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<document id="6A08251437E308873DB010CF29FAC374" ID-DOI="10.5252/g2012n3a7" ID-ISSN="1638-9395" ID-Zenodo-Dep="5378120" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630284107944" checkinUser="marcus" docAuthor="Mohamed Abdelhamid, Marouf Abdel-Aty &amp; Moustafa Azab, Mahmoud" docDate="2012" docId="03B78039FFB7A705FD2CFC9EC34D01C7" docLanguage="en" docName="Geodiversitas.2012.34.3.575-615.pdf" docOrigin="Geodiversitas 34 (3)" docSource="http://dx.doi.org/10.5252/g2012n3a7" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Trochalosoma (Trochalosoma) gharamulensis Abdelhamid &amp; Azab 2003" docType="treatment" docVersion="4" lastPageNumber="595" masterDocId="FF8EF841FFA4A711FF96FFAFC105036A" masterDocTitle="Turonian-Santonian echinoids from Egypt" masterLastPageNumber="615" masterPageNumber="575" pageNumber="594" updateTime="1699213872600" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="7430ABF18822F83455A729712F664BE6">Turonian-Santonian echinoids from Egypt</mods:title>
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<mods:namePart id="E63A6158ABF85DF7AF4B67DEA1D3F3B4">Mohamed Abdelhamid, Marouf Abdel-Aty</mods:namePart>
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<paragraph id="8BA1312FFFB7A702FD2CFC9EC2E600E1" blockId="19.[698,1188,817,907]" pageId="19" pageNumber="594">
<heading id="D0E98643FFB7A702FD2CFC9EC2E600E1" centered="true" fontSize="11" level="2" pageId="19" pageNumber="594" reason="2">
<taxonomicName id="4C1E4AACFFB7A702FD2CFC9EC5470001" ID-CoL="596J8" authority="Abdelhamid &amp; Azab, 2003" authorityName="Abdelhamid &amp; Azab" authorityYear="2003" class="Echinoidea" family="Phymosomatidae" genus="Trochalosoma" kingdom="Animalia" order="Phymosomatoida" pageId="19" pageNumber="594" phylum="Echinodermata" rank="species" species="gharamulensis" subGenus="Trochalosoma">
<emphasis id="B96AED3DFFB7A702FD2CFC9EC2570026" bold="true" box="[698,850,817,844]" italics="true" pageId="19" pageNumber="594">Trochalosoma</emphasis>
(
<emphasis id="B96AED3DFFB7A702FCF4FC9EC2FF0026" bold="true" box="[866,1018,817,844]" italics="true" pageId="19" pageNumber="594">Trochalosoma</emphasis>
)
<emphasis id="B96AED3DFFB7A702FB9CFC9EC5A10026" bold="true" box="[1034,1188,817,844]" italics="true" pageId="19" pageNumber="594">gharamulensis</emphasis>
<bibRefCitation id="EF8F4CDEFFB7A702FC8AFCFEC5470001" author="ABDELHAMID M. A. M. &amp; AZAB M. M." box="[796,1090,849,875]" pageId="19" pageNumber="594" pagination="851 - 876" refId="ref17825" refString="ABDELHAMID M. A. M. &amp; AZAB M. M. 2003. - Aptian-Cenomanian echinoids from Egypt. Revue de Paleobiologie, Geneve 22 (2): 851 - 876." type="journal article" year="2003">Abdelhamid &amp; Azab, 2003</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13252DAAFFB7A702FC12FCDEC2DC00E1" box="[900,985,881,907]" captionStart="FIG" captionStartId="13.[113,124,1498,1515]" captionText="FIG. 8. — A, B, Phymosoma roachensis (Fourtau, 1914), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; C, D, Phymosoma thevestense (Péron &amp; Gauthier, 1879), Abu Roash, Coniacian-Santonian, Ostrea and Plicatula unit, REH4, adapical and side views; E, F, Rachiosoma irregulare Fourtau, 1921, Wadi Abu Qaada, Turonian, Wata Formation, QEH4, adapical and side views; G, Rachiosoma rectilineatum (Péron &amp; Gauthier, 1881),Wadi Dakhl, Coniacian-Santonian, Matulla Formation, DEH3, adapical view; H, I, Thylechinus (T.) sinaiensis n. sp., Wadi Abu