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<mods:title id="26CCF93F063BF1C97D3BB2B1816C70E9">The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n.</mods:title>
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<taxonomicName id="4C154D050E14FFCBFF44C3D7CD307709" ID-CoL="63MXN" ID-ENA="54968" box="[151,302,1263,1289]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C3D7CD307709" bold="true" box="[151,302,1263,1289]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="4C154D050E14FFCBFE63C3D7CEB77709" authority="Kieffer" authorityName="Kieffer" box="[432,681,1263,1289]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFE63C3D7CE567709" bold="true" box="[432,584,1263,1289]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
Kieffer
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</emphasis>
</heading>
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<subSubSection id="C30F650D0E14FFCBFF44C200CDDC77CF" pageId="10" pageNumber="11" type="reference_group">
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<taxonomicName id="4C154D050E14FFCBFF44C200CDC3774F" ID-CoL="63MXN" authority="Kieffer, 1913: 15" authorityName="Kieffer" authorityPageNumber="15" authorityYear="1913" box="[151,477,1336,1359]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C200CD00774F" box="[151,286,1336,1359]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
<treatmentCitation id="0AB410970E14FFCBFEF7C200CDC3774F" author="Kieffer" box="[292,477,1336,1359]" page="15" pageId="10" pageNumber="11" year="1913">
<bibRefCitation id="EF844B770E14FFCBFEF7C200CDA8774F" author="Kieffer" box="[292,438,1336,1359]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1913) Nouveaux Chironomides (Tendipedides) d'Allemagne. Bulletin de la Societe d'Histoire naturelle de Metz, 28, 7 - 35." type="journal article" year="1913">Kieffer, 1913</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
; not
<treatmentCitation id="0AB410970E14FFCBFDC1C200CEC6774F" author="Kieffer" box="[530,728,1336,1359]" page="41" pageId="10" pageNumber="11" year="1912">
<bibRefCitation id="EF844B770E14FFCBFDC1C200CEC6774F" author="Kieffer" box="[530,728,1336,1359]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1912) Tendipedidae (Chironomidae) (Dipt.). H. Sauter's Formosa-Ausbeute. Supplementa entomologica, 1, 27 - 43." type="journal article" year="1912">Kieffer (1912: 41)</bibRefCitation>
</treatmentCitation>
.
<taxonomicName id="4C154D050E14FFCBFF44C260CDCE776F" authority="Kieffer, 1922: 1" authorityName="Kieffer" authorityPageNumber="1" authorityYear="1922" box="[151,464,1368,1391]" class="Insecta" family="Chironomidae" genus="Kribiocharis" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C260CD01776F" box="[151,287,1368,1391]" italics="true" pageId="10" pageNumber="11">Kribiocharis</emphasis>
<treatmentCitation id="0AB410970E14FFCBFEF5C260CDCE776F" author="Kieffer" box="[294,464,1368,1391]" page="1" pageId="10" pageNumber="11" year="1922">
<bibRefCitation id="EF844B770E14FFCBFEF5C260CDA6776F" author="Kieffer" box="[294,440,1368,1391]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1922) Chironomides de l'Afrique equatoriale (2 e partie). Annales de la Societe entomologique de France, 91, 1 - 72." type="journal article" year="1922">Kieffer, 1922</bibRefCitation>
: 1
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BAA36860E14FFCBFF44C240CE7E778F" blockId="10.[151,868,1336,1487]" box="[151,608,1400,1423]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB0511A80E14FFCBFF44C240CE7E778F" box="[151,608,1400,1423]" pageId="10" pageNumber="11">
<taxonomicName id="4C154D050E14FFCBFF44C240CDFA778F" authority="Kieffer, 1921b: 98" authorityName="Kieffer" authorityPageNumber="98" authorityYear="1921" box="[151,484,1400,1423]" class="Insecta" family="Chironomidae" genus="Pentapelma" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C240CD08778F" box="[151,278,1400,1423]" italics="true" pageId="10" pageNumber="11">Pentapelma</emphasis>
<treatmentCitation id="0AB410970E14FFCBFECEC240CDFA778F" author="Kieffer" box="[285,484,1400,1423]" page="98" pageId="10" pageNumber="11" year="1921">
<bibRefCitation id="EF844B770E14FFCBFECEC240CDA3778F" author="Kieffer" box="[285,445,1400,1423]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1921 b) Nouveaux representants du genre Polypedilum (Chironomides). Annales de la Societe scientifique Bruxelles, 41, 97 - 101." type="journal article" year="1921" yearSuffix="b">Kieffer, 1921b</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
; 1921c: 33.
