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<document id="BCD01715E37181F72617BBD843BDE1DA" ID-DOI="10.11646/zootaxa.5374.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10158722" ID-ZooBank="EEFF13BD-664A-4FD8-9FED-4E4B21A475E4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700469135046" checkinUser="plazi" docAuthor="Ahrens, Dirk, Lukic, Daniel, Pham, Phu, Li, Wei &amp; Liu, Wangang" docDate="2023" docId="03B73169BE62275ED9E40E26FCF2FC8C" docLanguage="en" docName="zootaxa.5374.4.1.pdf" docOrigin="Zootaxa 5374 (4)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetraserica phamanhi Pham &amp; Ahrens 2023, new species" docType="treatment" docVersion="2" lastPageNumber="470" masterDocId="FF8E4911BE73274DD9730A19FFFAFFD1" masterDocTitle="Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)" masterLastPageNumber="486" masterPageNumber="451" pageNumber="468" updateTime="1700469620343" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="33153B537504FC1A75468EE211960B0E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0D15FAF5DA73B1EF7B4F17148548C00">
<mods:title id="D039D706BE329B38788684A79E5C6F74">Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="B6EBA05E5B2D25EE4F33B7EA51B877FE" type="personal">
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<mods:namePart id="73775B1B0B0762B327EA449508B97E58">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="D9EA619EB488A944A12F6CA0B4CF5E83" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:affiliation id="DDC6DB33F804B368D32B86E08103B714">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="6C4B9017E8C58600EC3BA45EE0CCFBCE" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="08CD6BA6B786A691350CFBE1E88F760E" type="personal">
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<mods:namePart id="E8EEF7828FE6B61BFC1CE34EEC9027EA">Lukic, Daniel</mods:namePart>
<mods:affiliation id="B2AA1C0295C0E61C90830367681BD5F2">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; University of Salzburg, Salzburg, Austria</mods:affiliation>
</mods:name>
<mods:name id="83E62C382F68A34859EE7E93A9237168" type="personal">
<mods:role id="3A80FEB4F4B88BD708DEFA55B0056E20">
<mods:roleTerm id="B30E90FB5B5AE360BABE39CD3E8880AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23A32770C10FEB23A8D11D1F1615BB41">Pham, Phu</mods:namePart>
<mods:affiliation id="B99068F4E1FD1E908176937D7F84C494">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
</mods:name>
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<mods:role id="26BDBF70D90AFA562AD763683FC5F7FA">
<mods:roleTerm id="977DC2F0D3EB606DB1884994669D3422">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECABCB4EA90B0C737F44D7E07058D26C">Li, Wei</mods:namePart>
<mods:nameIdentifier id="6105AD693F6A0F3077D7B1676F280B32" type="ORCID">0009-0001-0195-7889</mods:nameIdentifier>
<mods:affiliation id="83E342B8E47EA84EFFEA94A2801E4662">Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liweiv @ stumail. nwu. edu. cn; https: // orcid. org / 0009 - 0001 - 0195 - 7889</mods:affiliation>
<mods:nameIdentifier id="752A64186EBAE5182369E25F952EDFAD" type="email">liweiv@stumail.nwu.edu.cn</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="19CBFCF48FA51D0DF58879A167905AFD">Liu, Wangang</mods:namePart>