Qaada, Turonian, Wata Formation, QEH3, (holotype) adapical and side views; J, K, Thylechinus (O.) quincuncialis Gregory, 1906, Wadi Abu Qaada, Turonian, Wata Formation, QEH2, adapical and side views; L, M, Goniopygus innesi Gauthier,1901,Abu Roash, Coniacian-Santonian,Ostrea and Plicatula unit,REH4, adapical view and a spine; N, Trochalosoma (T.) gharamulensis Abdelhamid &amp; Azab, 2003, Wadi Abu Qada, Turonian, Wata Formation, QEH1. Scale bars: A, B, D, 2 mm; C, H. I, M, 3.3 mm; E, 4 mm; F, G, 5 mm; J, 10 mm; K, 6.7 mm; L, 2.5 mm; N, 5 mm." pageId="19" pageNumber="594">Fig. 8N</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFB7A702FD33FC05C252009D" pageId="19" pageNumber="594" type="reference_group">
<paragraph id="8BA1312FFFB7A702FD33FC05C252009D" blockId="19.[675,1208,938,1015]" pageId="19" pageNumber="594">
<treatmentCitationGroup id="AB0E1601FFB7A702FD33FC05C252009D" pageId="19" pageNumber="594">
<treatmentCitation id="0ABF173EFFB7A702FD33FC05C24600B6" author="ABDELHAMID M. A. M. &amp; AZAB M. M." page="860" pageId="19" pageNumber="594" year="2003">
<taxonomicName id="4C1E4AACFFB7A702FD33FC05C24600B6" ID-CoL="596J8" authority="Abdelhamid &amp; Azab, 2003: 860" authorityName="Abdelhamid &amp; Azab" authorityPageNumber="860" authorityYear="2003" class="Echinoidea" family="Phymosomatidae" genus="Trochalosoma" kingdom="Animalia" order="Phymosomatoida" pageId="19" pageNumber="594" phylum="Echinodermata" rank="species" species="gharamulensis" subGenus="Trochalosoma">
<emphasis id="B96AED3DFFB7A702FD33FC05C21B00A8" box="[677,798,938,962]" italics="true" pageId="19" pageNumber="594">Trochalosoma</emphasis>
(
<emphasis id="B96AED3DFFB7A702FCBFFC05C2A700A8" box="[809,930,938,962]" italics="true" pageId="19" pageNumber="594">Trochalosoma</emphasis>
)
<emphasis id="B96AED3DFFB7A702FC38FC05C52E00A8" box="[942,1067,938,962]" italics="true" pageId="19" pageNumber="594">gharamulensis</emphasis>
<bibRefCitation id="EF8F4CDEFFB7A702FBB8FC05C24600B6" author="ABDELHAMID M. A. M. &amp; AZAB M. M." pageId="19" pageNumber="594" pagination="851 - 876" refId="ref17825" refString="ABDELHAMID M. A. M. &amp; AZAB M. M. 2003. - Aptian-Cenomanian echinoids from Egypt. Revue de Paleobiologie, Geneve 22 (2): 851 - 876." type="journal article" year="2003">Abdelhamid&amp; Azab, 2003: 860</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. III, figs B-D. —
<treatmentCitation id="0ABF173EFFB7A702FB9EFC6BC5B100B7" author="EL QOT G. M." box="[1032,1204,964,989]" page="138" pageId="19" pageNumber="594" year="2006">
<bibRefCitation id="EF8F4CDEFFB7A702FB9EFC6BC5B100B7" author="EL QOT G. M." box="[1032,1204,964,989]" pageId="19" pageNumber="594" refId="ref18478" refString="EL QOT G. M. 2006. - Late Cretaceous macrofossils from Sinai. Beringeria 36, 163 p, 34 pls." type="book" year="2006">El Qot 2006: 138</bibRefCitation>
</treatmentCitation>
, pl. 31, figs 11, 12.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFB7A702FD33FBA0C207071C" pageId="19" pageNumber="594" type="materials_examined">
<paragraph id="8BA1312FFFB7A702FD33FBA0C207071C" blockId="19.[677,1211,1039,1142]" pageId="19" pageNumber="594">
MATERIAL EXAMINED. — Three specimens from the Turonian of Wadi Abu Qaada (Wata Formation),
<collectionCode id="ED0FA9EAFFB7A702FB01FB86C3F40736" pageId="19" pageNumber="594">AS- UAQE</collectionCode>
70-72, Wadi Abu Qaada echinoid horizon 1 (
<collectionCode id="ED0FA9EAFFB7A702FD38FBF1C3E9071C" box="[686,748,1118,1142]" pageId="19" pageNumber="594">QEH</collectionCode>
1).