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</paragraph>
<paragraph id="8BAA36860E14FFCBFF44C2A0CDDC77CF" blockId="10.[151,868,1336,1487]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB0511A80E14FFCBFF44C2A0CDDC77CF" pageId="10" pageNumber="11">
<taxonomicName id="4C154D050E14FFCBFF44C2A0CDD877AF" authority="Kieffer, 1921a: 275" authorityName="Kieffer" authorityPageNumber="275" authorityYear="1921" box="[151,454,1432,1455]" class="Magnoliopsida" family="Asteraceae" genus="Rosenia" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C2A0CCF377AF" box="[151,237,1432,1455]" italics="true" pageId="10" pageNumber="11">Rosenia</emphasis>
<treatmentCitation id="0AB410970E14FFCBFF20C2A0CDD877AF" author="Kieffer" box="[243,454,1432,1455]" page="275" pageId="10" pageNumber="11" year="1921">
<bibRefCitation id="EF844B770E14FFCBFF20C2A0CD8F77AF" author="Kieffer" box="[243,401,1432,1455]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1921 a) Synopse de la tribu des Chironomariae (Dipteres). Annales de la Societe scientifique Bruxelles, 40, 269 - 277." type="journal article" year="1921" yearSuffix="a">Kieffer, 1921a</bibRefCitation>
: 275
</treatmentCitation>
</taxonomicName>
; not
<treatmentCitation id="0AB410970E14FFCBFE29C2A0CF7E77AF" author="Waagen" box="[506,864,1432,1455]" page="943" pageId="10" pageNumber="11" year="1887">
<bibRefCitation id="EF844B770E14FFCBFE29C2A0CF7E77AF" author="Waagen" box="[506,864,1432,1455]" pageId="10" pageNumber="32" refString="Waagen, W. &amp; Wentzel, J. (1887) Class Hydrozoa. Salt Range Fossils VI. Productus Limestone fossils. Memoirs of the Geological Survey of India, Series 13 (1), 925 - 962." type="journal article" year="1887">Waagen and Wentzel (1887: 943)</bibRefCitation>
</treatmentCitation>
.