<mods:nameIdentifier id="C9C0BD657C72C6CFC1FC0B557D41F7A2" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:affiliation id="81E9FF9F3AC6DFB6A98B4F4DBB1AC63E">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany &amp; Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. &amp; liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967</mods:affiliation>
<mods:nameIdentifier id="32EEFC4A15701E792D6E4F3438944C3B" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:typeOfResource id="4CB62CDA3B663E620F6EFE7B51CBFB31">text</mods:typeOfResource>
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<mods:title id="FFA3033F5F50D4BD53EA7B73AB836393">Zootaxa</mods:title>
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<mods:part id="DC75E647F71D9BCC743CA68C7B18547C">
<mods:date id="FE3DB415427652F9A30F76A39DCE83A3">2023</mods:date>
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<mods:number id="C1F42E8FE3DBF977F63C5293559AEE68">2023-11-20</mods:number>
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<mods:number id="9C15B4496C2DD8A4145B87B8E0E1CB73">5374</mods:number>
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<mods:url id="93390AC51D680B40E6E75E7E7D62B2B9">https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301</mods:url>
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<mods:classification id="AF0A3DB261658A78BB82B3EA42D5B610">journal article</mods:classification>
<mods:identifier id="39BCA103F0990AC31CEA917B74DA612A" type="DOI">10.11646/zootaxa.5374.4.1</mods:identifier>
<mods:identifier id="EAB5E33C25FE7ABCEADEE3825EC1405F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A6FD2708B3D0148A1BF730F7101FF619" type="Zenodo-Dep">10158722</mods:identifier>
<mods:identifier id="2F092878AC709B8E3EB0CD95DEA0508F" type="ZooBank">EEFF13BD-664A-4FD8-9FED-4E4B21A475E4</mods:identifier>
</mods:mods>
<treatment id="03B73169BE62275ED9E40E26FCF2FC8C" LSID="urn:lsid:plazi:treatment:03B73169BE62275ED9E40E26FCF2FC8C" httpUri="http://treatment.plazi.org/id/03B73169BE62275ED9E40E26FCF2FC8C" lastPageId="19" lastPageNumber="470" pageId="17" pageNumber="468">
<subSubSection id="C304D3F4BE62275CD9E40E26FCEAFB89" box="[151,784,1086,1113]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8BA1807FBE62275CD9E40E26FCEAFB89" blockId="17.[151,784,1086,1149]" box="[151,784,1086,1113]" pageId="17" pageNumber="468">
<heading id="D0E93713BE62275CD9E40E26FCEAFB89" bold="true" box="[151,784,1086,1113]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="B96A5C6DBE62275CD9E40E26FCEAFB89" bold="true" box="[151,784,1086,1113]" pageId="17" pageNumber="468">
<taxonomicName id="4C1EFBFCBE62275CD9E40E26FD89FB89" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[151,627,1086,1113]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="phamanhi" status="sp. nov.">
<emphasis id="B96A5C6DBE62275CD9E40E26FE5BFB88" bold="true" box="[151,417,1086,1113]" italics="true" pageId="17" pageNumber="468">Tetraserica phamanhi</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE62275CDBF30E26FCEAFB89" box="[640,784,1086,1113]" pageId="17" pageNumber="468" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE62275CD9E40E7AFEF5FBAC" box="[151,271,1123,1149]" pageId="17" pageNumber="468" type="description">
