271 lines
26 KiB
XML
271 lines
26 KiB
XML
<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E35FE223FE73A02E5E88FD85" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Melanella indicaformis DeVries 2019, sp. nov." docType="treatment" docVersion="20" lastPageNumber="1556" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1555" updateTime="1636522412784" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DeVries, Thomas J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-12-17</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1533</mods:start>
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<mods:end>1584</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
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<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671248" ID-GBIF-Taxon="162146491" ID-Zenodo-Dep="3671248" LSID="urn:lsid:plazi:treatment:03BB2338E35FE223FE73A02E5E88FD85" httpUri="http://treatment.plazi.org/id/03BB2338E35FE223FE73A02E5E88FD85" lastPageId="24" lastPageNumber="1556" pageId="23" pageNumber="1555">
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<subSubSection box="[466,849,365,390]" pageId="23" pageNumber="1555" type="nomenclature">
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<paragraph blockId="23.[466,849,365,424]" box="[466,849,365,390]" pageId="23" pageNumber="1555">
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<heading box="[466,849,365,390]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="1555" reason="2">
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<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[466,748,365,390]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="indicaformis" status="sp. nov.">
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<emphasis bold="true" box="[466,748,365,390]" italics="true" pageId="23" pageNumber="1555">Melanella indicaformis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[757,849,365,390]" pageId="23" pageNumber="1555" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[574,741,399,424]" pageId="23" pageNumber="1555" type="description">
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<paragraph blockId="23.[466,849,365,424]" box="[574,741,399,424]" pageId="23" pageNumber="1555">
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<heading box="[574,741,399,424]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="1555" reason="5">
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(
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<figureCitation box="[582,682,399,424]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="23" pageNumber="1555">Figure 4</figureCitation>
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(h
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<emphasis box="[704,718,399,424]" italics="true" pageId="23" pageNumber="1555">–</emphasis>
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l))
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="1555" type="diagnosis">
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<paragraph blockId="23.[160,1155,473,568]" box="[160,276,473,499]" pageId="23" pageNumber="1555">
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<emphasis bold="true" box="[160,276,473,499]" italics="true" pageId="23" pageNumber="1555">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="23.[160,1155,473,568]" pageId="23" pageNumber="1555">Shell small, elongate. Whorls smooth, flat-sided. Aperture oval, elongate, without anterior or posterior canals. Columella smooth, umbilicus absent.</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="1555" type="description">
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<paragraph blockId="23.[160,1156,617,920]" box="[160,294,617,643]" pageId="23" pageNumber="1555">
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<emphasis bold="true" box="[160,294,617,643]" italics="true" pageId="23" pageNumber="1555">Description</emphasis>
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</paragraph>
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<paragraph blockId="23.[160,1156,617,920]" pageId="23" pageNumber="1555">Shell length less than 15 mm, elongate, spire angle about 15°. Protoconch conical, about two smooth whorls. Teleoconch with about five whorls; whorls smooth, glossy, flat-sided to slightly convex, sometimes weakly turreted, thick; spire sometimes bent. Sutures adpressed; short crimped sutural collar sometimes overlaps preceding whorl. No spiral or axial sculpture. Base smooth, evenly rounded, set apart from rest of whorl by weak or moderately strong angulation. Aperture oval, twice as long as wide. Parietal callus strong; columellar callus thin; basal lip thickened, continuous with base of columella. Outer lip moderately thick, without teeth. Umbilicus absent.</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="1555" type="discussion">
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<paragraph blockId="23.[160,1156,968,1653]" box="[160,262,968,994]" pageId="23" pageNumber="1555">
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<emphasis bold="true" box="[160,262,968,994]" italics="true" pageId="23" pageNumber="1555">Remarks</emphasis>
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</paragraph>
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<paragraph blockId="23.[160,1156,968,1653]" pageId="23" pageNumber="1555">
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Species of
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<taxonomicName authorityName="Bowdichi" authorityYear="1822" box="[278,381,1005,1029]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="genus">
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<emphasis box="[278,381,1005,1029]" italics="true" pageId="23" pageNumber="1555">Melanella</emphasis>
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</taxonomicName>
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have been reported from Paleocene rocks in