<taxonomicName id="4C154D050E14FFCBFF44C280CDA377CF" authority="Lenz, 1937: 13" authorityName="Lenz" authorityPageNumber="13" authorityYear="1937" box="[151,445,1464,1487]" class="Insecta" family="Chironomidae" genus="Propedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFF44C280CD0A77CF" box="[151,276,1464,1487]" italics="true" pageId="10" pageNumber="11">Propedilum</emphasis>
<treatmentCitation id="0AB410970E14FFCBFEC9C280CDA377CF" author="Lenz" box="[282,445,1464,1487]" page="13" pageId="10" pageNumber="11" year="1937">
<bibRefCitation id="EF844B770E14FFCBFEC9C280CD8877CF" author="Lenz" box="[282,406,1464,1487]" pageId="10" pageNumber="30" refString="Lenz, F. (1937) Chironomariae aus Niederlandisch Indien. Larven und Puppen. Archiv fur Hydrobiologie, Supplement, 15, 1 - 29." type="journal article" year="1937">Lenz, 1937</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F650D0E14FFCBFF44C2C5CD357417" box="[151,299,1533,1559]" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BAA36860E14FFCBFF44C2C5CD357417" blockId="10.[151,752,1533,1657]" box="[151,299,1533,1559]" pageId="10" pageNumber="11">
<typeStatus id="54AE88240E14FFCBFF44C2C5CCD17417" box="[151,207,1533,1559]" pageId="10" pageNumber="11">Type</typeStatus>
species
</paragraph>
</subSubSection>
<subSubSection id="C30F650D0E14FFCAFF44C118CE4572FC" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BAA36860E14FFCBFF44C118CEB87479" blockId="10.[151,752,1533,1657]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB0511A80E14FFCBFF44C118CEB87479" pageId="10" pageNumber="11">
<taxonomicName id="4C154D050E14FFCBFF44C118CEEE743A" ID-CoL="8DG9P" authority="Meigen" authorityName="Meigen" box="[151,752,1568,1594]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nubeculosum" subGenus="Polypedilum">
<emphasis id="B961EA940E14FFCBFF44C118CE627439" box="[151,636,1568,1593]" italics="true" pageId="10" pageNumber="11">Polypedilum (Polypedilum) nubeculosum</emphasis>
(Meigen)
</taxonomicName>
<taxonomicName id="4C154D050E14FFCBFF14C17ACE877459" ID-CoL="87WQ8" authority="Meigen, 1804: 18" authorityName="Meigen" authorityPageNumber="18" authorityYear="1804" box="[199,665,1602,1625]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nubeculosus">
<emphasis id="B961EA940E14FFCBFF14C17ACDCA7459" box="[199,468,1602,1625]" italics="true" pageId="10" pageNumber="11">Chironomus nubeculosus</emphasis>
<treatmentCitation id="0AB410970E14FFCBFE09C17ACE877459" author="Meigen" box="[474,665,1602,1625]" page="18" pageId="10" pageNumber="11" year="1804">
<bibRefCitation id="EF844B770E14FFCBFE09C17ACE6C7459" author="Meigen" box="[474,626,1602,1625]" pageId="10" pageNumber="30" refString="Meigen, J. W. (1804) Klassifikazion und Beschreibung der europaischen zweiflugeligen Insekten. (Diptera Linn.). Erster Band, Abteilung 1. Reichard, Braunsweig, xxviii + 152 pp." type="book" year="1804">Meigen, 1804</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="4C154D050E14FFCBFF14C15ACEBC7479" ID-CoL="8C6QB" authority="Kieffer, 1913: 16" authorityName="Kieffer" authorityPageNumber="16" authorityYear="1913" box="[199,674,1634,1657]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="emarginatum">
<emphasis id="B961EA940E14FFCBFF14C15ACDFF7479" box="[199,481,1634,1657]" italics="true" pageId="10" pageNumber="11">Polypedilum emarginatum</emphasis>
<treatmentCitation id="0AB410970E14FFCBFE3BC15ACEBC7479" author="Kieffer" box="[488,674,1634,1657]" page="16" pageId="10" pageNumber="11" year="1913">
<bibRefCitation id="EF844B770E14FFCBFE3BC15ACE647479" author="Kieffer" box="[488,634,1634,1657]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1913) Nouveaux Chironomides (Tendipedides) d'Allemagne. Bulletin de la Societe d'Histoire naturelle de Metz, 28, 7 - 35." type="journal article" year="1913">Kieffer, 1913</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36860E14FFCAFF44C190CE4572FC" blockId="10.[151,1436,1704,2028]" lastBlockId="11.[151,1437,152,1002]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