<paragraph id="8BA1807FBE62275CD9E40E7AFEF5FBAC" blockId="17.[151,784,1086,1149]" box="[151,271,1123,1149]" pageId="17" pageNumber="468">
<figureCitation id="13259CFABE62275CD9E40E7AFEF5FBAC" box="[151,271,1123,1149]" captionStart="FIGURE 7" captionStartId="18.[152,255,1468,1492]" captionTargetBox="[162,1434,192,1435]" captionTargetId="figure-23@18.[151,1436,181,1445]" captionTargetPageId="18" captionText="FIGURE 7. A–D: Tetraserica phamanhi Pham &amp; Ahrens, new species (holotype); E–H: T. semikontumensis Pham &amp; Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158739" httpUri="https://zenodo.org/record/10158739/files/figure.png" pageId="17" pageNumber="468">Fig. 7A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE62275CD9E40EB2FD89FA84" pageId="17" pageNumber="468" type="materials_examined">
<paragraph id="8BA1807FBE62275CD9E40EB2FD89FA84" blockId="17.[151,1437,1194,1977]" pageId="17" pageNumber="468">
<emphasis id="B96A5C6DBE62275CD9E40EB2FE45FB15" bold="true" box="[151,447,1194,1221]" pageId="17" pageNumber="468">Type material examined.</emphasis>
<materialsCitation id="3B768A22BE62275CD8B50EB2FDF2FB38" collectingDate="2016-04-27" collectingDateMax="2016-04-28" collectingDateMin="2016-04-27" collectionCode="IEBR" collectorName="H. V. Tru" country="Vietnam" location="Son La" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Son La" typeStatus="holotype">
<typeStatus id="54A53EDDBE62275CD8B50EB2FDC9FB14" box="[454,563,1195,1221]" pageId="17" pageNumber="468" type="holotype">Holotype</typeStatus>
: ♁ “[
<collectingCountry id="F309C0EFBE62275CDB190EB2FD2BFB14" box="[618,721,1195,1221]" name="Vietnam" pageId="17" pageNumber="468">Vietnam</collectingCountry>
]
<collectingRegion id="49DA4E9DBE62275CDBAC0EB2FCD5FB14" box="[735,815,1195,1221]" country="Vietnam" name="Son La" pageId="17" pageNumber="468">Son La</collectingRegion>
Pro., Thuan Chau Dis., Copia NR.;
<date id="FFA0A6BFBE62275CDDB60EB2FA93FB14" box="[1221,1385,1195,1221]" pageId="17" pageNumber="468" value="2016-04-27" valueMax="2016-04-28" valueMin="2016-04-27">
<collectingDate id="EFE45F57BE62275CDDB60EB2FA93FB14" box="[1221,1385,1195,1221]" pageId="17" pageNumber="468" value="2016-04-27" valueMax="2016-04-28" valueMin="2016-04-27">27–28.iv.2016</collectingDate>
</date>
;
<collectingMethod id="525FF868BE62275CDC070EB2FEFBFB38" pageId="17" pageNumber="468">UV light trap</collectingMethod>
;
<collectorName id="26EBE5A9BE62275CD87F0ED6FE97FB38" box="[268,365,1231,1257]" pageId="17" pageNumber="468">H.V. Tru</collectorName>
leg.” (
<collectionCode id="ED0F18BABE62275CD8C90ED6FDFAFB38" box="[442,512,1231,1257]" pageId="17" pageNumber="468">IEBR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B768A22BE62275CDB610ED6FDC0FADC" collectingDate="2016-05-01" collectionCode="IEBR" collectorName="H. V. Tru" country="Vietnam" location="Son La" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
<typeStatus id="54A53EDDBE62275CDB610ED6FD79FB38" box="[530,643,1231,1257]" pageId="17" pageNumber="468" type="paratype">Paratypes</typeStatus>
: 1 ♁ “[
