum</emphasis>
. Total L (max) – 0.98; collum: L (median) – 0.18, max W – 0.49; midlobe: L (max) – 0.44, W (max) – 0.89; hindlobe: L (max) – 0.36, L (median) – 0.31, W (max) – 0.98. 
<emphasis id="B961EAA2B34E5A12FD753B26DD3EF9F2" bold="true" box="[762,869,1566,1590]" italics="true" pageId="1" pageNumber="4">Forewing</emphasis>
. Max L – 2.95. 
<emphasis id="B961EAA2B34E5A11FB853B26DEE3FF30" bold="true" italics="true" lastPageId="2" lastPageNumber="5" pageId="1" pageNumber="4">Fore leg</emphasis>
. Total L – 1.07, max W – 0.42; 
<emphasis id="B961EAA2B34D5A11FD883DE5DC78FF3E" box="[519,547,221,250]" italics="true" pageId="2" pageNumber="5">
Ti 
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</emphasis>
: L – 0.87, max W – 0.41;. 
<emphasis id="B961EAA2B34D5A11FCB83DE5DD02FF3E" box="[823,857,221,250]" italics="true" pageId="2" pageNumber="5">
Ta 
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</emphasis>
: L – 0.29, max W – 0.17; anterior claw, L (basis – apex) – 0.31; posterior claw, L (basis – apex) – 0.26. 
<emphasis id="B961EAA2B34D5A11FC483DC3DA60FED0" bold="true" box="[967,1083,251,276]" italics="true" pageId="2" pageNumber="5">Middle leg</emphasis>
. 
<emphasis id="B961EAA2B34D5A11FBC83DC5DA04FEDE" box="[1095,1119,253,282]" italics="true" pageId="2" pageNumber="5">
F 
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</emphasis>
: L – 0.84, max W – 0.18; 
<emphasis id="B961EAA2B34D5A11FE293C25DF98FEFE" box="[422,451,285,314]" italics="true" pageId="2" pageNumber="5">
Ti 
<subScript id="179134F5B34D5A11FE343C15DF98FEFE" attach="left" box="[443,451,301,314]" fontSize="6" pageId="2" pageNumber="5">2</subScript>
</emphasis>
: L – 0.69, max W – 0.13; 
<emphasis id="B961EAA2B34D5A11FD523C25DCA5FEFE" box="[733,766,285,314]" italics="true" pageId="2" pageNumber="5">
Ta 
<subScript id="179134F5B34D5A11FD793C15DCA5FEFE" attach="left" box="[758,766,301,314]" fontSize="6" pageId="2" pageNumber="5">2</subScript>
</emphasis>
: L (without claw) – 0.24, max W – 0.09. 
<emphasis id="B961EAA2B34D5A11FF673C03DF12FE90" bold="true" box="[232,329,315,340]" italics="true" pageId="2" pageNumber="5">Hind leg</emphasis>
. 
<emphasis id="B961EAA2B34D5A11FED83C05DF34FE9E" box="[343,367,317,346]" italics="true" pageId="2" pageNumber="5">
F 
<subScript id="179134F5B34D5A11FEE83C75DF34FE9E" attach="left" box="[359,367,333,346]" fontSize="6" pageId="2" pageNumber="5">3</subScript>
</emphasis>
: L – 1.02, max W – 0.29; 
<emphasis id="B961EAA2B34D5A11FD033C05DCF3FE9E" box="[652,680,317,346]" italics="true" pageId="2" pageNumber="5">
Ti 
<subScript id="179134F5B34D5A11FD2F3C75DCF3FE9E" attach="left" box="[672,680,333,346]" fontSize="6" pageId="2" pageNumber="5">3</subScript>
</emphasis>
: L – 1.16, max W – 0.14; 
<emphasis id="B961EAA2B34D5A11FC4A3C05DDBCFE9E" box="[965,999,317,346]" italics="true" pageId="2" pageNumber="5">
Ta 
<subScript id="179134F5B34D5A11FC503C75DDBCFE9E" attach="left" box="[991,999,333,346]" fontSize="6" pageId="2" pageNumber="5">3</subScript>
</emphasis>
: L (without claw) – 0.38, max W – 0.09.
</paragraph>
<paragraph id="8BAA36B0B34D5A11FF3A3C43DDDBFE70" blockId="2.[151,1127,219,1588]" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FF3A3C43DF6BFE57" bold="true" box="[181,304,379,403]" pageId="2" pageNumber="5">Coloration</emphasis>
(
<figureCitation id="132E2A35B34D5A11FEB03C44DF22FE50" box="[319,377,380,404]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="2" pageNumber="5">Fig.1</figureCitation>
). Body nearly unicolorous, between light-brown (antennae, middle and hind legs) and brown to dark-brown (head and midlobe of pronotum).
</paragraph>
<paragraph id="8BAA36B0B34D5A11FF3A3C84DA09FD30" blockId="2.[151,1127,219,1588]" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FF3A3C84DF4FFE10" bold="true" box="[181,276,444,468]" pageId="2" pageNumber="5">Texture.</emphasis>
Moderately shiny (including extremities), head lustrous. Dorsum of head with irregularly distributed, rather sparse, small setigerous tubercles, lateral and ventral sides of head, prothoracic collum, and dorsum of pronotal midlobe with a continuous cover of regularly distributed, minute setigerous tubercles (the latter particularly distinct in preocular area, on posterior cephalic lobe and on midlobe of pronotum, creating their shagreened appearance). Setigerous tubercles replaced by indistinct, large and shallow alveoles laterally on parts of prothorax and dorsum of pronotal hindlobe, the latter slightly rugulose. Mesoscutellum, wings and abdomen without particular structures. Antennae, labium, 
<emphasis id="B961EAA2B34D5A11FCA33FA5DD0AFD7E" box="[812,849,669,698]" italics="true" pageId="2" pageNumber="5">
Cx 
<subScript id="179134F5B34D5A11FCC63F95DD0AFD7E" attach="left" box="[841,849,685,698]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
, 
<emphasis id="B961EAA2B34D5A11FCD13FA5DD25FD7E" box="[862,894,669,698]" italics="true" pageId="2" pageNumber="5">
Tr 
<subScript id="179134F5B34D5A11FCF93F95DD25FD7E" attach="left" box="[886,894,685,698]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
, 
<emphasis id="B961EAA2B34D5A11FC053FA5DDF7FD7E" box="[906,940,669,698]" italics="true" pageId="2" pageNumber="5">
Ta 
<subScript id="179134F5B34D5A11FC2B3F95DDF7FD7E" attach="left" box="[932,940,685,698]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
, middle legs, and hind legs smooth. Anterior and posterior faces of 
<emphasis id="B961EAA2B34D5A11FD2D3F85DCE0FD1E" box="[674,699,701,730]" italics="true" pageId="2" pageNumber="5">
F 
<subScript id="179134F5B34D5A11FD3C3FF5DCE0FD1E" attach="left" box="[691,699,717,730]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34D5A11FD623F85DD51FD1E" box="[749,778,701,730]" italics="true" pageId="2" pageNumber="5">
Ti 
<subScript id="179134F5B34D5A11FC8D3FF5DD51FD1E" attach="left" box="[770,778,717,730]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
with scattered minute setigerous tubercles; longitudinal depression of anterior (mesal) face of 
<emphasis id="B961EAA2B34D5A11FCAC3FE5DD1BFD3E" box="[803,832,733,762]" italics="true" pageId="2" pageNumber="5">
Ti 
<subScript id="179134F5B34D5A11FCB73FD5DD1BFD3E" attach="left" box="[824,832,749,762]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
with transverse wrinkles.
</paragraph>
<paragraph id="8BAA36B0B34D5A10FF3A3FC3DDF4FCF0" blockId="2.[151,1127,219,1588]" lastBlockId="3.[113,1090,220,1620]" lastPageId="3" lastPageNumber="6" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FF3A3FC3DF79FCD7" bold="true" box="[181,290,763,787]" pageId="2" pageNumber="5">Vestiture.</emphasis>
Macrotrichia golden, ‘soft’, mostly short, straight, oblique, on some body parts (particularly posterior lobe of head, pronotum, and forewings) moderately to strongly curved. Macrotrichia mixed with much longer, diagonal, semierect to erect, conspicuous, mostly straight to only slightly curved 
<emphasis id="B961EAA2B34D5A11FE6C3E65DFA7FCB0" box="[483,508,861,884]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae with bilaterally symmetrical position; some of these may represent true trichobothria although the bothrium itself was not observed. Scales absent, except on 
<emphasis id="B961EAA2B34D5A11FE8F3EA5DF7DFC7E" box="[256,294,925,954]" italics="true" pageId="2" pageNumber="5">
Cx 
<subScript id="179134F5B34D5A11FE913E95DF7DFC7E" attach="left" box="[286,294,941,954]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
(see below). 
<emphasis id="B961EAA2B34D5A11FE203EA3DC10FC70" bold="true" box="[431,587,923,948]" italics="true" pageId="2" pageNumber="5">Head, labrum,</emphasis>
and 
<emphasis id="B961EAA2B34D5A11FDF43EA3DC95FC70" bold="true" box="[635,718,923,948]" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FDF43EA3DC9CFC70" bold="true" box="[635,711,923,948]" italics="true" pageId="2" pageNumber="5">labium</emphasis>
.