The genus
<taxonomicName id="4C154D050E14FFCBFECEC190CE0874C2" ID-CoL="63MXN" authority="Kieffer" authorityName="Kieffer" box="[285,534,1704,1730]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFECEC190CDA874C1" box="[285,438,1704,1729]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
Kieffer
</taxonomicName>
was described in 1913 and
<taxonomicName id="4C154D050E14FFCBFCBFC190C94774C2" ID-CoL="8C6QB" authority="Kieffer, 1913" authorityName="Kieffer" authorityYear="1913" box="[876,1369,1704,1730]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="emarginatum">
<emphasis id="B961EA940E14FFCBFCBFC190C8B074C1" box="[876,1198,1704,1729]" italics="true" pageId="10" pageNumber="11">Polypedilum emarginatum</emphasis>
<bibRefCitation id="EF844B770E14FFCBFB64C190C94774C2" author="Kieffer" box="[1207,1369,1704,1730]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1913) Nouveaux Chironomides (Tendipedides) d'Allemagne. Bulletin de la Societe d'Histoire naturelle de Metz, 28, 7 - 35." type="journal article" year="1913">Kieffer, 1913</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="4C154D050E14FFCBFA55C190CDE174E7" ID-CoL="8DG9P" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nubeculosum">
<emphasis id="B961EA940E14FFCBFA55C190CD2974E6" italics="true" pageId="10" pageNumber="11">P. nubeculosum</emphasis>
(Meigen, 1818)
</taxonomicName>
] designated as the
<typeStatus id="54AE88240E14FFCBFD3AC1F5CF0374E7" box="[745,797,1741,1767]" pageId="10" pageNumber="11">type</typeStatus>
of the genus (
<bibRefCitation id="EF844B770E14FFCBFC1EC1F5C88174E7" author="Kieffer" box="[973,1183,1741,1767]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1913) Nouveaux Chironomides (Tendipedides) d'Allemagne. Bulletin de la Societe d'Histoire naturelle de Metz, 28, 7 - 35." type="journal article" year="1913">Kieffer 1913: 15</bibRefCitation>
). As shown by
<bibRefCitation id="EF844B770E14FFCBFA8DC1F5CCF9750C" author="Ashe" pageId="10" pageNumber="29" refString="Ashe, P. (1981) A taxonomic note on the genus Polypedilum Kieffer (Diptera: Chironomidae) with an earlier date for the genus and designation of a type species. Aquatic Insects, 3, 57 - 59." type="journal article" year="1981">Ashe (1981)</bibRefCitation>
, however, the
<typeStatus id="54AE88240E14FFCBFE42C1CACDDD750C" box="[401,451,1778,1804]" pageId="10" pageNumber="11">type</typeStatus>
designation was invalid as two species were assigned to the genus
<taxonomicName id="4C154D050E14FFCBFB06C1CBC977750C" ID-CoL="63MXN" box="[1237,1385,1779,1804]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E14FFCBFB06C1CBC977750C" box="[1237,1385,1779,1804]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
</taxonomicName>
in a paper published a year earlier (
<bibRefCitation id="EF844B770E14FFCBFDC5C020CEF77532" author="Kieffer" box="[534,745,1816,1842]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1912) Tendipedidae (Chironomidae) (Dipt.). H. Sauter's Formosa-Ausbeute. Supplementa entomologica, 1, 27 - 43." type="journal article" year="1912">Kieffer 1912: 41</bibRefCitation>
). Of the two species,
<taxonomicName id="4C154D050E14FFCBFC25C020C9577532" authority="Kieffer, 1912" authorityName="Kieffer" authorityYear="1912" box="[1014,1353,1816,1842]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pelostolum">
<emphasis id="B961EA940E14FFCBFC25C020C8857531" box="[1014,1179,1816,1841]" italics="true" pageId="10" pageNumber="11">P. pelostolum</emphasis>