<collectingCountry id="F309C0EFBE62275CDBA30ED6FCCDFB38" box="[720,823,1231,1257]" name="Vietnam" pageId="17" pageNumber="468">Vietnam</collectingCountry>
]
<collectingRegion id="49DA4E9DBE62275CDA360ED6FC6CFB38" box="[837,918,1231,1257]" country="Vietnam" name="Son La" pageId="17" pageNumber="468">Son La</collectingRegion>
Pro., Thuan Chau Dis., Copia NR.;
<date id="FFA0A6BFBE62275CDC5E0ED6FA62FB38" box="[1325,1432,1231,1257]" pageId="17" pageNumber="468" value="2016-05-01">
<collectingDate id="EFE45F57BE62275CDC5E0ED6FA62FB38" box="[1325,1432,1231,1257]" pageId="17" pageNumber="468" value="2016-05-01">1.v. 2016</collectingDate>
</date>
;
<collectingMethod id="525FF868BE62275CD9E40EEAFEC8FADC" box="[151,306,1267,1293]" pageId="17" pageNumber="468">UV light trap</collectingMethod>
;
<collectorName id="26EBE5A9BE62275CD84D0EEAFE65FADC" box="[318,415,1267,1293]" pageId="17" pageNumber="468">H.V. Tru</collectorName>
leg.” (
<collectionCode id="ED0F18BABE62275CD89E0EEAFDC8FADC" box="[493,562,1267,1293]" pageId="17" pageNumber="468">IEBR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B768A22BE62275CDB340EEAFDC9FAE0" collectingDate="2016-04-18" collectingDateMax="2016-05-02" collectingDateMin="2016-04-18" collectionCode="IEBR" collectorName="H. V. Tru" country="Vietnam" location="Son La" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
1 ♁ “[
<collectingCountry id="F309C0EFBE62275CDBF90EEAFD0BFADC" box="[650,753,1267,1293]" name="Vietnam" pageId="17" pageNumber="468">Vietnam</collectingCountry>
]
<collectingRegion id="49DA4E9DBE62275CDA730EEAFCABFADC" box="[768,849,1267,1293]" country="Vietnam" name="Son La" pageId="17" pageNumber="468">Son La</collectingRegion>
Pro., Thuan Chau Dis., Copia NR.;
<date id="FFA0A6BFBE62275CDD9D0EEAFA62FADC" box="[1262,1432,1267,1293]" pageId="17" pageNumber="468" value="2016-04-18" valueMax="2016-05-02" valueMin="2016-04-18">
<collectingDate id="EFE45F57BE62275CDD9D0EEAFA62FADC" box="[1262,1432,1267,1293]" pageId="17" pageNumber="468" value="2016-04-18" valueMax="2016-05-02" valueMin="2016-04-18">18.iv–2.v.2016</collectingDate>
</date>
;
<collectingMethod id="525FF868BE62275CD9E40F0EFEDCFAE0" box="[151,294,1303,1329]" pageId="17" pageNumber="468">Malaise trap</collectingMethod>
;
<collectorName id="26EBE5A9BE62275CD8410F0EFE63FAE0" box="[306,409,1303,1329]" pageId="17" pageNumber="468">H. V. Tru</collectorName>
leg.” (
<collectionCode id="ED0F18BABE62275CD8950F0EFDD1FAE0" box="[486,555,1303,1329]" pageId="17" pageNumber="468">IEBR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B768A22BE62275CDB4C0F0EFD97FA84" collectingDate="2017-05-14" collectionCode="IEBR, ZFMK" collectorName="P. V. Pham" country="Vietnam" location="Son La" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
2 ♁♁ “[
<collectingCountry id="F309C0EFBE62275CDBE50F0EFD07FAE0" box="[662,765,1303,1329]" name="Vietnam" pageId="17" pageNumber="468">Vietnam</collectingCountry>
]
<collectingRegion id="49DA4E9DBE62275CDA780F0EFCA6FAE0" box="[779,860,1303,1329]" country="Vietnam" name="Son La" pageId="17" pageNumber="468">Son La</collectingRegion>
Pro., Thuan Chau Dis., Copia NR.;
<date id="FFA0A6BFBE62275CDD870F0EFA9FFAE0" box="[1268,1381,1303,1329]" pageId="17" pageNumber="468" value="2017-05-14">