</emphasis>
Distribution of 
<emphasis id="B961EAA2B34D5A11FCFD3EA5DDD0FC70" box="[882,907,925,948]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae (longitudinally arranged if more than 1+1) as follows. Head: several 
<emphasis id="B961EAA2B34D5A11FD5E3E85DCB1FC10" box="[721,746,957,980]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae on and alongside anteclypeus and on labrum; 2+2 preocular (proximad to antennifers); 1+1 close to mesal eye margins, 2+2 postocellar; labial segment 1: dorsum/venter 1+1/0; 2: 3+3/0; 3: 3+3/2+2; 4: 3+3/3+3 (all strongly oblique and curved, similar to normal, elongate macrotrichia). Setation of ventral surface of head erect, short; longer and curved hairs on buccular part, gradually more curved and longer on proximal part close to neck, none trichobothrium-like. 
<emphasis id="B961EAA2B34D5A11FCE93964DD8AFBB0" bold="true" box="[870,977,1116,1140]" italics="true" pageId="2" pageNumber="5">Antennae</emphasis>
with uniform, short, straight, oblique setae; 
<emphasis id="B961EAA2B34D5A11FE453945DFB8FB50" box="[458,483,1149,1172]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae 1+1 on segment 1, then occurring from base of segment 2 to apex of segment 4, more elongate, denser and more erect distally. 
<emphasis id="B961EAA2B34D5A11FC1239A3DA55FB70" bold="true" box="[925,1038,1179,1204]" italics="true" pageId="2" pageNumber="5">Prothorax</emphasis>
. Dense, ‘soft’, short, mostly curved setae; 
<emphasis id="B961EAA2B34D5A11FE713985DC4CFB10" box="[510,535,1213,1236]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae: collum 1+1 anterolaterally, 1+1 posteromedially; midlobe: 1+1 posteromedially. 
<emphasis id="B961EAA2B34D5A11FE6639E3DCCEFB30" bold="true" box="[489,661,1243,1268]" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FE6639E3DCD4FB30" bold="true" box="[489,655,1243,1268]" italics="true" pageId="2" pageNumber="5">Mesoscutellum</emphasis>
.
</emphasis>
<emphasis id="B961EAA2B34D5A11FD1239E5DCEDFB30" box="[669,694,1245,1268]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae 2+2, in proximal part. 
<emphasis id="B961EAA2B34D5A11FC7E39E3DA3DFB37" bold="true" box="[1009,1126,1243,1267]" italics="true" pageId="2" pageNumber="5">Forewings</emphasis>
with sparse, short, curved microtrichia only on veins, none on wing membrane. 
<emphasis id="B961EAA2B34D5A11FC5539C4DA67FAD0" bold="true" box="[986,1084,1275,1300]" pageId="2" pageNumber="5">
<emphasis id="B961EAA2B34D5A11FC5539C4DA6DFAD0" bold="true" box="[986,1078,1275,1300]" italics="true" pageId="2" pageNumber="5">Fore leg</emphasis>
.
</emphasis>
<emphasis id="B961EAA2B34D5A11FBCE39C5DA3CFADE" box="[1089,1127,1277,1306]" italics="true" pageId="2" pageNumber="5">
Cx 
<subScript id="179134F5B34D5A11FBD03835DA3CFADE" attach="left" box="[1119,1127,1293,1306]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
with curved, nearly adpressed pubescence and two long, distiventral, diagonally pointing 
<emphasis id="B961EAA2B34D5A11FBC23825DA3DFAF0" box="[1101,1126,1309,1332]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae; a long, narrow, moderately ectally curved scale (L 0.12, 
<geoCoordinate id="EE215077B34D5A11FCAC3804DD2AFA90" box="[803,881,1340,1364]" degrees="0.02" direction="west" orientation="longitude" pageId="2" pageNumber="5" precision="555" value="-0.02">W 0.02</geoCoordinate>
) subapically on ventral surface, nearer mesal face (
<figureCitation id="132E2A35B34D5A11FE343864DC5DFAB0" box="[443,518,1372,1396]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="2" pageNumber="5">Figs. 3</figureCitation>
, 
<figureCitation id="132E2A35B34D5A11FD9D3863DC79FAB7" box="[530,546,1371,1395]" captionStart="Figs" captionStartId="5.[113,152,896,915]" captionTargetBox="[133,1073,218,864]" captionTargetId="figure-190@5.[104,1079,198,868]" captionTargetPageId="5" captionText="Figs. 4-7. Systelloderes loebli sp. nov., holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: aprSCx – anterior prosupracoxale; ar – coxo-trochanteral articulation; atNPtl – articulatory tubercles of neopatella; bst – probasisternum; C – collum; Cx1 – fore coxa; F1 – fore femur and its articulatory processes; ltC – lateral tubercle of collum; NPtl – central part of neopatella; pc – precollum; pprSCx – (edge of) posterior prosupracoxale; s – coxal scale; se – stridulatory edge of basisternum; stll –prosternellum; Ti – fore tibia with a subdivided apex." figureDoi="http://doi.org/10.5281/zenodo.4503570" httpUri="https://zenodo.org/record/4503570/files/figure.png" pageId="2" pageNumber="5">6</figureCitation>
). 
<emphasis id="B961EAA2B34D5A11FDBA3865DC0EFABE" box="[565,597,1373,1402]" italics="true" pageId="2" pageNumber="5">
Tr 
<subScript id="179134F5B34D5A11FDC23855DC0EFABE" attach="left" box="[589,597,1389,1402]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
with curved, long setae; on ventral face, with 3-4 long, curved 
<emphasis id="B961EAA2B34D5A11FEAF3845DF62FA50" box="[288,313,1405,1428]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae: the most distal one inserted at identical point as coxal scale; a few setae directed proximad, the other distad. 
<emphasis id="B961EAA2B34D5A11FD9638A5DC6AFA7E" box="[537,561,1437,1466]" italics="true" pageId="2" pageNumber="5">
F 
<subScript id="179134F5B34D5A11FDA63895DC6AFA7E" attach="left" box="[553,561,1453,1466]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
with short, oblique setae on dorsal and ventral faces, anterior and posterior faces nearly bare; erect 
<emphasis id="B961EAA2B34D5A11FDF83885DCCBFA10" box="[631,656,1469,1492]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae as follows: about 5+ 
<quantity id="4CED9B55B34D5A11FC2A3883DD97FA10" box="[933,972,1467,1492]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="5" unit="in" value="5.0">5 in</quantity>
double-row in distal half of dorsal face and multiple terminal dorsal cluster, and about 5+5 
<emphasis id="B961EAA2B34D5A11FC2138E5DD9CFA30" box="[942,967,1501,1524]" italics="true" pageId="2" pageNumber="5">trl</emphasis>
setae regularly distributed on ventral face. 
<emphasis id="B961EAA2B34D5A11FE3A38C5DF89F9DE" box="[437,466,1533,1562]" italics="true" pageId="2" pageNumber="5">
Ti 
<subScript id="179134F5B34D5A11FE453B35DF89F9DE" attach="left" box="[458,466,1549,1562]" fontSize="6" pageId="2" pageNumber="5">1</subScript>
</emphasis>
anterior face nearly bare, posterior and ventral faces with long, straight, oblique setae (particularly in distal half), dorsal and ventral faces and distal edge with numerous, irregularly distributed 
<emphasis id="B961EAA2B34C5A10FE433DE5DFBEFF30" box="[460,485,221,244]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae; other conspicuous macrotrichia: a single subpatellar 
<emphasis id="B961EAA2B34C5A10FFFE3DC5DED1FED0" box="[113,138,253,276]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
seta, and three very long, curly setae at distiventral angle. 
<emphasis id="B961EAA2B34C5A10FD7E3DC5DD48FEDE" box="[753,787,253,282]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FC843C35DD48FEDE" attach="left" box="[779,787,269,282]" fontSize="6" pageId="3" pageNumber="6">1</subScript>
</emphasis>
with long, straight, diagonal setae, especially dense and mostly curly on ventral face; conspicuous 
<emphasis id="B961EAA2B34C5A10FCED3C25DD20FEF0" box="[866,891,285,308]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae: 1+1 dorsal, subterminal, strikingly long, and 1+1 apicilateral claw-guarding setae (anterior one, close to the shorter claw, conspicuously shorter). 
<emphasis id="B961EAA2B34C5A10FDA23C63DC21FEB0" bold="true" box="[557,634,347,372]" italics="true" pageId="3" pageNumber="6">Middle</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FD3E3C63DD4DFEB0" bold="true" box="[689,790,347,372]" italics="true" pageId="3" pageNumber="6">hind legs</emphasis>
densely covered with short oblique to semierect setae on all faces; tibial and tarsal setae radiating relative to tibial and tarsal axes. 