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<collectingCountry box="[883,1029,1005,1029]" name="New Zealand" pageId="23" pageNumber="1555">New Zealand</collectingCountry>
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(
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<bibRefCitation author="Finlay HJi Marwick J" pageId="23" pageNumber="1555" pagination="1 - 140" refId="ref23494" refString="Finlay HJi Marwick J. 1937. The Wangaloan and associated molluscan faunas of Kaitangata - green Island Subdivision. New Zealand Geol Survey Paleontological Bull. 15: 1 - 140." type="journal article" year="1937">Finlay and Marwick 1937</bibRefCitation>
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) and
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<collectingCountry box="[378,489,1040,1064]" name="Antarctica" pageId="23" pageNumber="1555">Antarctica</collectingCountry>
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(
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<bibRefCitation author="Stillwell JDi Zinsmeister WJi Olenik AE" box="[503,712,1040,1064]" pageId="23" pageNumber="1555" pagination="1 - 89" refId="ref26753" refString="Stillwell JDi Zinsmeister WJi Olenik AE. 2004. Early Paleocene mollusks of Antarctica: systematicsi paleoecology and paleobiogeographic significance. Bulletins Am Paleontol. 367: 1 - 89." type="journal article" year="2004">Stillwell et al. 2004</bibRefCitation>
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) and Eocene and Oligocene strata from North America (
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<bibRefCitation author="MacNeil FSi Dockery DT III." box="[332,628,1074,1098]" pageId="23" pageNumber="1555" pagination="1 - 415" refId="ref25028" refString="MacNeil FSi Dockery DT III. 1984. Lower Oligocene Gastropodai Scaphopodai and Cephalopoda of the Vicksburg Group in Mississippi. Mississippi Dept Nat Resour Bur Geol Bull. 124: 1 - 415." type="journal article" year="1984">MacNeil and Dockery 1984</bibRefCitation>
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;
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<bibRefCitation author="Garvie CL" box="[640,769,1074,1098]" pageId="23" pageNumber="1555" pagination="1 - 177" refId="ref23803" refString="Garvie CL. 1996. The molluscan macrofauna of the Reklaw Formationi Marquez Member (Eocene: Lower Claibornian) i in Texas. Bulletins Am Paleontol. 111 (352): 1 - 177." type="journal article" year="1996">Garvie 1996</bibRefCitation>
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) and
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<collectingCountry box="[833,977,1074,1098]" name="New Zealand" pageId="23" pageNumber="1555">New Zealand</collectingCountry>
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(
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<bibRefCitation author="Maxwell PA" box="[990,1143,1074,1098]" pageId="23" pageNumber="1555" pagination="1 - 280" refId="ref25206" refString="Maxwell PA. 1992. Eocene Mollusca from the vicinity of McCulloch ' s Bridgei Waihao Riveri South Canterburyi New Zealand: Paleoecology and systematics. New Zealand Geol Survey Paleontological Bull. 65: 1 - 280." type="journal article" year="1992">Maxwell 1992</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="23.[160,1156,968,1653]" pageId="23" pageNumber="1555">
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A species similar to
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[413,654,1109,1133]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="indicaformis">
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<emphasis box="[413,654,1109,1133]" italics="true" pageId="23" pageNumber="1555">Melanella indicaformis</emphasis>
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</taxonomicName>
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was provisionally assigned to
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<taxonomicName authorityName="Pictet & Campiche" authorityYear="1862" box="[993,1155,1109,1133]" class="Gastropoda" family="Pseudomelaniidae" genus="Pseudomelania" higherTaxonomySource="GBIF" kingdom="Animalia" order="[unassigned] Caenogastropoda" pageId="23" pageNumber="1555" phylum="Mollusca" rank="genus">
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<emphasis box="[993,1155,1109,1133]" italics="true" pageId="23" pageNumber="1555">Pseudomelania</emphasis>
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</taxonomicName>
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by
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<bibRefCitation author="Olsson AA" box="[195,346,1144,1168]" pageId="23" pageNumber="1555" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
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:
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<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[358,612,1144,1168]" class="Gastropoda" family="Pseudomelaniidae" genus="Pseudomelania" higherTaxonomySource="GBIF" kingdom="Animalia" order="[unassigned] Caenogastropoda" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="simplex">
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<emphasis box="[358,465,1144,1168]" italics="true" pageId="23" pageNumber="1555">P. simplex</emphasis>
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Olsson, 1944
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</taxonomicName>
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, found in upper Campanian beds of the
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<collectingCountry box="[1069,1156,1144,1168]" name="Haiti" pageId="23" pageNumber="1555">Tortuga</collectingCountry>
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Formation in the Sechura Basin of northern
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<collectingCountry box="[647,700,1178,1202]" name="Peru" pageId="23" pageNumber="1555">Peru</collectingCountry>
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. The
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<collectingCountry box="[758,844,1178,1202]" name="Haiti" pageId="23" pageNumber="1555">Tortuga</collectingCountry>