<collectingDate id="EFE45F57BE62275CDD870F0EFA9FFAE0" box="[1268,1381,1303,1329]" pageId="17" pageNumber="468" value="2017-05-14">14.v.2017</collectingDate>
</date>
;
<collectorName id="26EBE5A9BE62275CDC020F0EFF2DFA84" pageId="17" pageNumber="468">P.V. Pham</collectorName>
leg.,
<collectingMethod id="525FF868BE62275CD8660F22FE56FA84" box="[277,428,1339,1365]" pageId="17" pageNumber="468">UV light trap</collectingMethod>
” (
<collectionCode id="ED0F18BABE62275CD8B60F22FDFDFA84" box="[453,519,1339,1365]" pageId="17" pageNumber="468">IEBR</collectionCode>
,
<collectionCode id="ED0F18BABE62275CDB660F22FD99FA84" box="[533,611,1339,1365]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="468" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE62275ED9B40F47FCA1FE24" lastPageId="19" lastPageNumber="470" pageId="17" pageNumber="468" type="description">
<paragraph id="8BA1807FBE62275CD9B40F47FC9AFA4C" blockId="17.[151,1437,1194,1977]" pageId="17" pageNumber="468">
<emphasis id="B96A5C6DBE62275CD9B40F47FDE6FAA8" bold="true" box="[199,540,1374,1401]" pageId="17" pageNumber="468">
Description of the
<typeStatus id="54A53EDDBE62275CD8DD0F46FDEDFAA8" box="[430,535,1375,1401]" pageId="17" pageNumber="468" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CE62D9ABE62275CDBFA0F46FD1EFAA8" box="[649,740,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="17" pageNumber="468" unit="mm" value="8.1">8.1 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE62275CDACA0F46FBEEFAA8" box="[953,1044,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="17" pageNumber="468" unit="mm" value="6.7">6.7 mm</quantity>
, width:
<quantity id="4CE62D9ABE62275CDD050F47FB29FAA8" box="[1142,1235,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="17" pageNumber="468" unit="mm" value="5.1">5.1 mm</quantity>
. Body oval, dark reddish brown, labroclypeus and dorsal surface shiny, glabrous.
</paragraph>
<paragraph id="8BA1807FBE62275CD9B40FBEFD49F948" blockId="17.[151,1437,1194,1977]" pageId="17" pageNumber="468">Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex, anterior angles very strongly obtuse; anterior margin emarginate medially; lateral margin and ocular canthus producing a distinct blunt angle, margins moderately reflexed; surface medially convex, shiny, moderately and densely punctate, without setae; frontoclypeal suture moderately impressed and moderately bent medially; smooth area in front of eye approximately 3 times as wide as long; ocular canthus moderate and triangular(1/3 of ocular diameter), weakly punctate, with one terminal seta. Frons weakly shiny and smooth, with only a few erect setae beside eyes. Eyes small, ratio of diameter/ interocular width: 0.57.</paragraph>
<paragraph id="8BA1807FBE62275CD9B40CBAFE13F8D4" blockId="17.[151,1437,1194,1977]" pageId="17" pageNumber="468">
Antenna yellowish brown, with ten antennomeres, club in male with four antennomeres, little longer than remaining antennomeres combined, first joint of club slightly shorter than club.
<collectingRegion id="49DA4E9DBE62275CDD630CDEFB95F930" box="[1040,1135,1735,1761]" country="Myanmar" name="Chin" pageId="17" pageNumber="468">Mentum</collectingRegion>
anteriorly weakly elevated and weakly convex in middle.