<emphasis id="B961EAA2B34C5A10FF613CA5DF55FE7E" box="[238,270,413,442]" italics="true" pageId="3" pageNumber="6">
Tr 
<subScript id="179134F5B34C5A10FE893C95DF55FE7E" attach="left" box="[262,270,429,442]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FECE3CA5DF3AFE7E" box="[321,353,413,442]" italics="true" pageId="3" pageNumber="6">
Tr 
<subScript id="179134F5B34C5A10FED63C95DF3AFE7E" attach="left" box="[345,353,429,442]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
with two conspicuous erect ventral 
<emphasis id="B961EAA2B34C5A10FD6D3CA5DCA0FE70" box="[738,763,413,436]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae each. Ventral faces of 
<emphasis id="B961EAA2B34C5A10FBA73CA5DA1AFE7E" box="[1064,1089,413,442]" italics="true" pageId="3" pageNumber="6">
F 
<subScript id="179134F5B34C5A10FBB63C95DA1AFE7E" attach="right" box="[1081,1089,429,442]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FF2F3C85DEE3FE1E" box="[160,184,445,474]" italics="true" pageId="3" pageNumber="6">
F 
<subScript id="179134F5B34C5A10FF3F3CF5DEE3FE1E" attach="left" box="[176,184,461,474]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
each with one basal (adtrochanteral), one postmedial, and several distal suberect to erect 
<emphasis id="B961EAA2B34C5A10FF233CE5DE9EFE30" box="[172,197,477,500]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae, dorsal faces of 
<emphasis id="B961EAA2B34C5A10FE3D3CE5DF91FE3E" box="[434,458,477,506]" italics="true" pageId="3" pageNumber="6">
F 
<subScript id="179134F5B34C5A10FE4D3CD5DF91FE3E" attach="left" box="[450,458,493,506]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FE713CE5DC4CFE3E" box="[510,535,477,506]" italics="true" pageId="3" pageNumber="6">
F 
<subScript id="179134F5B34C5A10FD803CD5DC4CFE3E" attach="left" box="[527,535,493,506]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
each with 1+1 erect subterminal (adtibial) 
<emphasis id="B961EAA2B34C5A10FC6B3CE5DDA6FE30" box="[996,1021,477,500]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae. 
<emphasis id="B961EAA2B34C5A10FFFE3CC5DED5FDDE" box="[113,142,509,538]" italics="true" pageId="3" pageNumber="6">
Ti 
<subScript id="179134F5B34C5A10FF093F35DED5FDDE" attach="left" box="[134,142,525,538]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FF4F3CC5DE86FDDE" box="[192,221,509,538]" italics="true" pageId="3" pageNumber="6">
Ti 
<subScript id="179134F5B34C5A10FF5A3F35DE86FDDE" attach="left" box="[213,221,525,538]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
with long, oblique to erect 
<emphasis id="B961EAA2B34C5A10FE703CC5DC43FDD0" box="[511,536,509,532]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae developed along entire length,&gt;15 and&gt; 
<quantity id="4CED9B55B34C5A10FB843CC4DA1AFDD0" box="[1035,1089,508,532]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="6" unit="in" value="20.0">20 in</quantity>
number, respectively. 
<emphasis id="B961EAA2B34C5A10FED73F25DF22FDFE" box="[344,377,541,570]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FEFE3F15DF22FDFE" attach="left" box="[369,377,557,570]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B34C5A10FE263F25DF90FDFE" box="[425,459,541,570]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FE4C3F15DF90FDFE" attach="left" box="[451,459,557,570]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
(second ’segments’) with about 10 strongly oblique 
<emphasis id="B961EAA2B34C5A10FC623F25DA5DFDF0" box="[1005,1030,541,564]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae on 
<emphasis id="B961EAA2B34C5A10FF1C3F05DEEFFD9E" box="[147,180,573,602]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FF233F75DEEFFD9E" attach="left" box="[172,180,589,602]" fontSize="6" pageId="3" pageNumber="6">2</subScript>
</emphasis>
, over 12 on 
<emphasis id="B961EAA2B34C5A10FEB53F05DF07FD9E" box="[314,348,573,602]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FEDB3F75DF07FD9E" attach="left" box="[340,348,589,602]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
; setae on the latter undoubtedly homologous but not thin and 
<emphasis id="B961EAA2B34C5A10FC7D3F05DA50FD90" box="[1010,1035,573,596]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
like, and nearly spiniform instead. The only visually undoubtedly 
<emphasis id="B961EAA2B34C5A10FD753F65DD48FDB0" box="[762,787,605,628]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
seta on 
<emphasis id="B961EAA2B34C5A10FCE53F65DDD7FDBE" box="[874,908,605,634]" italics="true" pageId="3" pageNumber="6">
Ta 
<subScript id="179134F5B34C5A10FC0B3F55DDD7FDBE" attach="left" box="[900,908,621,634]" fontSize="6" pageId="3" pageNumber="6">3</subScript>
</emphasis>
is a ventral erect seta on its first ‘segment’. 
<emphasis id="B961EAA2B34C5A10FE1F3F43DFA2FD50" bold="true" box="[400,505,635,660]" italics="true" pageId="3" pageNumber="6">Abdomen</emphasis>
. Dorsum with short setae, without scales. Venter with golden short to moderately long, straight to slightly curved semierect macrotrichia (length and density increasing distally) intermixed with short, blackish, adpressed hairs. Some 
<emphasis id="B961EAA2B34C5A10FC643F85DA5FFD10" box="[1003,1028,701,724]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae occurring marginally; distribution of long 
<emphasis id="B961EAA2B34C5A10FDA03FE5DC13FD30" box="[559,584,733,756]" italics="true" pageId="3" pageNumber="6">trl</emphasis>
setae on posterior segments as follows: ventrite 7 – 1+1 at posterolateral angles, 1+1 submedial near posterior margin; subgenital plate: 1+1 submedial (extremely long) in basal part, 1+1 (very long) at posterior margin.
</paragraph>
<paragraph id="8BAA36B0B34C5A10FF1F3E03DD7CFAB0" blockId="3.[113,1090,220,1620]" pageId="3" pageNumber="6">
<emphasis id="B961EAA2B34C5A10FF1F3E03DF1AFC90" bold="true" box="[144,321,827,852]" pageId="3" pageNumber="6">
Structure. 
<emphasis id="B961EAA2B34C5A10FE893E03DF1AFC90" bold="true" box="[262,321,827,852]" italics="true" pageId="3" pageNumber="6">Head</emphasis>
</emphasis>
(
<figureCitation id="132E2A35B34C5A10FEC23E04DFF2FC90" box="[333,425,828,852]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="3" pageNumber="6">Figs. 1-2</figureCitation>
) strikingly narrow and long, slightly longer than pronotum (1.04 times as long as pronotum). Anterior lobe markedly longer than posterior lobe, 2.3 times as long. Lateral margin of preocular region parallel-sided proximally, its long distal part slightly convex, diverging towards antennifers; anteclypeus very long, narrow. Eye 0.31 times as long as distance between eye and antennifer. Postocular impression broad, shallow; lateral margin of postocular part of anterior cephalic lobe straight, diverging towards convex side of posterior lobe – postocular impression not marked in lateral outline. Posterior lobe transverse, 0.82 times as long as wide, its dorsum strongly convex, lateral sides moderately so. Eyes small, in lateral view not exceeding dorsal or ventral outlines of head; facets individually convex. Ocelli large, ocellar tubercles low. Ventral outline of head continuous, very slightly concave, only apex (fused bucculae) and basis (association with neck) outstanding. Dorsal ocular index 6.0, ventral ocular index 4.0. 
<emphasis id="B961EAA2B34C5A10FE2539A4DC4EFB70" bold="true" box="[426,533,1180,1204]" italics="true" pageId="3" pageNumber="6">Antennae</emphasis>
moderately long, thin; first segment strikingly long, cylindrical; segments 2-4 terete (not flagelliform, neither segment 4 subfusiform); antennal formula (longest segment first) 2, 3, 4, 1; segment 2 being 1.11 times as long as segment 3. 
<emphasis id="B961EAA2B34C5A10FFFE39C3DE9CFAD0" bold="true" box="[113,199,1275,1300]" italics="true" pageId="3" pageNumber="6">Labium</emphasis>
(
<figureCitation id="132E2A35B34C5A10FF5839C4DF43FAD0" box="[215,280,1276,1300]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="3" pageNumber="6">Fig. 2</figureCitation>
) moderately long, rather thin, directed anterad (segments 1, 2) and ventrad (segments 3, 4), without particular structures, labial formula (longest segment first) 3, 2, 4, 1. Segment 3: 2.2 times as long as segment 2. Ventral outline of segment 2 emarginate near base and at midlength. 
<emphasis id="B961EAA2B34C5A10FEE73863DF9AFAB0" bold="true" box="[360,449,1371,1396]" italics="true" pageId="3" pageNumber="6">Labrum</emphasis>
reaching to middle of segment 2.