<bibRefCitation id="EF844B770E14FFCBFB77C020C9577532" author="Kieffer" box="[1188,1353,1816,1842]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1912) Tendipedidae (Chironomidae) (Dipt.). H. Sauter's Formosa-Ausbeute. Supplementa entomologica, 1, 27 - 43." type="journal article" year="1912">Kieffer, 1912</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C154D050E14FFCBFA55C020CDC97557" authority="Kieffer, 1912" authorityName="Kieffer" authorityYear="1912" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ceylanicum">
<emphasis id="B961EA940E14FFCBFA55C020CD037556" italics="true" pageId="10" pageNumber="11">P. ceylanicum</emphasis>
(
<bibRefCitation id="EF844B770E14FFCBFEFFC005CDD17557" author="Kieffer" box="[300,463,1853,1879]" pageId="10" pageNumber="30" refString="Kieffer, J. J. (1912) Tendipedidae (Chironomidae) (Dipt.). H. Sauter's Formosa-Ausbeute. Supplementa entomologica, 1, 27 - 43." type="journal article" year="1912">Kieffer, 1912</bibRefCitation>
)
</taxonomicName>
, at least the first according to
<bibRefCitation id="EF844B770E14FFCBFCE9C005CFE97557" author="Freeman" box="[826,1015,1853,1879]" pageId="10" pageNumber="29" refString="Freeman, P. (1961) The Chironomidae (Diptera) of Australia. British Museum (Natural History), London, pp. 611 - 737." type="book chapter" year="1961">Freeman (1961)</bibRefCitation>
is a synonym of
<taxonomicName id="4C154D050E14FFCBFB11C005C92A7556" ID-CoL="8BGPP" box="[1218,1332,1853,1878]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B961EA940E14FFCBFB11C005C92A7556" box="[1218,1332,1853,1878]" italics="true" pageId="10" pageNumber="11">P. nubifer</emphasis>
</taxonomicName>
and was selected as the
<typeStatus id="54AE88240E14FFCBFE95C05ACD66757C" box="[326,376,1890,1916]" pageId="10" pageNumber="11">type</typeStatus>
species. However,
<bibRefCitation id="EF844B770E14FFCBFD88C05ACF08757C" author="Freeman" box="[603,790,1890,1916]" pageId="10" pageNumber="29" refString="Freeman, P. (1961) The Chironomidae (Diptera) of Australia. British Museum (Natural History), London, pp. 611 - 737." type="book chapter" year="1961">Freeman (1961)</bibRefCitation>
stated that
<taxonomicName id="4C154D050E14FFCBFC48C05BC812757C" ID-CoL="8BGPP" box="[923,1036,1891,1916]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B961EA940E14FFCBFC48C05BC812757C" box="[923,1036,1891,1916]" italics="true" pageId="10" pageNumber="11">P. nubifer</emphasis>
</taxonomicName>
is without frontal tubercles, while the presence of frontal tubercles is one of the most significant features of the species. Two
<typeStatus id="54AE88240E14FFCBFB21C0B0C94275A2" box="[1266,1372,1928,1954]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C154D050E14FFCBFA55C0B0CD0675C6" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pelostolum">
<emphasis id="B961EA940E14FFCBFA55C0B0CD0675C6" italics="true" pageId="10" pageNumber="11">P. pelostolum</emphasis>
</taxonomicName>
from The Natural History Museum in London were examined. The specimens were lacking wings, antennae, hypopygia and most legs, but one head with frontal tubercles intact remained. Accordingly the synonymy with
<taxonomicName id="4C154D050E15FFCAFEB2C7A0CDC272B1" ID-CoL="8BGPP" box="[353,476,152,177]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B961EA940E15FFCAFEB2C7A0CDC272B1" box="[353,476,152,177]" italics="true" pageId="11" pageNumber="12">P. nubifer</emphasis>
</taxonomicName>
has to be accepted. As shown here
<taxonomicName id="4C154D050E15FFCAFC72C7A0C80572B1" ID-CoL="8BGPP" box="[929,1051,152,177]" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nubifer">
<emphasis id="B961EA940E15FFCAFC72C7A0C80572B1" box="[929,1051,152,177]" italics="true" pageId="11" pageNumber="12">P. nubifer</emphasis>
</taxonomicName>
, however, should be placed in