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Formation species is longer than the Caballas Formation species (length
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<quantity box="[650,728,1213,1237]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="1555" unit="mm" value="18.0">18 mm</quantity>
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vs
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<quantity box="[764,843,1213,1237]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="1555" unit="mm" value="12.0">12 mm</quantity>
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) and has nearly microscopic spiral threads, but the convexity of the spire, the glossy surface, and thickened inner lip are shared characters. A late Eocene species with a similar form,
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<taxonomicName authority="Woodringi 1973" authorityName="Woodringi" authorityYear="1973" box="[818,1155,1282,1306]" class="Gastropoda" family="Pseudomelaniidae" genus="Bayania" higherTaxonomySource="GBIF" kingdom="Animalia" order="[unassigned] Caenogastropoda" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="epelys">
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<emphasis box="[818,975,1282,1306]" italics="true" pageId="23" pageNumber="1555">Bayania epelys</emphasis>
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Woodring, 1973
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</taxonomicName>
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from the Canal Zone of Panama, differs from the Caballas Formation
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[915,1155,1317,1341]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="indicaformis">
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<emphasis box="[915,1155,1317,1341]" italics="true" pageId="23" pageNumber="1555">Melanella indicaformis</emphasis>
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</taxonomicName>
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by having faint spiral sculpture on the last whorl and a very shallow recurved siphonal notch. An early Oligocene species from the Talara Basin of northern Peru,
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<taxonomicName authority="Olssoni 1931" authorityName="Olssoni" authorityYear="1931" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="canoaensis">
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<emphasis italics="true" pageId="23" pageNumber="1555">Pachychilus canoaensis</emphasis>
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Olsson, 1931
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</taxonomicName>
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, from the estuarine and freshwater Bravo Grits of the Mancora Formation, also resembles
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<taxonomicName authorityName="DeVries" authorityYear="2019" box="[455,618,1456,1480]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="species" species="indicaformis">
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<emphasis box="[455,618,1456,1480]" italics="true" pageId="23" pageNumber="1555">M. indicaformis</emphasis>
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</taxonomicName>
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and may be referable to
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<taxonomicName authorityName="Bowdichi" authorityYear="1822" box="[898,1001,1456,1480]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="genus">
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<emphasis box="[898,1001,1456,1480]" italics="true" pageId="23" pageNumber="1555">Melanella</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="23.[160,1156,968,1653]" pageId="23" pageNumber="1555">
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Modern species of Eulimidae, including
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<taxonomicName authorityName="Bowdichi" authorityYear="1822" box="[658,763,1490,1514]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="genus">
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<emphasis box="[658,763,1490,1514]" italics="true" pageId="23" pageNumber="1555">Melanella</emphasis>
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</taxonomicName>
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, are noted for their parasitism of echinoderms (
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<bibRefCitation author="Waren A" box="[324,467,1525,1549]" pageId="23" pageNumber="1555" pagination="1 - 96" refId="ref27079" refString="Waren A. 1983. A generic revision of the family Eulimidae (Gastropodai Prosobranchia). J Molluscan Stud Suppl. 13: 1 - 96." type="journal article" year="1983">Warén 1983</bibRefCitation>
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). Echinoderms are noted for their intolerance of brackish conditions, although some extant brackish-water echinoderms do exist (
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<bibRefCitation author="Turner RLi Meyer CE" pageId="23" pageNumber="1555" pagination="249 - 256" refId="ref27020" refString="Turner RLi Meyer CE. 1980. Salinity tolerance of the brackish-water echinoderm Ophiophragmus filograneus (Ophiuroidea). Mar Ecol Progr Ser. 2: 249 - 256." type="book chapter" year="1980">Turner and Meyer 1980</bibRefCitation>
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). No echinoderms have been encountered in beds of the Cuenca Member, despite an abundance of
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<taxonomicName authorityName="Bowdichi" authorityYear="1822" box="[453,558,1629,1653]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="1555" phylum="Mollusca" rank="genus">
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<emphasis box="[453,558,1629,1653]" italics="true" pageId="23" pageNumber="1555">Melanella</emphasis>
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</taxonomicName>
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shells.