</paragraph>
<paragraph id="8BAA36B0B34C5A16FF1F3844DA76F9D1" blockId="3.[113,1090,220,1620]" lastBlockId="5.[113,1089,1117,1557]" lastPageId="5" lastPageNumber="8" pageId="3" pageNumber="6">
<emphasis id="B961EAA2B34C5A10FF1F3844DEA4FA57" bold="true" box="[144,255,1404,1427]" italics="true" pageId="3" pageNumber="6">Pronotum</emphasis>
(
<figureCitation id="132E2A35B34C5A10FE823844DF34FA50" box="[269,367,1404,1428]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="3" pageNumber="6">Figs. 1-2</figureCitation>
). 
<emphasis id="B961EAA2B34C5A10FE0C3843DF8EFA50" bold="true" box="[387,469,1403,1428]" italics="true" pageId="3" pageNumber="6">Collum</emphasis>
short, 2.45 times as wide as long, with narrow precollum, dorsum with a linear impressed median area and pair of low and broad elevations, lateral area with low ‘pleural’ tubercle not visible in dorsal view. Collar constriction sharply delimited. 
<emphasis id="B961EAA2B34C5A10FFFE38E3DE90FA30" bold="true" box="[113,203,1499,1524]" italics="true" pageId="3" pageNumber="6">Midlobe</emphasis>
(dorsal side) with a linear, nearly percurrent median impression terminating just in front of posterior margin; disc with markedly plastic relief, with i) an inversely triangular anteromedial depression, ii) broad, not distinctly delimited, subcircular posteromedial depression, and iii) paired deep lateral pits emitting a lateral depression each; lateral margins broadly convex, interrupted, slightly notched because of lateral pits; posterior margin entire, trisinuate, sublateral shallowly concave parts slightly depressed, medial convex part broadly rounded, without edge. No traces of Y-shaped impressions. Constriction between midlobe and hindlobe broad, sharply demarcated. Midlobe 2.5 times as long as collum, 1.2 and 1.45 times as long as hindlobe maximum and median length, respectively. Midlobe 2.0 times as long as wide. 
<emphasis id="B961EAA2B34B5A17FEC23843DFE8FA50" bold="true" box="[333,435,1403,1428]" italics="true" pageId="4" pageNumber="7">Hindlobe</emphasis>
ample, its median twice as long as collum, indicated neither by ridge nor impression, lateral margins rounded, posterolateral angles (in strictly dorsal view) subrectangular; posterior margin bisinuate (tetrasinuate, if moderately protruding posterolateral angles are counted), medially broadly and shallowly concave, moderately convex sublaterally. Hindlobe 3.2 times as wide as medially long. ‘ 
<emphasis id="B961EAA2B34B5A17FC9638C3DD8CF9D0" bold="true" box="[793,983,1531,1556]" italics="true" pageId="4" pageNumber="7">Proepimeral lobe</emphasis>
’ (see 
<bibRefCitation id="EF844B41B34B5A17FB9738C3DF45F9F0" author="BANAR P. &amp; STYS P." pageId="4" pageNumber="7" pagination="7 - 14" refId="ref6316" refString="BANAR P. &amp; STYS P. 2006: A new species of Pseudohenschiella (Heteroptera: Enicocephalidae) from Madagascar. Acta Entomologica Musei Nationalis Pragae 46: 7 - 14." type="journal article" year="2006">ŠTYS &amp; BAŇAŘ 2006</bibRefCitation>
) extensive, distinctly exceeding posterior prosupracoxale posteroventrad, but not enclosing fore acetabula. 
<emphasis id="B961EAA2B34B5A17FE5C3B03DC22F990" bold="true" box="[467,633,1595,1620]" italics="true" pageId="4" pageNumber="7">Mesoscutellum</emphasis>
. Concave central part equilaterally triangular, produced in long, apically rounded mucro and included into a larger triangle due to lateral association with forewing grooves. 
<emphasis id="B961EAA2B34A5A16FE7B3944DCE8FB50" bold="true" box="[500,691,1148,1173]" italics="true" pageId="5" pageNumber="8">Prothoracic coxa</emphasis>
, 
<emphasis id="B961EAA2B34A5A16FD4D3944DD37FB51" bold="true" box="[706,876,1148,1173]" italics="true" pageId="5" pageNumber="8">prosupracoxale</emphasis>
, 
<emphasis id="B961EAA2B34A5A16FCF43944DA50FB51" bold="true" box="[891,1035,1148,1173]" italics="true" pageId="5" pageNumber="8">proacetabula</emphasis>
, and 
<emphasis id="B961EAA2B34A5A16FFFE39A5DEACFB70" bold="true" box="[113,247,1181,1204]" italics="true" pageId="5" pageNumber="8">prosternum.</emphasis>
For details see 
<figureCitation id="132E2A35B34A5A16FE1339A5DC7DFB70" box="[412,550,1180,1205]" captionStart="Figs" captionStartId="5.[113,152,896,915]" captionTargetBox="[133,1073,218,864]" captionTargetId="figure-190@5.[104,1079,198,868]" captionTargetPageId="5" captionText="Figs. 4-7. Systelloderes loebli sp. nov., holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: aprSCx – anterior prosupracoxale; ar – coxo-trochanteral articulation; atNPtl – articulatory tubercles of neopatella; bst – probasisternum; C – collum; Cx1 – fore coxa; F1 – fore femur and its articulatory processes; ltC – lateral tubercle of collum; NPtl – central part of neopatella; pc – precollum; pprSCx – (edge of) posterior prosupracoxale; s – coxal scale; se – stridulatory edge of basisternum; stll –prosternellum; Ti – fore tibia with a subdivided apex." figureDoi="http://doi.org/10.5281/zenodo.4503570" httpUri="https://zenodo.org/record/4503570/files/figure.png" pageId="5" pageNumber="8">Figs. 4 and 6</figureCitation>
. 
<emphasis id="B961EAA2B34A5A16FDBF39A6DC0DFB7F" box="[560,598,1182,1211]" italics="true" pageId="5" pageNumber="8">
Cx 
<subScript id="179134F5B34A5A16FDC13996DC0DFB7F" attach="left" box="[590,598,1198,1211]" fontSize="6" pageId="5" pageNumber="8">1</subScript>
</emphasis>
situated within proacetabulum, the latter open anteriorly and closed elsewhere. Proximal parts of lateral, anterior, and most of mesal faces of 
<emphasis id="B961EAA2B34A5A16FF0039E6DEEFFB3F" box="[143,180,1246,1275]" italics="true" pageId="5" pageNumber="8">
Cx 
<subScript id="179134F5B34A5A16FF2339D6DEEFFB3F" attach="left" box="[172,180,1262,1275]" fontSize="6" pageId="5" pageNumber="8">1</subScript>
</emphasis>
fully enclosed and externally delimited by prosupracoxalia. Anterior prosupracoxale excessively developed, anteromesally extended, embracing 
<emphasis id="B961EAA2B34A5A16FD5039C6DD5FFADF" box="[735,772,1278,1307]" italics="true" pageId="5" pageNumber="8">
Cx 
<subScript id="179134F5B34A5A16FD733836DD5FFADF" attach="left" box="[764,772,1294,1307]" fontSize="6" pageId="5" pageNumber="8">1</subScript>
</emphasis>
anterolaterally, anteriorly, and anteromesally. Transverse anterior part of anterior prosupracoxale fused with ventral part of collum (forming a not fully understood system of anteroventral prothoracic evaginations); anteromesal part of anterior prosupracoxale directed mesocaudad, fusing with triangular probasisternum, and taking part in formation of its strengthened, horizontal, sharp-edged lateral sides, here termed ‘ 
<emphasis id="B961EAA2B34A5A16FE0238A4DC89FA71" bold="true" box="[397,722,1436,1461]" italics="true" pageId="5" pageNumber="8">prosternal strigilatory edge(s)</emphasis>
’ (see Discussion). This composite triangular probasisternum horizontal, clearly delimited by its sharp edges; prosternellum adhering to probasisternum, sinuate in sagittal plane, more dorsal than probasisternum, with only the posterior, rounded, tongue-shaped part posteriad to coxae visible in ventral view.
</paragraph>
<caption id="DF6A6638B34B5A17FF1839BBDF55FB75" ID-DOI="http://doi.org/10.5281/zenodo.4503568" ID-Zenodo-Dep="4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="4" pageNumber="7" startId="4.[151,190,1155,1174]" targetBox="[152,1126,215,1129]" targetPageId="4">
<paragraph id="8BAA36B0B34B5A17FF1839BBDF55FB75" blockId="4.[151,1127,1155,1201]" pageId="4" pageNumber="7">
Figs. 1-3. 
<taxonomicName id="4C154D33B34B5A17FF6039BCDFCDFB52" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[239,406,1156,1174]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="7" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B34B5A17FF6039BCDFCDFB52" box="[239,406,1156,1174]" italics="true" pageId="4" pageNumber="7">Systelloderes loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B34B5A17FE1339BBDFBAFB52" box="[412,481,1155,1174]" pageId="4" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view.