<taxonomicName id="4C154D050E15FFCAFF44C785CD0772D6" box="[151,281,189,214]" class="Insecta" family="Chironomidae" genus="Tripedilum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E15FFCAFF44C785CD0772D6" box="[151,281,189,214]" italics="true" pageId="11" pageNumber="12">Tripedilum</emphasis>
</taxonomicName>
. A case will be made to ICZN for rejecting the
<typeStatus id="54AE88240E15FFCAFC86C785CF9672D7" box="[853,904,189,215]" pageId="11" pageNumber="12">type</typeStatus>
designation of
<bibRefCitation id="EF844B770E15FFCAFB93C785C8CA72D7" author="Ashe" box="[1088,1236,189,215]" pageId="11" pageNumber="29" refString="Ashe, P. (1981) A taxonomic note on the genus Polypedilum Kieffer (Diptera: Chironomidae) with an earlier date for the genus and designation of a type species. Aquatic Insects, 3, 57 - 59." type="journal article" year="1981">Ashe (1981)</bibRefCitation>
and maintaining the
<typeStatus id="54AE88240E15FFCAFF11C7DACCEB72FC" box="[194,245,226,252]" pageId="11" pageNumber="12">type</typeStatus>
designation of
<bibRefCitation id="EF844B770E15FFCAFE79C7DACE4972FC" author="Kieffer" box="[426,599,226,252]" pageId="11" pageNumber="30" refString="Kieffer, J. J. (1913) Nouveaux Chironomides (Tendipedides) d'Allemagne. Bulletin de la Societe d'Histoire naturelle de Metz, 28, 7 - 35." type="journal article" year="1913">Kieffer (1913)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F650D0E15FFCAFF15C63FC930712F" pageId="11" pageNumber="12" type="description">
<paragraph id="8BAA36860E15FFCAFF15C63FCF6C7394" blockId="11.[151,1437,152,1002]" pageId="11" pageNumber="12">
<emphasis id="B961EA940E15FFCAFF15C63FCDFE7321" bold="true" box="[198,480,263,289]" pageId="11" pageNumber="12">Diagnostic characters.</emphasis>
The male imagines of the subgenus can be distinguished by having no frontal tubercles, bare wing membrane, wing vein R2+3 usually relatively well separated from R1, projection of superior volsella long compared to the short base and nearly always with outer seta and anal point never trifid or with lateral teeth or shoulders to each side of the anal point.
</paragraph>
<paragraph id="8BAA36860E15FFCAFF17C698C8E973DF" blockId="11.[151,1437,152,1002]" pageId="11" pageNumber="12">The few female imagines sufficiently described have nearly straight spermathecal ducts and the dorsomesal lobes of gonapophysis VIII are not reaching much caudal of the ventrolateral lobes.</paragraph>
<paragraph id="8BAA36860E15FFCAFF17C6D2CEFC704F" blockId="11.[151,1437,152,1002]" pageId="11" pageNumber="12">The pupa has no frontal tubercles and no or vestigial cephalic tubercles, less than 20 branches in thoracic horn, single spur on caudolateral margin of segment VIII, taeniae on segments V and VI may be reduced in size or number and anal lobe without dorsal setae.</paragraph>
<paragraph id="8BAA36860E15FFCAFF17C562C930712F" blockId="11.[151,1437,152,1002]" pageId="11" pageNumber="12">
The larva has opposing Lauterborn organs; third antennal segment distinctly longer to somewhat shorter than fourth;
<taxonomicName id="4C154D050E15FFCAFEF8C5B8CE02709A" box="[299,540,640,666]" class="Bivalvia" family="Pectinidae" genus="Pecten" kingdom="Animalia" order="Pectinoida" pageId="11" pageNumber="22" phylum="Mollusca" rank="species" species="epipharyngis">pecten epipharyngis</taxonomicName>
occasionally with less than 3 teeth in median scale, usually 35; mandible occasionally without dorsal tooth, usually with; occasionally mola smooth, usually serrate; mentum usually with first lateral teeth distinctly lower than second and median teeth, but often at least as high as median and/ or second lateral teeth; and ventromental plates without posterior lobes, with median apices pointing towards each other and distance between plates usually at least as long as combined width of 4 median teeth.