</paragraph>
</subSubSection>
<subSubSection id="C30462A4FFB7A702FD33FB20C2BC05C9" pageId="19" pageNumber="594" type="description">
<paragraph id="8BA1312FFFB7A702FD33FB20C23007CD" blockId="19.[675,1212,1167,1699]" box="[677,821,1167,1193]" pageId="19" pageNumber="594">DESCRIPTION</paragraph>
<paragraph id="8BA1312FFFB7A702FD33FB01C2BC05C9" blockId="19.[675,1212,1167,1699]" pageId="19" pageNumber="594">Outline pentagonal to round. Adapical surface flattened to feebly convex. Adoral surface not depressed around peristome. Ambulacral area wide, feebly swollen. Poriferous zones biserial and wide adapically constituting most of ambulacrum width near apical disc, narrow and uniserial at ambitus, wide and crowded near peristome forming oblique triads of pore pairs.Major ambulacral plates trigeminate near peristome, quadrigeminate at ambitus, polygeminate with five pore pairs near apical disc. Interambulacral area with two columns of imperforate, noncrenulated primary tubercles; beside two external columns of secondary, nonconfluent tubercles as large as the primary tubercles but don’t persist until peristome and apical disc. Gill slits conspicuously developed.</paragraph>
</subSubSection>
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<paragraph id="8BA1312FFFB0A705FF2AFF74C5740386" blockId="20.[188,1137,219,236]" box="[188,1137,219,236]" pageId="20" pageNumber="595">
TABLE 7. — Measurements (in mm) of
<taxonomicName id="4C1E4AACFFB0A705FE64FF74C2080386" authority="Gauthier, 1901" authorityName="Gauthier" authorityYear="1901" box="[498,781,219,236]" class="Echinoidea" family="Acropeltidae" genus="Goniopygus" kingdom="Animalia" order="Arbacioida" pageId="20" pageNumber="595" phylum="Echinodermata" rank="species" species="innesi">
<emphasis id="B96AED3DFFB0A705FE64FF74C3890386" box="[498,652,219,236]" italics="true" pageId="20" pageNumber="595">Goniopygus innesi</emphasis>
Gauthier, 1901
</taxonomicName>
. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BA1312FFFB0A705FEAEFE88C58E021C" pageId="20" pageNumber="595">
<table id="F91EC38FFFB058EEFFE5FE88C5A6021C" box="[115,1187,295,374]" gridcols="8" gridrows="3" pageId="20" pageNumber="595" rowsContinueFrom="19.[115,1176,295,374]" rowsContinueIn="21.[115,634,317,397]">
<tr id="352E336DFFB058EEFFE5FE88C5A60251" box="[115,1187,295,315]" gridrow="0" pageId="20" pageNumber="595" rowspan-0="1">
<th id="76FF5A11FFB058EEFE8EFE88C06D0251" box="[280,360,295,315]" gridcol="1" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FEAEFE88C04D0251" bold="true" box="[312,328,295,315]" pageId="20" pageNumber="595">D</emphasis>
</th>
<th id="76FF5A11FFB058EEFE0DFE88C0F30251" box="[411,502,295,315]" gridcol="2" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FE23FE88C0D90251" bold="true" box="[437,476,295,315]" pageId="20" pageNumber="595">H/D</emphasis>
</th>
<th id="76FF5A11FFB058EEFDB2FE88C3850251" box="[548,640,295,315]" gridcol="3" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FDA7FE88C3770251" bold="true" box="[561,626,295,315]" pageId="20" pageNumber="595">Wa/Wi</emphasis>
</th>
<th id="76FF5A11FFB058EEFD53FE88C3F50251" box="[709,752,295,315]" gridcol="4" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FD5BFE88C3EC0251" bold="true" box="[717,745,295,315]" pageId="20" pageNumber="595">Na</emphasis>
</th>
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<emphasis id="B96AED3DFFB0A705FCCFFE88C26B0251" bold="true" box="[857,878,295,315]" pageId="20" pageNumber="595">Ni</emphasis>
</th>
<th id="76FF5A11FFB058EEFC29FE88C51E0251" box="[959,1051,295,315]" gridcol="6" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FC42FE88C5000251" bold="true" box="[980,1029,295,315]" pageId="20" pageNumber="595">La/D</emphasis>
</th>
<th id="76FF5A11FFB058EEFBDEFE88C5A60251" box="[1096,1187,295,315]" gridcol="7" gridrow="0" pageId="20" pageNumber="595">
<emphasis id="B96AED3DFFB0A705FBCDFE88C5950251" bold="true" box="[1115,1168,295,315]" pageId="20" pageNumber="595">Dp/D</emphasis>
</th>