</paragraph>
</caption>
<caption id="DF6A6638B34A5A16FFFE3EB8DDB3FBF7" ID-DOI="http://doi.org/10.5281/zenodo.4503570" ID-Zenodo-Dep="4503570" httpUri="https://zenodo.org/record/4503570/files/figure.png" pageId="5" pageNumber="8" startId="5.[113,152,896,915]" targetBox="[133,1073,218,864]" targetPageId="5">
<paragraph id="8BAA36B0B34A5A16FFFE3EB8DCF5FBF7" blockId="5.[113,1090,896,1075]" pageId="5" pageNumber="8">
Figs. 4-7. 
<taxonomicName id="4C154D33B34A5A16FF463EB9DF2BFC57" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[201,368,897,915]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B34A5A16FF463EB9DF2BFC57" box="[201,368,897,915]" italics="true" pageId="5" pageNumber="8">Systelloderes loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B34A5A16FEFA3EB8DFE0FC57" box="[373,443,896,915]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: 
<emphasis id="B961EAA2B34A5A16FFFE3EF7DEECFC27" bold="true" box="[113,183,975,995]" pageId="5" pageNumber="8">aprSCx</emphasis>
– anterior prosupracoxale; 
<emphasis id="B961EAA2B34A5A16FE133EE8DFEBFC27" bold="true" box="[412,432,976,995]" pageId="5" pageNumber="8">ar</emphasis>
– coxo-trochanteral articulation; 
<emphasis id="B961EAA2B34A5A16FD473EE8DD58FC27" bold="true" box="[712,771,976,995]" pageId="5" pageNumber="8">atNPtl</emphasis>
– articulatory tubercles of neopatella; 
<emphasis id="B961EAA2B34A5A16FFFE3ED3DED7FC3A" bold="true" box="[113,140,1003,1022]" pageId="5" pageNumber="8">bst</emphasis>
– probasisternum; 
<emphasis id="B961EAA2B34A5A16FEA43ED2DF61FC3A" bold="true" box="[299,314,1002,1022]" pageId="5" pageNumber="8">C</emphasis>
– collum; 
<emphasis id="B961EAA2B34A5A16FE1E3ED2DFEBFBC7" bold="true" box="[401,432,1002,1027]" pageId="5" pageNumber="8">
Cx 
<subScript id="179134F5B34A5A16FE253ECFDFEBFBC7" attach="left" box="[426,432,1015,1027]" fontSize="5" pageId="5" pageNumber="8">1</subScript>
</emphasis>
– fore coxa; 
<emphasis id="B961EAA2B34A5A16FD953ED3DC76FBC7" bold="true" box="[538,557,1003,1027]" pageId="5" pageNumber="8">
F 
<subScript id="179134F5B34A5A16FDA83ECFDC76FBC7" attach="left" box="[551,557,1015,1027]" fontSize="5" pageId="5" pageNumber="8">1</subScript>
</emphasis>
– fore femur and its articulatory processes; 
<emphasis id="B961EAA2B34A5A16FC193ED2DDE9FC3A" bold="true" box="[918,946,1002,1022]" pageId="5" pageNumber="8">ltC</emphasis>
– lateral tubercle of collum; 
<emphasis id="B961EAA2B34A5A16FF42393DDEADFBDC" bold="true" box="[205,246,1029,1048]" pageId="5" pageNumber="8">NPtl</emphasis>
– central part of neopatella; 
<emphasis id="B961EAA2B34A5A16FE6A393DDFA1FBDC" bold="true" box="[485,506,1029,1048]" pageId="5" pageNumber="8">pc</emphasis>
– precollum; 
<emphasis id="B961EAA2B34A5A16FDE0393DDCEDFBDD" bold="true" box="[623,694,1029,1049]" pageId="5" pageNumber="8">pprSCx</emphasis>
– (edge of) posterior prosupracoxale; 
<emphasis id="B961EAA2B34A5A16FC78393DDDA4FBDC" bold="true" box="[1015,1023,1029,1048]" pageId="5" pageNumber="8">s</emphasis>
– coxal scale; 
<emphasis id="B961EAA2B34A5A16FF283918DEE2FBF7" bold="true" box="[167,185,1056,1075]" pageId="5" pageNumber="8">se</emphasis>
– stridulatory edge of basisternum; 
<emphasis id="B961EAA2B34A5A16FE623918DC53FBF7" bold="true" box="[493,520,1056,1075]" pageId="5" pageNumber="8">stll</emphasis>
–prosternellum; 
<emphasis id="B961EAA2B34A5A16FD153927DCF5FBF7" bold="true" box="[666,686,1055,1075]" pageId="5" pageNumber="8">Ti</emphasis>
</paragraph>
<paragraph id="8BAA36B0B34A5A16FD213915DDB3FBF7" box="[686,1000,1056,1081]" lastBlockId="5.[113,1090,896,1075]" pageId="5" pageNumber="8">
<emphasis id="B961EAA2B34A5A16FD213915DCEFFBFD" bold="true" box="[686,692,1069,1081]" pageId="5" pageNumber="8">1</emphasis>
– fore tibia with a subdivided apex.
</paragraph>
</caption>
<paragraph id="8BAA36B0B3495A15FF3A3DE4DA7DFC37" blockId="6.[151,1127,219,1274]" pageId="6" pageNumber="9">
<emphasis id="B961EAA2B3495A15FF3A3DE4DEB2FF37" bold="true" box="[181,233,220,243]" italics="true" pageId="6" pageNumber="9">Legs</emphasis>
rather short. 
<emphasis id="B961EAA2B3495A15FEFA3DE4DF8AFF30" bold="true" box="[373,465,219,244]" italics="true" pageId="6" pageNumber="9">Fore leg</emphasis>
(
<figureCitation id="132E2A35B3495A15FE503DE4DC72FF30" box="[479,553,220,244]" captionStart="Figs" captionStartId="4.[151,190,1155,1174]" captionTargetBox="[152,1126,215,1129]" captionTargetId="figure-10@4.[152,1129,201,1132]" captionTargetPageId="4" captionText="Figs. 1-3. Systelloderes loebli sp. nov., holotype, female. 1 – total view; 2 – head, lateral view; 3 – right fore leg, anterior view." figureDoi="http://doi.org/10.5281/zenodo.4503568" httpUri="https://zenodo.org/record/4503568/files/figure.png" pageId="6" pageNumber="9">Figs. 3</figureCitation>
, 
<figureCitation id="132E2A35B3495A15FDBB3DE3DC3DFF37" box="[564,614,219,243]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="5.[113,152,896,915]" captionStartId-1="7.[113,152,1295,1314]" captionStartId-2="8.[152,191,640,659]" captionTargetBox-0="[133,1073,218,864]" captionTargetBox-1="[166,1053,217,1271]" captionTargetBox-2="[155,1122,221,613]" captionTargetId-0="figure-190@5.[104,1079,198,868]" captionTargetId-1="figure-89@7.[142,1060,195,1271]" captionTargetId-2="figure-294@8.[76,1134,210,619]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figs. 4-7. Systelloderes loebli sp. nov., holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: aprSCx – anterior prosupracoxale; ar – coxo-trochanteral articulation; atNPtl – articulatory tubercles of neopatella; bst – probasisternum; C – collum; Cx1 – fore coxa; F1 – fore femur and its articulatory processes; ltC – lateral tubercle of collum; NPtl – central part of neopatella; pc – precollum; pprSCx – (edge of) posterior prosupracoxale; s – coxal scale; se – stridulatory edge of basisternum; stll –prosternellum; Ti – fore tibia with a subdivided apex." captionText-1="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." captionText-2="Figs. 15-16. Systelloderes loebli sp. nov., holotype, female, right fore leg (anterior view); vestiture omitted. 15 – apicitibial armature; 16 – tarsal armature." figureDoi-0="http://doi.org/10.5281/zenodo.4503570" figureDoi-1="http://doi.org/10.5281/zenodo.4503574" figureDoi-2="http://doi.org/10.5281/zenodo.4503576" httpUri-0="https://zenodo.org/record/4503570/files/figure.png" httpUri-1="https://zenodo.org/record/4503574/files/figure.png" httpUri-2="https://zenodo.org/record/4503576/files/figure.png" pageId="6" pageNumber="9">6-16</figureCitation>
) extremely stout, femur and tibia incrassate. 