</paragraph>
</subSubSection>
<subSubSection id="C30F650D0E15FFCAFF15C402C97471EA" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BAA36860E15FFCAFF15C402C97471EA" blockId="11.[151,1437,152,1002]" pageId="11" pageNumber="12">
<emphasis id="B961EA940E15FFCAFF15C402CD217154" bold="true" box="[198,319,826,852]" pageId="11" pageNumber="12">Remarks.</emphasis>
The pupae of
<emphasis id="B961EA940E15FFCAFE3EC403CEDC7154" box="[493,706,827,852]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C154D050E15FFCAFE3EC403CE9F7154" box="[493,641,827,852]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" sensu="stricto">Polypedilum</taxonomicName>
<taxonomicNameLabel id="A25257EF0E15FFCAFD59C403CEDC7154" box="[650,706,827,852]" pageId="11" pageNumber="12" sensu="stricto">s. str</taxonomicNameLabel>
</emphasis>
. are not separable from those of
<taxonomicName id="4C154D050E15FFCAFB98C403C8F07154" box="[1099,1262,827,852]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E15FFCAFB98C403C8F07154" box="[1099,1262,827,852]" italics="true" pageId="11" pageNumber="12">Pentapedilum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C154D050E15FFCAFB2EC403C9857154" box="[1277,1435,827,852]" class="Insecta" family="Chironomidae" genus="Uresipedilum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E15FFCAFB2EC403C9857154" box="[1277,1435,827,852]" italics="true" pageId="11" pageNumber="12">Uresipedilum</emphasis>
</taxonomicName>
and
<emphasis id="B961EA940E15FFCAFF19C458CD217179" box="[202,319,864,889]" italics="true" pageId="11" pageNumber="12">Probolum</emphasis>
. Most larvae cannot be distinguished from those of
<taxonomicName id="4C154D050E15FFCAFC7EC458C84F7179" box="[941,1105,864,889]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B961EA940E15FFCAFC7EC458C84F7179" box="[941,1105,864,889]" italics="true" pageId="11" pageNumber="12">Pentapedilum</emphasis>
</taxonomicName>
. Some larvae, however, are separable from other known larvae except
<taxonomicName id="4C154D050E15FFCAFD4DC4BEC897719F" authority="Oyewo et Jacobsen, 2007" authorityName="Oyewo et Jacobsen" authorityYear="2007" box="[670,1161,901,927]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="epleri" subGenus="Pe.">
<emphasis id="B961EA940E15FFCAFD4DC4BECF56719F" box="[670,840,902,927]" italics="true" pageId="11" pageNumber="12">P. (Pe.) epleri</emphasis>
Oyewo
<emphasis id="B961EA940E15FFCAFC7DC4BECFDD719F" box="[942,963,902,927]" italics="true" pageId="11" pageNumber="12">et</emphasis>
Jacobsen, 2007
</taxonomicName>
, by having first lateral teeth of mentum at least as high as median and/or second lateral teeth. From
<taxonomicName id="4C154D050E15FFCAFB97C493C8F071C4" box="[1092,1262,939,964]" class="Insecta" family="Chironomidae" genus="Pentapedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="epleri" subGenus="Pe.">
<emphasis id="B961EA940E15FFCAFB97C493C8F071C4" box="[1092,1262,939,964]" italics="true" pageId="11" pageNumber="12">P. (Pe.) epleri</emphasis>
</taxonomicName>
they differ by either having mandible with a dorsal tooth, mola without serrations or well developed Lauterborn organs.
</paragraph>
</subSubSection>
</treatment>
</document>