</paragraph>
<paragraph id="8BA1807FBE62275CD9B40D16FEF6F868" blockId="17.[151,1437,1194,1977]" pageId="17" pageNumber="468">Pronotum transverse, widest just before base, lateral margins moderately convex and evenly convergent anteriorly, anterior angles distinctly produced, posterior angles strongly convex. Anterior margin slightly convex medially, with a fine, complete marginal line. Surface densely and finely punctate, with minute setae in punctures; lateral and anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, with fine and dense punctures.</paragraph>
<caption id="DF61D0F7BE61275FD9EB0FA5FD21F9CC" ID-DOI="http://doi.org/10.5281/zenodo.10158739" ID-Zenodo-Dep="10158739" httpUri="https://zenodo.org/record/10158739/files/figure.png" pageId="18" pageNumber="469" startId="18.[152,255,1468,1492]" targetBox="[162,1434,192,1435]" targetPageId="18" targetType="figure">
<paragraph id="8BA1807FBE61275FD9EB0FA5FD21F9CC" blockId="18.[151,1436,1468,1565]" pageId="18" pageNumber="469">
<emphasis id="B96A5C6DBE61275FD9EB0FA5FEADFA05" bold="true" box="[152,343,1468,1493]" pageId="18" pageNumber="469">FIGURE 7. A–D:</emphasis>
<taxonomicName id="4C1EFBFCBE61275FD82D0FA4FD1DFA04" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[350,743,1469,1493]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="phamanhi" status="sp. nov.">
<emphasis id="B96A5C6DBE61275FD82D0FA4FDC6FA04" box="[350,572,1469,1493]" italics="true" pageId="18" pageNumber="469">Tetraserica phamanhi</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE61275FDB820FA4FC91FA04" box="[753,875,1469,1493]" pageId="18" pageNumber="469" rank="species">new species</taxonomicNameLabel>
(holotype);
<emphasis id="B96A5C6DBE61275FDA9A0FA4FBD8FA04" bold="true" box="[1001,1058,1469,1493]" pageId="18" pageNumber="469">E–H:</emphasis>
<taxonomicName id="4C1EFBFCBE61275FDD5A0FA4FA63FA04" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[1065,1433,1469,1493]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="semikontumensis" status="sp. nov.">
<emphasis id="B96A5C6DBE61275FDD5A0FA4FB14FA04" box="[1065,1262,1469,1493]" italics="true" pageId="18" pageNumber="469">T. semikontumensis</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE61275FD9E40FF8FEEBFA28" box="[151,273,1505,1529]" pageId="18" pageNumber="469" rank="species">new species</taxonomicNameLabel>
(holotype).
<emphasis id="B96A5C6DBE61275FD8FE0FF9FE3CFA28" bold="true" box="[397,454,1504,1529]" pageId="18" pageNumber="469">A, E:</emphasis>
aedeagus, left side lateral view;
<emphasis id="B96A5C6DBE61275FDA670FF8FCB1FA28" bold="true" box="[788,843,1505,1529]" pageId="18" pageNumber="469">B, F:</emphasis>
parameres, dorsal view;
<emphasis id="B96A5C6DBE61275FDD390FF9FB7FFA28" bold="true" box="[1098,1157,1504,1529]" pageId="18" pageNumber="469">C, G:</emphasis>
aedeagus, right side lateral view;
<emphasis id="B96A5C6DBE61275FD9A50C1CFEEBF9CC" bold="true" box="[214,273,1541,1565]" pageId="18" pageNumber="469">D, H:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BA1807FBE61275FD9B40C76FDDCF924" blockId="18.[151,1436,1647,2033]" pageId="18" pageNumber="469">Elytra oblong, widest at posterior third, striae moderately impressed, finely and densely punctate, intervals moderately convex, with fine, dense punctures, with some erect, short setae, punctures with minute setae; epipleural edge robust, ending at convex external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of short microtrichomes.</paragraph>
<paragraph id="8BA1807FBE61275FD9B40CE6FEC2F81C" blockId="18.[151,1436,1647,2033]" pageId="18" pageNumber="469">Ventral surface dull, reddish brown, with large and dense punctures, sparsely setose; metasternum sparsely covered with fine, short, or minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.6. Pygidium moderately convex and dull, coarsely densely punctate, without smooth midline, punctures with fine setae along apical margin.</paragraph>
<paragraph id="8BA1807FBE61275ED9B40DCEFB44FE00" blockId="18.[151,1436,1647,2033]" lastBlockId="19.[151,1437,151,861]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">Legs moderately wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur wide, moderately shiny, its anterior margin acute, lacking an adjacent serrated line; ventral posterior margin straight, smooth and moderately widened in apical half; dorsal posterior margin smooth, without short setae basally. Metatibia short, widest at middle; ratio width/ length: 1/ 2.9; dorsal margin weakly longitudinally convex, with two groups of spines, basal one at first third, apical one at two thirds of metatibial length, basally with a few short single setae; lateral face weakly finely punctate; ventral margin finely serrate, with four robust equidistant setae; medial face smooth; apex interiorly near tarsal articulation moderately concave. Tarsomeres in distal half with fine, dense setae ventrally, dorsally smooth; metatarsomeres with a strong serrated ridge and glabrous ventrally, first metatarsomere as long as following two tarsomeres combined, distinctly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BA1807FBE60275ED9B40BC2FCA1FE24" blockId="19.[151,1437,151,861]" box="[199,859,475,501]" pageId="19" pageNumber="470">
Aedeagus.