<emphasis id="B961EAA2B3495A15FBCE3DE5DA3CFF3E" box="[1089,1127,221,250]" italics="true" pageId="6" pageNumber="9">
Cx 
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</emphasis>
(
<figureCitation id="132E2A35B3495A15FF103DC4DEBBFED0" box="[159,224,252,276]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Fig. 8</figureCitation>
) conical, anteroventral face with dense, prominent, drop-like cuticular thorns (
<figureCitation id="132E2A35B3495A15FBB43DC4DEFEFEF0" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Fig. 9</figureCitation>
) resembling and arranged like rasping files. Antero- and especially posterodorsal proximal parts of fore coxa with dense rows of different cuticular processes (
<figureCitation id="132E2A35B3495A15FCC73C04DDCEFE90" box="[840,917,316,340]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Fig. 10</figureCitation>
), resembling serrate rasping files, each of those with several apical teeth, thus very similar to 
<taxonomicName id="4C154D33B3495A15FC113C65DFEBFE57" authority="Banar &amp; Stys 2006" authorityName="Banar &amp; Stys" authorityYear="2006" class="Insecta" family="Enicocephalidae" genus="Pseudohenschiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="9" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="B961EAA2B3495A15FC113C65DEBDFE50" italics="true" pageId="6" pageNumber="9">Pseudohenschiella hauseri</emphasis>
<bibRefCitation id="EF844B41B3495A15FF643C44DFEBFE57" author="BANAR P. &amp; STYS P." box="[235,432,379,404]" pageId="6" pageNumber="9" pagination="7 - 14" refId="ref6316" refString="BANAR P. &amp; STYS P. 2006: A new species of Pseudohenschiella (Heteroptera: Enicocephalidae) from Madagascar. Acta Entomologica Musei Nationalis Pragae 46: 7 - 14." type="journal article" year="2006">Baňař &amp; Štys 2006</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F3027620B3495A15FE633C44DC37FE50" box="[492,620,380,404]" name="Madagascar" pageId="6" pageNumber="9">Madagascar</collectingCountry>
(cf. 
<bibRefCitation id="EF844B41B3495A15FD163C44DD36FE57" author="BANAR P. &amp; STYS P." box="[665,877,379,404]" pageId="6" pageNumber="9" pagination="7 - 14" refId="ref6316" refString="BANAR P. &amp; STYS P. 2006: A new species of Pseudohenschiella (Heteroptera: Enicocephalidae) from Madagascar. Acta Entomologica Musei Nationalis Pragae 46: 7 - 14." type="journal article" year="2006">BAŇAŘ &amp; ŠTYS 2006</bibRefCitation>
). Proximal region of 
<emphasis id="B961EAA2B3495A15FBCA3C44DA3CFE5E" bold="true" box="[1093,1127,380,410]" italics="true" pageId="6" pageNumber="9">
Tr 
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</emphasis>
very narrow, more so than robust distal part, both regions separated by concave impression; entire adcoxal (dorsal) surface deeply concave (accommodating distal part of 
<emphasis id="B961EAA2B3495A15FC233C85DD8AFE1E" box="[940,977,445,474]" italics="true" pageId="6" pageNumber="9">
Cx 
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</emphasis>
during flexion of 
<emphasis id="B961EAA2B3495A15FF3A3CE5DE8EFE3E" box="[181,213,477,506]" italics="true" pageId="6" pageNumber="9">
Tr 
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</emphasis>
) and delimited by sharp, lateral, free anterodorsal and posterodorsal edges (about as long as one third of 
<emphasis id="B961EAA2B3495A15FEE83CC5DFD2FDD0" box="[359,393,509,532]" italics="true" pageId="6" pageNumber="9">tr-f</emphasis>
junction). 
<emphasis id="B961EAA2B3495A15FE753CC4DC4FFDDE" bold="true" box="[506,532,508,538]" italics="true" pageId="6" pageNumber="9">
F 
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</emphasis>
2.5 times as long as wide, with basidorsal angle forming subrectangularly produced, apically rounded dorsal extension, the latter rising strikingly above dorsal surface of 
<emphasis id="B961EAA2B3495A15FE193F05DFEDFD9E" box="[406,438,573,602]" italics="true" pageId="6" pageNumber="9">
Tr 
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</emphasis>
, and as high as its proximal diameter. 
<emphasis id="B961EAA2B3495A15FCDE3F04DDD2FD97" bold="true" box="[849,905,572,595]" italics="true" pageId="6" pageNumber="9">Knee</emphasis>
. Distinct remnant of a tripartite 
<emphasis id="B961EAA2B3495A15FE843F63DF26FDB0" bold="true" box="[267,381,603,628]" italics="true" pageId="6" pageNumber="9">neopatella</emphasis>
* (new term) visible dorsally in intersegmental 
<emphasis id="B961EAA2B3495A15FCFC3F65DDEBFDBE" box="[883,944,605,634]" italics="true" pageId="6" pageNumber="9">
F 
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-Ti 
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</emphasis>
membrane when tibia maximally bent towards femur (
<figureCitation id="132E2A35B3495A15FDA93F44DC33FD50" box="[550,616,636,660]" captionStart="Figs" captionStartId="5.[113,152,896,915]" captionTargetBox="[133,1073,218,864]" captionTargetId="figure-190@5.[104,1079,198,868]" captionTargetPageId="5" captionText="Figs. 4-7. Systelloderes loebli sp. nov., holotype, female. 4 – prosternum and associated structures, posteroventral view, diagrammatic; 5 – prosternum, sagittal section, diagrammatic; 6 – right fore coxa with a scale; anterior (mesal) view; diagrammatic; 7 – neopatella; lateral (dorsal) view on bent ‘knee’ of left fore leg; diagrammatic. Lettering: aprSCx – anterior prosupracoxale; ar – coxo-trochanteral articulation; atNPtl – articulatory tubercles of neopatella; bst – probasisternum; C – collum; Cx1 – fore coxa; F1 – fore femur and its articulatory processes; ltC – lateral tubercle of collum; NPtl – central part of neopatella; pc – precollum; pprSCx – (edge of) posterior prosupracoxale; s – coxal scale; se – stridulatory edge of basisternum; stll –prosternellum; Ti – fore tibia with a subdivided apex." figureDoi="http://doi.org/10.5281/zenodo.4503570" httpUri="https://zenodo.org/record/4503570/files/figure.png" pageId="6" pageNumber="9">Fig. 7</figureCitation>
). 
<emphasis id="B961EAA2B3495A15FDF33F43DCC0FD5E" bold="true" box="[636,667,635,666]" italics="true" pageId="6" pageNumber="9">
Ti 
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</emphasis>
broadly triangular, 2.1 times long as wide, compressed in anteroposterior plane, both anterior and posterior faces each with vaguely delimited longitudinal depression. Cleaning comb short, formed by tightly packed short setae; three ventralmost spiniform setae longer and stouter. Intersegmental tibiotarsal membrane forming an evaginated pocket stretching far ventrad beneath tarsus itself (enabling apparently its close appression towards distal margin of tibia). Distiventral, armature-bearing process absent. 
<emphasis id="B961EAA2B3495A15FF673E03DF9CFC97" bold="true" box="[232,455,827,852]" italics="true" pageId="6" pageNumber="9">Apicitibial armature</emphasis>
(
<figureCitation id="132E2A35B3495A15FE5A3E04DC7AFC97" box="[469,545,827,852]" captionStart="Figs" captionStartId="8.[152,191,640,659]" captionTargetBox="[155,1122,221,613]" captionTargetId="figure-294@8.[76,1134,210,619]" captionTargetPageId="8" captionText="Figs. 15-16. Systelloderes loebli sp. nov., holotype, female, right fore leg (anterior view); vestiture omitted. 15 – apicitibial armature; 16 – tarsal armature." figureDoi="http://doi.org/10.5281/zenodo.4503576" httpUri="https://zenodo.org/record/4503576/files/figure.png" pageId="6" pageNumber="9">Fig. 15</figureCitation>
) consisting of seven spiniform setae: two short ventral (straight), three long subventral (all slightly oblique towards tarsus), and two short dorsal (strongly oblique towards tarsus). 
<emphasis id="B961EAA2B3495A15FD893E44DC72FC5E" bold="true" box="[518,553,892,922]" italics="true" pageId="6" pageNumber="9">
Ta 
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</emphasis>
cylindrical, 1.7 times as long as wide, ventral surface slightly concave, 
<emphasis id="B961EAA2B3495A15FEDE3EA3DFA1FC77" bold="true" box="[337,506,923,948]" italics="true" pageId="6" pageNumber="9">tarsal armature</emphasis>
(
<figureCitation id="132E2A35B3495A15FD873EA4DC0FFC77" box="[520,596,923,948]" captionStart="Figs" captionStartId="8.[152,191,640,659]" captionTargetBox="[155,1122,221,613]" captionTargetId="figure-294@8.[76,1134,210,619]" captionTargetPageId="8" captionText="Figs. 15-16. Systelloderes loebli sp. nov., holotype, female, right fore leg (anterior view); vestiture omitted. 15 – apicitibial armature; 16 – tarsal armature." figureDoi="http://doi.org/10.5281/zenodo.4503576" httpUri="https://zenodo.org/record/4503576/files/figure.png" pageId="6" pageNumber="9">Fig. 16</figureCitation>
) of 1+1 proximal, curved spiniform setae and 1+1 distal setae (anterior one semicircular, posterior one broadly spiniform, shorter than proximal setae). 
<emphasis id="B961EAA2B3495A15FF6C3EE3DF7DFC30" bold="true" box="[227,294,987,1012]" italics="true" pageId="6" pageNumber="9">Claws</emphasis>
all of same shape, regularly curved, posterior one shorter and narrower.