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<th id="76FF5A11FFB058EEFFE5FEE5C1B40234" box="[115,177,330,350]" gridcol="0" gridrow="1" pageId="20" pageNumber="595">Range</th>
<td id="76FF5A11FFB058EEFE8EFEE5C06D0234" box="[280,360,330,350]" gridcol="1" gridrow="1" pageId="20" pageNumber="595">6.6-14.2</td>
<td id="76FF5A11FFB058EEFE0DFEE5C0F30234" box="[411,502,330,350]" gridcol="2" gridrow="1" pageId="20" pageNumber="595">0.46-0.58</td>
<td id="76FF5A11FFB058EEFDB2FEE5C3850234" box="[548,640,330,350]" gridcol="3" gridrow="1" pageId="20" pageNumber="595">0.46-0.59</td>
<td id="76FF5A11FFB058EEFD53FEE5C3F50234" box="[709,752,330,350]" gridcol="4" gridrow="1" pageId="20" pageNumber="595">7-11</td>
<td id="76FF5A11FFB058EEFCC2FEE5C2710234" box="[852,884,330,350]" gridcol="5" gridrow="1" pageId="20" pageNumber="595">6-7</td>
<td id="76FF5A11FFB058EEFC29FEE5C51E0234" box="[959,1051,330,350]" gridcol="6" gridrow="1" pageId="20" pageNumber="595">0.41-0.59</td>
<td id="76FF5A11FFB058EEFBDEFEE5C5A60234" box="[1096,1187,330,350]" gridcol="7" gridrow="1" pageId="20" pageNumber="595">0.50-0.56</td>
</tr>
<tr id="352E336DFFB058EEFFE5FECDC5A6021C" box="[115,1187,354,374]" gridrow="2" pageId="20" pageNumber="595">
<th id="76FF5A11FFB058EEFFE5FECDC1B4021C" box="[115,177,354,374]" gridcol="0" gridrow="2" pageId="20" pageNumber="595">Mean</th>
<td id="76FF5A11FFB058EEFE8EFECDC06D021C" box="[280,360,354,374]" gridcol="1" gridrow="2" pageId="20" pageNumber="595">6.7</td>
<td id="76FF5A11FFB058EEFE0DFECDC0F3021C" box="[411,502,354,374]" gridcol="2" gridrow="2" pageId="20" pageNumber="595">0.52</td>
<td id="76FF5A11FFB058EEFDB2FECDC385021C" box="[548,640,354,374]" gridcol="3" gridrow="2" pageId="20" pageNumber="595">0.51</td>
<td id="76FF5A11FFB058EEFD53FECDC3F5021C" box="[709,752,354,374]" gridcol="4" gridrow="2" pageId="20" pageNumber="595">9</td>
<td id="76FF5A11FFB058EEFCC2FECDC271021C" box="[852,884,354,374]" gridcol="5" gridrow="2" pageId="20" pageNumber="595">6.5</td>
<td id="76FF5A11FFB058EEFC29FECDC51E021C" box="[959,1051,354,374]" gridcol="6" gridrow="2" pageId="20" pageNumber="595">0.5</td>
<td id="76FF5A11FFB058EEFBDEFECDC5A6021C" box="[1096,1187,354,374]" gridcol="7" gridrow="2" pageId="20" pageNumber="595">0.53</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30462A4FFB0A705FFE7FE19C34D01C7" pageId="20" pageNumber="595" type="discussion">
<paragraph id="8BA1312FFFB0A705FFE7FE19C3860125" blockId="20.[113,648,438,591]" pageId="20" pageNumber="595">
REMARKS
<bibRefCitation id="EF8F4CDEFFB0A705FFE7FE7AC0A80285" author="ABDELHAMID M. A. M. &amp; AZAB M. M." box="[113,429,469,495]" pageId="20" pageNumber="595" pagination="851 - 876" refId="ref17825" refString="ABDELHAMID M. A. M. &amp; AZAB M. M. 2003. - Aptian-Cenomanian echinoids from Egypt. Revue de Paleobiologie, Geneve 22 (2): 851 - 876." type="journal article" year="2003">Abdelhamid &amp; Azab (2003)</bibRefCitation>
erected this species from the Cenomanian of Gabal Gharamul, the Eastern Desert,
<collectingCountry id="F30971BFFFB0A705FE8BFDBAC0650145" box="[285,352,533,559]" name="Egypt" pageId="20" pageNumber="595">Egypt</collectingCountry>
. It is recorded here from the Turonian of Wadi Abu Qaada, west
<collectingRegion id="49DAFFCDFFB0A705FE61FD9BC3350124" box="[503,560,564,590]" country="Egypt" name="Shamal Sina'" pageId="20" pageNumber="595">Sinai</collectingRegion>
,
<collectingCountry id="F30971BFFFB0A705FDADFD9AC37B0125" box="[571,638,565,591]" name="Egypt" pageId="20" pageNumber="595">Egypt</collectingCountry>
.
</paragraph>
<paragraph id="8BA1312FFFB0A705FF39FD3CC34D01C7" blockId="20.[175,584,659,749]" box="[175,584,659,685]" pageId="20" pageNumber="595">
Order
<taxonomicName id="4C1E4AACFFB0A705FF6FFD3CC34D01C7" authority="Gregory, 1900" authorityName="Gregory" authorityYear="1900" box="[249,584,659,685]" class="Echinoidea" kingdom="Animalia" order="Arbacioida" pageId="20" pageNumber="595" phylum="Echinodermata" rank="order">ARBACIOIDA Gregory, 1900</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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