<figureCitation id="13259CFABE60275ED8370BC2FE47FE24" box="[324,445,475,501]" captionStart="FIGURE 7" captionStartId="18.[152,255,1468,1492]" captionTargetBox="[162,1434,192,1435]" captionTargetId="figure-23@18.[151,1436,181,1445]" captionTargetPageId="18" captionText="FIGURE 7. A–D: Tetraserica phamanhi Pham &amp; Ahrens, new species (holotype); E–H: T. semikontumensis Pham &amp; Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158739" httpUri="https://zenodo.org/record/10158739/files/figure.png" pageId="19" pageNumber="470">Fig. 7A–C</figureCitation>
. Habitus:
<figureCitation id="13259CFABE60275EDB5D0BC2FD7DFE24" box="[558,647,475,501]" captionStart="FIGURE 7" captionStartId="18.[152,255,1468,1492]" captionTargetBox="[162,1434,192,1435]" captionTargetId="figure-23@18.[151,1436,181,1445]" captionTargetPageId="18" captionText="FIGURE 7. A–D: Tetraserica phamanhi Pham &amp; Ahrens, new species (holotype); E–H: T. semikontumensis Pham &amp; Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158739" httpUri="https://zenodo.org/record/10158739/files/figure.png" pageId="19" pageNumber="470">Fig. 7D</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE60275ED9B40BE7FDBDFD1C" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="8BA1807FBE60275ED9B40BE7FDBDFD1C" blockId="19.[151,1437,151,861]" pageId="19" pageNumber="470">
<emphasis id="B96A5C6DBE60275ED9B40BE7FEB8FDC9" bold="true" box="[199,322,510,536]" pageId="19" pageNumber="470">Diagnosis.</emphasis>
<taxonomicName id="4C1EFBFCBE60275ED83B0BE6FD06FDC8" authority="Pham &amp; Ahrens" authorityName="Pham &amp; Ahrens" authorityYear="2023" box="[328,764,511,537]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="phamanhi" status="sp. nov.">
<emphasis id="B96A5C6DBE60275ED83B0BE6FDC4FDC8" box="[328,574,511,537]" italics="true" pageId="19" pageNumber="470">Tetraserica phamanhi</emphasis>
Pham &amp; Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE60275EDA740BE6FC77FDC8" box="[775,909,511,537]" pageId="19" pageNumber="470" rank="species">new species</taxonomicNameLabel>
is in the shape of aedeagus very similar to that of
<taxonomicName id="4C1EFBFCBE60275ED9CB083AFCE2FDEC" authority="Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014" authorityName="Liu, Fabrizi, Bai, Yang &amp; Ahrens" authorityYear="2014" box="[184,792,547,573]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="mengeana">
<emphasis id="B96A5C6DBE60275ED9CB083AFEB2FDED" box="[184,328,547,572]" italics="true" pageId="19" pageNumber="470">T. mengeana</emphasis>
<bibRefCitation id="EF8FFD8EBE60275ED821083AFCE2FDEC" author="Liu, W. &amp; Fabrizi, S. &amp; Bai, M. &amp; Yang, X. - K. &amp; Ahrens, D." box="[338,792,547,573]" pageId="19" pageNumber="470" pagination="83 - 121" refId="ref44587" refString="Liu, W., Fabrizi, S., Bai, M., Yang, X. - K. &amp; Ahrens, D. (2014) A taxonomic review of the Tetraserica Ahrens, 2004 species of China (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 448, 83 - 121. https: // doi. org / 10.3897 / zookeys. 443.8429" type="journal article" year="2014">Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1EFBFCBE60275EDA21083AFA6DFDEC" authority="Fabrizi, Dalstein &amp; Ahrens, 2019" authorityName="Fabrizi, Dalstein &amp; Ahrens" authorityYear="2019" box="[850,1431,547,573]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="allomengeana">