</paragraph>
<paragraph id="8BAA36B0B3495A15FF3A3EC4DFC3FB37" blockId="6.[151,1127,219,1274]" pageId="6" pageNumber="9">
<emphasis id="B961EAA2B3495A15FF3A3EC4DF2AFBD0" bold="true" box="[181,369,1019,1044]" italics="true" pageId="6" pageNumber="9">Fore leg sensilla</emphasis>
on coxa, femur, and trochanter. Basal rim of 
<emphasis id="B961EAA2B3495A15FCE53EC5DDD4FBDE" box="[874,911,1021,1050]" italics="true" pageId="6" pageNumber="9">
Cx 
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</emphasis>
anteromesally with coxal rim organ (
<figureCitation id="132E2A35B3495A15FEDD3924DFC8FBF0" box="[338,403,1052,1076]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Fig. 8</figureCitation>
), consisting of cluster of several (5-7) differently directed, straight setae and one distant short seta. Condylar trochanteral organ (
<figureCitation id="132E2A35B3495A15FCB83904DD2CFB90" box="[823,887,1084,1108]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Fig. 8</figureCitation>
) consisting of several (six?) poorly visible short setae. Anterior trochanteral organ (
<figureCitation id="132E2A35B3495A15FCA63964DDC7FBB0" box="[809,924,1116,1140]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Figs. 8, 11</figureCitation>
) consisting of 6+1 campaniform sensilla (group of six, one isolated; posterior trochanteral organ (
<figureCitation id="132E2A35B3495A15FC6C3944DA05FB50" box="[995,1118,1148,1172]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Figs. 13-14</figureCitation>
) consisting of six campaniform sensilla (five in straight row, one isolated). Anterior femoral organ (
<figureCitation id="132E2A35B3495A15FF6D3984DF08FB10" box="[226,339,1212,1236]" captionStart="Figs" captionStartId="7.[113,152,1295,1314]" captionTargetBox="[166,1053,217,1271]" captionTargetId="figure-89@7.[142,1060,195,1271]" captionTargetPageId="7" captionText="Figs. 8-14. Systelloderes loebli sp. nov., holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, cto – condylar trochanteral organ, cro – coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ." figureDoi="http://doi.org/10.5281/zenodo.4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="6" pageNumber="9">Figs. 8, 12</figureCitation>
) consisting of 3+1 campaniform sensilla (group of three and one isolated) very close to base of 
<emphasis id="B961EAA2B3495A15FEF639E5DFCAFB3E" box="[377,401,1245,1274]" italics="true" pageId="6" pageNumber="9">
F 
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</emphasis>
.
</paragraph>
<paragraph id="8BAA36B0B3495A15FF183814DC41F9D1" blockId="6.[151,1126,1324,1562]" pageId="6" pageNumber="9">
<emphasis id="B961EAA2B3495A15FF183814DEA3FAFB" bold="true" box="[151,248,1324,1343]" pageId="6" pageNumber="9">Neopatella</emphasis>
(new term) = sclerite(s) situated within the dorsal section of the 
<emphasis id="B961EAA2B3495A15FCBB3815DD02FAFB" box="[820,857,1325,1343]" italics="true" pageId="6" pageNumber="9">F-Ti</emphasis>
intersegmental membrane in a position where the original patellar limb segment or its remnants would be situated (see Discussion). In fore leg of female 
<taxonomicName id="4C154D33B3495A15FF61385BDF6EFAB1" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[238,309,1379,1397]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="9" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B3495A15FF61385BDF6EFAB1" box="[238,309,1379,1397]" italics="true" pageId="6" pageNumber="9">S. loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B3495A15FEB5385ADF24FAB1" box="[314,383,1378,1397]" pageId="6" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, the dorsal apex of 
<emphasis id="B961EAA2B3495A15FDA9385BDC62FABE" box="[550,569,1379,1402]" italics="true" pageId="6" pageNumber="9">
F 
<subScript id="179134F5B3495A15FDBC3857DC62FABE" attach="left" box="[563,569,1391,1402]" fontSize="5" pageId="6" pageNumber="9">1</subScript>
</emphasis>
is concavely excised and the small dorsolateral projections of its posterior margin provide articulation with the neopatellar articulatory tubercles.The area between these projections is filled with a tripartite neopatella, its medial part forming a narrow arcuate strip adhering to the concavity of the apex of 
<emphasis id="B961EAA2B3495A15FF22388BDE9BFA0E" box="[173,192,1459,1482]" italics="true" pageId="6" pageNumber="9">
F 
<subScript id="179134F5B3495A15FF353887DE9BFA0E" attach="left" box="[186,192,1471,1482]" fontSize="5" pageId="6" pageNumber="9">1</subScript>
</emphasis>
, and its lateral parts being formed by strongly sclerotized, prominent, dorsally projecting articulatory tubercles associated with the medial strip of neopatella and articular projections of 
<emphasis id="B961EAA2B3495A15FC8438F5DD45FA21" box="[779,798,1485,1509]" italics="true" pageId="6" pageNumber="9">
F 
<subScript id="179134F5B3495A15FC9738E2DD45FA21" attach="left" box="[792,798,1498,1509]" fontSize="5" pageId="6" pageNumber="9">1</subScript>
</emphasis>
. The distal margin of the neopatella is associated with a short intersegmental 
<emphasis id="B961EAA2B3495A15FE5538D0DFA4FA3E" box="[474,511,1512,1530]" italics="true" pageId="6" pageNumber="9">F-Ti</emphasis>
membrane; the latter separates the neopatella from a produced and feebly delimited process of basidorsal margin of 
<emphasis id="B961EAA2B3495A15FE713B3BDC41F9D1" box="[510,538,1539,1562]" italics="true" pageId="6" pageNumber="9">
Ti 
<subScript id="179134F5B3495A15FD813B37DC4FF9DE" attach="left" box="[526,532,1551,1562]" fontSize="5" pageId="6" pageNumber="9">1</subScript>
.
</emphasis>
</paragraph>
<caption id="DF6A6638B3485A14FFFE3837DFD4FA49" ID-DOI="http://doi.org/10.5281/zenodo.4503574" ID-Zenodo-Dep="4503574" httpUri="https://zenodo.org/record/4503574/files/figure.png" pageId="7" pageNumber="10" startId="7.[113,152,1295,1314]" targetBox="[166,1053,217,1271]" targetPageId="7">
<paragraph id="8BAA36B0B3485A14FFFE3837DFD4FA49" blockId="7.[113,1090,1295,1421]" pageId="7" pageNumber="10">
Figs. 8-14. 
<taxonomicName id="4C154D33B3485A14FF5D3828DF23FAE6" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[210,376,1296,1314]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="10" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B3485A14FF5D3828DF23FAE6" box="[210,376,1296,1314]" italics="true" pageId="7" pageNumber="10">Systelloderes loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B3485A14FEF23837DF9AFAE6" box="[381,449,1295,1314]" pageId="7" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female, right fore leg (8-12 anterior view, 13-14 posterior view). 8 – coxa, trochanter and base of femur (cleared, vestiture omitted, 
<emphasis id="B961EAA2B3485A14FD3C3812DC95FAF9" bold="true" box="[691,718,1322,1341]" pageId="7" pageNumber="10">cto</emphasis>
– condylar trochanteral organ, 
<emphasis id="B961EAA2B3485A14FC513812DDA0FAF9" bold="true" box="[990,1019,1322,1341]" pageId="7" pageNumber="10">cro</emphasis>
– coxal rim organ); 9 – droplet-shaped rasplike microsculpture on fore coxa; 10 – serrate rasplike microsculpture on fore coxa; 11 – anterior trochanteral organ; 12 – anterior femoral organ; 13 – trochanter (cleared, vestiture omitted); 14 – posterior trochanteral organ.
</paragraph>
</caption>
<paragraph id="8BAA36B0B3485A1BFF1F38FBDC57FCAA" blockId="7.[113,1089,1475,1602]" lastBlockId="8.[151,1128,726,1102]" lastPageId="8" lastPageNumber="11" pageId="7" pageNumber="10">
<emphasis id="B961EAA2B3485A14FF1F38FBDF5EFA18" bold="true" box="[144,261,1475,1500]" italics="true" pageId="7" pageNumber="10">Middle leg</emphasis>
and 
<emphasis id="B961EAA2B3485A14FEB538FBDFCEFA18" bold="true" box="[314,405,1475,1500]" italics="true" pageId="7" pageNumber="10">hind leg</emphasis>
. Lateral (dorsal, adfemoral) face of 
<emphasis id="B961EAA2B3485A14FC9338FDDD19FA26" box="[796,834,1477,1506]" italics="true" pageId="7" pageNumber="10">
Cx 
<subScript id="179134F5B3485A14FCB538EDDD19FA26" attach="left" box="[826,834,1493,1506]" fontSize="6" pageId="7" pageNumber="10">3</subScript>
</emphasis>
flat, largely covered by sandpaper-like file similar to that on 
<emphasis id="B961EAA2B3485A14FD8238DDDC69F9C6" box="[525,562,1509,1538]" italics="true" pageId="7" pageNumber="10">
Cx 
<subScript id="179134F5B3485A14FDA538CDDC69F9C6" attach="left" box="[554,562,1525,1538]" fontSize="6" pageId="7" pageNumber="10">1</subScript>
</emphasis>
. 