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</paragraph>
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</subSubSection>
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<subSection lastPageId="24" lastPageNumber="1556" pageId="23" pageNumber="1555" type="multiple">
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<subSubSection box="[160,1067,142,1727]" lastPageId="24" lastPageNumber="1556" pageId="23" pageNumber="1555" type="etymology">
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<paragraph blockId="23.[160,285,1701,1727]" box="[160,285,1701,1727]" pageId="23" pageNumber="1555">
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<emphasis bold="true" box="[160,285,1701,1727]" italics="true" pageId="23" pageNumber="1555">Etymology</emphasis>
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</paragraph>
|
||
<paragraph blockId="24.[160,1067,142,166]" box="[160,1067,142,166]" pageId="24" pageNumber="1556">
|
||
<emphasis box="[160,166,142,166]" italics="true" pageId="24" pageNumber="1556">‘</emphasis>
|
||
Indica
|
||
<emphasis box="[233,238,142,166]" italics="true" pageId="24" pageNumber="1556">’</emphasis>
|
||
, referencing the long-grained variety of rice that this species resembles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="1556" type="materials_examined">
|
||
<paragraph blockId="24.[160,1157,228,462]" box="[160,259,228,254]" pageId="24" pageNumber="1556">
|
||
<emphasis bold="true" box="[160,259,228,254]" italics="true" pageId="24" pageNumber="1556">Material</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="24.[160,1157,228,462]" pageId="24" pageNumber="1556">
|
||
<materialsCitation ID-GBIF-Occurrence="2572159715" box="[160,857,265,289]" collectionCode="UWBM" longitude="-4.1" pageId="24" pageNumber="1556" specimenCode="UWBM 107617" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode box="[160,330,265,289]" pageId="24" pageNumber="1556">UWBM 107617</specimenCode>
|
||
,
|
||
<typeStatus box="[342,445,265,289]" pageId="24" pageNumber="1556">holotype</typeStatus>
|
||
, B8769 (type locality), L 11.1, W 4.1
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2572159712" collectionCode="UWBM" longitude="-4.1" pageId="24" pageNumber="1556" specimenCode="UWBM 107618" specimenCount="1" typeStatus="paratype">
|
||
remainder are
|
||
<typeStatus box="[1040,1153,265,289]" pageId="24" pageNumber="1556">paratypes</typeStatus>
|
||
:
|
||
<specimenCode box="[160,329,299,323]" pageId="24" pageNumber="1556">UWBM 107618</specimenCode>
|
||
, B8772, L 10.6, W 4.1
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890717" box="[579,973,299,323]" collectionCode="UWBM" longitude="-3.6" pageId="24" pageNumber="1556" specimenCode="UWBM 107619" specimenCount="1">
|
||
<specimenCode box="[579,747,299,323]" pageId="24" pageNumber="1556">UWBM 107619</specimenCode>
|
||
, B8769, L 9.4, W 3.6
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890668" collectionCode="UWBM" longitude="-3.1" pageId="24" pageNumber="1556" specimenCode="UWBM 107620" specimenCount="1">
|
||
<specimenCode box="[984,1152,299,323]" pageId="24" pageNumber="1556">UWBM 107620</specimenCode>
|
||
, B8772, L 8.2, W 3.1
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890731" box="[385,779,334,358]" collectionCode="UWBM" longitude="-3.2" pageId="24" pageNumber="1556" specimenCode="UWBM 107621" specimenCount="1">
|
||
<specimenCode box="[385,554,334,358]" pageId="24" pageNumber="1556">UWBM 107621</specimenCode>
|
||
, B8772, L 8.7, W 3.2