<emphasis id="B96A5C6DBE60275EDA21083AFBF5FDEC" box="[850,1039,547,573]" italics="true" pageId="19" pageNumber="470">T. allomengeana</emphasis>
<bibRefCitation id="EF8FFD8EBE60275EDD6B083AFA6DFDEC" author="Fabrizi, S. &amp; Dalstein, V. &amp; Ahrens, D." box="[1048,1431,547,573]" pageId="19" pageNumber="470" pagination="1 - 155" refId="ref44465" refString="Fabrizi, S., Dalstein, V. &amp; Ahrens, D. (2019) A monograph on the genus Tetraserica from the Indochinese region (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 837, 1 - 155. https: // doi. org / 10.3897 / zookeys. 837.32057" type="journal article" year="2019">Fabrizi, Dalstein &amp; Ahrens, 2019</bibRefCitation>
</taxonomicName>
; the new species differs by the presence of a convex basal lobe at the base of the right paramere which is subequal in length the apical portion (distal to paramere’s bent); in the latter two species this basal portion is only slightly elevated and half as high as the distal portion of the paramere; additionally, in the new species the left paramere also lacks the blunt tooth on dorsal margin.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE60275ED9B408CFFAC1FD20" box="[199,1339,726,753]" pageId="19" pageNumber="470" type="description">
<paragraph id="8BA1807FBE60275ED9B408CFFAC1FD20" blockId="19.[151,1437,151,861]" box="[199,1339,726,753]" pageId="19" pageNumber="470">
<emphasis id="B96A5C6DBE60275ED9B408CFFEC5FD21" bold="true" box="[199,319,726,752]" pageId="19" pageNumber="470">Variation.</emphasis>
Length of body: 8.6–9.0 mm; length of elytra:
<quantity id="4CE62D9ABE60275EDA2308CEFC20FD20" box="[848,986,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="19" pageNumber="470" unit="mm" value="6.75" valueMax="6.8" valueMin="6.7">6.7–6.8 mm</quantity>
; maximum width:
<quantity id="4CE62D9ABE60275EDDDE08CFFACDFD20" box="[1197,1335,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="19" pageNumber="470" unit="mm" value="5.3" valueMax="5.4" valueMin="5.2">5.2–5.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE60275ED9B408E2FCF2FC8C" pageId="19" pageNumber="470" type="etymology">
<paragraph id="8BA1807FBE60275ED9B408E2FCF2FC8C" blockId="19.[151,1437,151,861]" pageId="19" pageNumber="470">
<emphasis id="B96A5C6DBE60275ED9B408E2FEB4FCC4" bold="true" box="[199,334,763,789]" pageId="19" pageNumber="470">Etymology.</emphasis>
This species is named after Associate Professor Pham Van Anh (Department of Environment, University of Science, VNU
<collectingRegion id="49DA4E9DBE60275ED8930906FDDCFCE8" box="[480,550,799,825]" country="Vietnam" name="Ha Noi" pageId="19" pageNumber="470">Hanoi</collectingRegion>
) who has arranged the expeditions in 2016 and 2017 which collected the
<typeStatus id="54A53EDDBE60275EDC1F0906FA66FCE8" box="[1388,1436,799,825]" pageId="19" pageNumber="470">type</typeStatus>
material of this species (noun in genitive case singular).
</paragraph>
</subSubSection>
</treatment>
</document>
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