<emphasis id="B961EAA2B3485A14FDCC38DDDC00F9C6" box="[579,603,1509,1538]" italics="true" pageId="7" pageNumber="10">
F 
<subScript id="179134F5B3485A14FDDC38CDDC00F9C6" attach="left" box="[595,603,1525,1538]" fontSize="6" pageId="7" pageNumber="10">3</subScript>
</emphasis>
moderately incrassate, its basidorsal margin arcuate, clearly extending above the level of dorsal face of 
<emphasis id="B961EAA2B3485A14FD503B3DDD5EF9D8" box="[735,773,1541,1570]" italics="true" pageId="7" pageNumber="10">
Tr 
<subScript id="179134F5B3485A14FD793B2DDCA5F9E6" attach="left" box="[758,766,1557,1570]" fontSize="6" pageId="7" pageNumber="10">3</subScript>
.
</emphasis>
Proximal segments otherwise without particular structures, anterior and posterior faces of 
<emphasis id="B961EAA2B3485A14FD763B1DDD4DF986" box="[761,790,1573,1602]" italics="true" pageId="7" pageNumber="10">
Ti 
<subScript id="179134F5B3485A14FC813B0DDD4DF986" attach="left" box="[782,790,1589,1602]" fontSize="6" pageId="7" pageNumber="10">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B3485A14FCC43B1DDD33F986" box="[843,872,1573,1602]" italics="true" pageId="7" pageNumber="10">
Ti 
<subScript id="179134F5B3485A14FCEF3B0DDD33F986" attach="left" box="[864,872,1589,1602]" fontSize="6" pageId="7" pageNumber="10">3</subScript>
</emphasis>
sulcate. 
<emphasis id="B961EAA2B3485A14FC473B1DDDB1F986" box="[968,1002,1573,1602]" italics="true" pageId="7" pageNumber="10">
Ta 
<subScript id="179134F5B3485A14FC6D3B0DDDB1F986" attach="left" box="[994,1002,1589,1602]" fontSize="6" pageId="7" pageNumber="10">2</subScript>
</emphasis>
and 
<emphasis id="B961EAA2B3485A14FB903B1DDA1AF986" box="[1055,1089,1573,1602]" italics="true" pageId="7" pageNumber="10">
Ta 
<subScript id="179134F5B3485A14FBB63B0DDA1AF986" attach="left" box="[1081,1089,1589,1602]" fontSize="6" pageId="7" pageNumber="10">3</subScript>
</emphasis>
two-segmented, segment 1 extremely short, without dorsal surface in lateral view (dorsal part of segment 1 visible as dorsal part of basitarsal ring filling up apex of tibia in anterodorsal view). Apices of both middle and hind tibiae each with strikingly short posteroventral and anteroventral setal combs; every comb terminating ventrally with a long spiniform seta. Claws and parempodial setae isomorphic.
</paragraph>
<caption id="DF6A6638B3475A1BFF173FB8DC51FD6B" ID-DOI="http://doi.org/10.5281/zenodo.4503576" ID-Zenodo-Dep="4503576" httpUri="https://zenodo.org/record/4503576/files/figure.png" pageId="8" pageNumber="11" startId="8.[152,191,640,659]" targetBox="[155,1122,221,613]" targetPageId="8">
<paragraph id="8BAA36B0B3475A1BFF173FB8DC51FD6B" blockId="8.[151,1126,640,687]" pageId="8" pageNumber="11">
Figs. 15-16. 
<taxonomicName id="4C154D33B3475A1BFE873FBADFEBFD50" authority="Štys &amp; Baňař, 2007" authorityName="Štys &amp; Baňař" authorityYear="2007" box="[264,432,642,660]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="11" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B961EAA2B3475A1BFE873FBADFEBFD50" box="[264,432,642,660]" italics="true" pageId="8" pageNumber="11">Systelloderes loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257D9B3475A1BFE383FB9DFA5FD50" box="[439,510,641,660]" pageId="8" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female, right fore leg (anterior view); vestiture omitted. 15 – apicitibial armature; 16 – tarsal armature.
</paragraph>
</caption>
<paragraph id="8BAA36B0B3475A1BFF3A3E4EDA1FFC0A" blockId="8.[151,1128,726,1102]" pageId="8" pageNumber="11">
<emphasis id="B961EAA2B3475A1BFF3A3E4EDF44FC4A" bold="true" box="[181,287,886,910]" italics="true" pageId="8" pageNumber="11">Forewing</emphasis>
as usual for 
<emphasis id="B961EAA2B3475A1BFE113E4FDC6BFC4A" box="[414,560,887,910]" italics="true" pageId="8" pageNumber="11">
<taxonomicName id="4C154D33B3475A1BFE113E4FDC70FC4A" authorityName="Blanchard" authorityYear="1852" box="[414,555,887,910]" class="Insecta" family="Enicocephalidae" genus="Systelloderes" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="11" phylum="Arthropoda" rank="genus">Systelloderes</taxonomicName>
.
</emphasis>
Pterostigma strikingly well formed, long and wide, RP arising from its middle; rp-mp (= the anterior crossvein) entering open discal cell strikingly more distad than CuA3+4 (= the posterior ‘crossvein’); AP in claval area not developed.
</paragraph>
<paragraph id="8BAA36B0B3475A1BFF3A3EEEDD11FB8A" blockId="8.[151,1128,726,1102]" pageId="8" pageNumber="11">
Ventral side of 
<emphasis id="B961EAA2B3475A1BFEDB3EEDDFE2FC2A" bold="true" box="[340,441,981,1006]" italics="true" pageId="8" pageNumber="11">abdomen</emphasis>
(distorted in the 
<typeStatus id="54AE8812B3475A1BFDE53EEEDC92FC2A" box="[618,713,982,1006]" pageId="8" pageNumber="11" type="holotype">holotype</typeStatus>
) with series of 1+1+1 large sclerites on ventrites 3-8. 
<taxonomicName id="4C154D33B3475A1BFEAC3ECDDFC2FBCA" box="[291,409,1013,1038]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="11" phylum="Tracheophyta" rank="genus">
<emphasis id="B961EAA2B3475A1BFEAC3ECDDFC2FBCA" bold="true" box="[291,409,1013,1038]" italics="true" pageId="8" pageNumber="11">Terminalia</emphasis>
</taxonomicName>
(distorted in the 
<typeStatus id="54AE8812B3475A1BFDCB3ECEDCFFFBCA" box="[580,676,1014,1038]" pageId="8" pageNumber="11" type="holotype">holotype</typeStatus>
). Posteromedial part of ventrite 7 thickened. Ventral laterotergite 8 distinct from subgenital plate, the latter strongly sclerotized, elongate, basal margin convex, distal margin concave in front of proctiger.
</paragraph>
</subSubSection>
<subSubSection id="C30F653BB3475A1BFF18395DDDB2FB19" pageId="8" pageNumber="11" type="etymology">
<paragraph id="8BAA36B0B3475A1BFF18395DDF28FB59" blockId="8.[151,1126,1125,1309]" pageId="8" pageNumber="11">
<emphasis id="B961EAA2B3475A1BFF18395DDF43FBB9" bold="true" box="[151,280,1125,1149]" pageId="8" pageNumber="11">Etymology.</emphasis>
Dedicated to Ivan Löbl (
<collectingRegion id="49D1F852B3475A1BFDA9395DDC26FBB9" box="[550,637,1125,1149]" country="Switzerland" name="Geneve" pageId="8" pageNumber="11">Genève</collectingRegion>
), an eminent coleopterist, our friend, and collector of the species.
</paragraph>
<paragraph id="8BAA36B0B3475A1BFF18399DDDB2FB19" blockId="8.[151,1126,1125,1309]" pageId="8" pageNumber="11">
<emphasis id="B961EAA2B3475A1BFF18399DDF4EFB79" bold="true" box="[151,277,1189,1213]" pageId="8" pageNumber="11">Bionomics.</emphasis>
The 
<typeStatus id="54AE8812B3475A1BFEC5399DDFFCFB79" box="[330,423,1189,1213]" pageId="8" pageNumber="11" type="holotype">holotype</typeStatus>
was collected by sieving leaf litter in a patch of a primary tropical New Caledonian rainforest surrounded by a secondary forest (I. Löbl, in epist.).
</paragraph>
</subSubSection>
<subSubSection id="C30F653BB3475A1BFF1839DDDF1FFAD9" pageId="8" pageNumber="11" type="distribution">
<paragraph id="8BAA36B0B3475A1BFF1839DDDF1FFAD9" blockId="8.[151,1126,1125,1309]" pageId="8" pageNumber="11">
<emphasis id="B961EAA2B3475A1BFF1839DDDF71FB39" bold="true" box="[151,298,1253,1277]" pageId="8" pageNumber="11">Distribution.</emphasis>
Known only from the 
<typeStatus id="54AE8812B3475A1BFDA839DEDC0EFB3A" box="[551,597,1254,1278]" pageId="8" pageNumber="11">type</typeStatus>
locality at Mount Koghi on the Grande Terre of 
<collectingCountry id="F3027620B3475A1BFF18383EDF1BFAD9" box="[151,320,1285,1309]" name="New Caledonia" pageId="8" pageNumber="11">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>