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890744" collectionCode="UWBM" pageId="24" pageNumber="1556" specimenCode="UWBM 107622" specimenCount="1">
|
||
<specimenCode box="[790,958,334,358]" pageId="24" pageNumber="1556">UWBM 107622</specimenCode>
|
||
, B8769, L (8.4), W (4.7)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890743" box="[225,638,369,393]" collectionCode="UWBM" longitude="-3.3" pageId="24" pageNumber="1556" specimenCode="UWBM 107623" specimenCount="1">
|
||
<specimenCode box="[225,396,369,393]" pageId="24" pageNumber="1556">UWBM 107623</specimenCode>
|
||
, B8769, L 8.2, W 3.3
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2572159716" box="[652,1065,369,393]" collectionCode="UWBM" longitude="-2.8" pageId="24" pageNumber="1556" specimenCode="UWBM 107624" specimenCount="1">
|
||
<specimenCode box="[652,824,369,393]" pageId="24" pageNumber="1556">UWBM 107624</specimenCode>
|
||
, B8772, L 6.0, W 2.8
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890723" collectionCode="UWBM" longitude="-2.6" pageId="24" pageNumber="1556" specimenCode="UWBM 107625" specimenCount="1">
|
||
<specimenCode pageId="24" pageNumber="1556">UWBM 107625</specimenCode>
|
||
, B8772, L 5.0, W 2.6
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890739" box="[478,869,403,427]" collectionCode="UWBM" longitude="-2.4" pageId="24" pageNumber="1556" specimenCode="UWBM 107626" specimenCount="1">
|
||
<specimenCode box="[478,646,403,427]" pageId="24" pageNumber="1556">UWBM 107626</specimenCode>
|
||
, B8770, L 4.5, W 2.4
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2565890679" collectionCode="MUSM" longitude="-3.3" pageId="24" pageNumber="1556" specimenCode="INV 266" specimenCount="1">
|
||
<collectionCode box="[880,953,403,427]" pageId="24" pageNumber="1556">MUSM</collectionCode>
|
||
<specimenCode box="[960,1051,403,427]" pageId="24" pageNumber="1556">INV 266</specimenCode>
|
||
, B8772, L 8.4, W 3.3
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2572159713" box="[286,704,438,462]" collectionCode="MUSM" longitude="-3.4" pageId="24" pageNumber="1556" specimenCode="INV 267" specimenCount="1">
|
||
<collectionCode box="[286,359,438,462]" pageId="24" pageNumber="1556">MUSM</collectionCode>
|
||
<specimenCode box="[368,459,438,462]" pageId="24" pageNumber="1556">INV 267</specimenCode>
|
||
, B8769, L (9.0), W 3.4
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2572159714" box="[717,1121,438,462]" collectionCode="MUSM" longitude="-2.6" pageId="24" pageNumber="1556" specimenCode="INV 268" specimenCount="1">
|
||
<collectionCode box="[717,790,438,462]" pageId="24" pageNumber="1556">MUSM</collectionCode>
|
||
<specimenCode box="[799,890,438,462]" pageId="24" pageNumber="1556">INV 268</specimenCode>
|
||
, B8772, L 6.1, W 2.6
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="24.[160,1157,228,462]" box="[1120,1124,438,462]" pageId="24" pageNumber="1556">.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[160,1156,524,585]" pageId="24" pageNumber="1556" type="description">
|
||
<paragraph blockId="24.[160,1156,524,585]" box="[160,292,524,550]" pageId="24" pageNumber="1556">
|
||
<emphasis bold="true" box="[160,292,524,550]" italics="true" pageId="24" pageNumber="1556">Occurrence</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="24.[160,1156,524,585]" box="[160,1156,561,585]" pageId="24" pageNumber="1556">
|
||
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
|
||
<collectingCountry box="[1099,1153,561,585]" name="Peru" pageId="24" pageNumber="1556">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |