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<document ID-DOI="http://doi.org/10.5281/zenodo.3792222" ID-GBIF-Dataset="6bed073a-175c-42e6-bb99-645c99d55b4f" ID-GBIF-Taxon="163750554" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792222" ID-ZooBank="C631A347-306E-4773-84A4-E4712329186B" LSID="urn:lsid:zoobank.org:act:AD950C98-9B09-40AC-85FE-04154475586F" checkinTime="1587984202003" checkinUser="plazi" docAuthor="Miller, Jeremy, Griswold, Charles &amp; Yin, Chang" docDate="2009" docId="03ACB915FFA2742AFF4E45FBFC1BFEBF" docLanguage="en" docName="ZK_article_1993.pdf" docOrigin="ZooKeys 11 (11)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Crassignatha pianma Miller, Griswold &amp; Yin 2009, sp. n." docType="treatment" docUuid="AD950C98-9B09-40AC-85FE-04154475586F" docUuidSource="ZooBank" docVersion="8" lastPageId="62" lastPageNumber="71" masterDocId="FF95C16DFF9F7414FFC2455DFFF1FFB8" masterDocTitle="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers" masterLastPageNumber="195" masterPageNumber="9" pageId="61" pageNumber="70" updateTime="1635491590422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
<mods:affiliation>California Academy of Sciences ,, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Chang</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University ,, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.11.160</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bed073a-175c-42e6-bb99-645c99d55b4f</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576459</mods:identifier>
<mods:identifier type="ZooBank">C631A347-306E-4773-84A4-E4712329186B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792222" ID-GBIF-Taxon="163750554" ID-Zenodo-Dep="3792222" LSID="urn:lsid:zoobank.org:act:AD950C98-9B09-40AC-85FE-04154475586F" httpUri="http://treatment.plazi.org/id/03ACB915FFA2742AFF4E45FBFC1BFEBF" lastPageId="62" lastPageNumber="71" pageId="61" pageNumber="70">
<subSubSection box="[140,756,166,193]" pageId="61" pageNumber="70" type="nomenclature">
<paragraph blockId="61.[140,962,166,264]" box="[140,756,166,193]" pageId="61" pageNumber="70">
<heading box="[140,756,166,193]" fontSize="11" level="2" pageId="61" pageNumber="70" reason="3">
<emphasis bold="true" box="[140,756,166,193]" pageId="61" pageNumber="70">
<taxonomicName authority="Miller, Griswold &amp; Yin" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[140,679,166,193]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="70" phylum="Arthropoda" rank="species" species="pianma" status="sp. n.">
<emphasis bold="true" box="[140,394,166,193]" italics="true" pageId="61" pageNumber="70">
<emphasis box="[140,295,166,193]" inLineHeading="true" pageId="61" pageNumber="70" reason="1">Crassignatha</emphasis>
pianma
</emphasis>
Miller, Griswold &amp; Yin
</taxonomicName>
,
<taxonomicNameLabel box="[691,756,167,193]" pageId="61" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="70" type="description">
<paragraph blockId="61.[140,962,166,264]" box="[140,962,202,229]" pageId="61" pageNumber="70">
<uri box="[140,962,202,229]" pageId="61" pageNumber="70">
urn:lsid:zoobank.org:act:
<uuid box="[416,962,202,229]" pageId="61" pageNumber="70">AD950C98-9B09-40AC-85FE-04154475586F</uuid>
</uri>
</paragraph>
<paragraph blockId="61.[140,962,166,264]" box="[140,772,237,264]" pageId="61" pageNumber="70">
<figureCitation box="[140,234,237,263]" captionStart="Figure 67" captionStartId="139.[140,212,960,982]" captionTargetBox="[140,1108,165,942]" captionTargetId="figure@139.[140,1109,164,940]" captionTargetPageId="139" captionText="Figure 67. A-D, Patu jidanweishi sp. n. from Chang Yan He; E, F, Patu qiqi from QiQi He; G, H; Patu xiaoxiao sp. n. from Chang Yan He; I, Crassignatha pianma sp. n. from Chang Yan He. A, E, G, I, epigynum, ventral view; B, epigynum, lateral view; C, F, H, cleared vulva, dorsal view; D, eggs." figureDoi="http://doi.org/10.5281/zenodo.3768876" httpUri="https://zenodo.org/record/3768876/files/figure.png" pageId="61" pageNumber="70">Figs 67I</figureCitation>
,
<figureCitation box="[246,277,237,263]" captionStart="Figure 72" captionStartId="144.[140,212,1112,1134]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@144.[140,1109,164,1092]" captionTargetPageId="144" captionText="Figure 72. Webs of symphytognathid spiders. A, Patu jidanweishi sp. n., female from Xiao Hei Shan Nature Reserve; B, Patu jidanweishi sp. n., male from Pianma; C, D, Crassignatha pianma sp. n., female from Chang Yan He; E, Crassignatha yinzhi sp. n., female from Xiao Hei Shan Nature Reserve." figureDoi="http://doi.org/10.5281/zenodo.3768884" httpUri="https://zenodo.org/record/3768884/files/figure.png" pageId="61" pageNumber="70">72</figureCitation>
C-D,
<figureCitation box="[343,414,237,264]" captionStart-0="Figure 74" captionStart-1="Figure 75" captionStartId-0="146.[140,212,1125,1147]" captionStartId-1="147.[140,212,1622,1644]" captionTargetBox-0="[163,1101,188,1093]" captionTargetBox-1="[180,1073,174,1590]" captionTargetId-0="figure@146.[140,1109,164,1105]" captionTargetId-1="figure@147.[168,1081,164,1602]" captionTargetPageId-0="146" captionTargetPageId-1="147" captionText-0="Figure 74. Crassignatha pianma sp. n. from Chang Yan He. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." captionText-1="Figure 75. Crassignatha pianma sp. n. from Chang Yan He, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." figureDoi-0="http://doi.org/10.5281/zenodo.3768888" figureDoi-1="http://doi.org/10.5281/zenodo.3768890" httpUri-0="https://zenodo.org/record/3768888/files/figure.png" httpUri-1="https://zenodo.org/record/3768890/files/figure.png" pageId="61" pageNumber="70">74-75</figureCitation>
,
<figureCitation box="[425,454,237,263]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">76</figureCitation>
A-B,
<figureCitation box="[515,547,238,264]" captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="61" pageNumber="70">77</figureCitation>
,
<figureCitation box="[559,588,237,263]" captionStart="Figure 78" captionStartId="150.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@150.[140,1108,164,1186]" captionTargetPageId="150" captionText="Figure 78. SEM. A-D, Crassignatha pianma sp. n. from Chang Yan He; E, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; F, Crassignatha quanqu sp. n. from Bang Bie village. A, female chelicera removed from prosoma, note single asymetrically bifid tooth; B, labrum, anterior, female; C, labrum, lateral, female; D, male metatarsus-tarsus joint, arrow indicates tarsal organ; E, epiandrous region; F, prosoma, lateral, male. Arrows in B and C indicate maximum extension of labrum." figureDoi="http://doi.org/10.5281/zenodo.3768896" httpUri="https://zenodo.org/record/3768896/files/figure.png" pageId="61" pageNumber="70">78</figureCitation>
A-D,
<figureCitation box="[653,681,237,263]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79</figureCitation>
A-B,
<figureCitation box="[743,772,237,263]" captionStart="Figure 98" captionStartId="169.[140,212,1410,1432]" captionTargetBox="[140,1108,165,1390]" captionTargetId="figure@169.[140,1109,164,1390]" captionTargetPageId="169" captionText="Figure 98. Distribution map, inset map shows study area. White: Simaoa maku sp. n.; blue: Simaoa bianjing sp. n.; red: Gaoligonga changya sp. n.; yellow:Gaoligonga zhusun sp. n.; orange: Crassignatha pianma sp. n.; purple: Crassignatha yinzhi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3768929" httpUri="https://zenodo.org/record/3768929/files/figure.png" pageId="61" pageNumber="70">98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="70" type="materials_examined">
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
<emphasis bold="true" box="[140,381,307,333]" pageId="61" pageNumber="70">
<emphasis box="[140,241,307,333]" inLineHeading="true" pageId="61" pageNumber="70" reason="1">Material</emphasis>
Examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608939458" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="HNU" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[393,504,308,334]" pageId="61" pageNumber="70">Holotype</typeStatus>
:
<collectingCountry box="[520,619,308,334]" name="China" pageId="61" pageNumber="70">CHINA</collectingCountry>
:
<collectingRegion box="[634,726,308,334]" country="China" name="Yunnan" pageId="61" pageNumber="70">Yunnan</collectingRegion>
:
<collectingCounty box="[741,875,308,334]" pageId="61" pageNumber="70">Lushui Co.</collectingCounty>
,
<collectingMunicipality box="[893,1103,308,334]" pageId="61" pageNumber="70">Pianma Township</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03ACB915FFA2742AFF4E45FBFC1BFEBF:8EDA5ED8FFA27429FF4E440AFEC7FEC9" box="[140,310,343,369]" country="China" county="Lushui Co." latitude="25.99363" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" name="Chang Yan He" pageId="61" pageNumber="70" stateProvince="Yunnan">Chang Yan He</location>
,
<locationDeviation box="[322,557,343,369]" location="Pianma" pageId="61" pageNumber="70">
<quantity box="[322,404,343,369]" metricMagnitude="3" metricUnit="m" metricValue="9.3" pageId="61" pageNumber="70" unit="km" value="9.3">9.3 km</quantity>
ESE Pianma
</locationDeviation>
,
<geoCoordinate box="[568,713,343,370]" degrees="25.99363" direction="north" orientation="latitude" pageId="61" pageNumber="70" precision="1" value="25.99363">25.99363°N</geoCoordinate>
,
<geoCoordinate box="[725,864,343,370]" degrees="98.66651" direction="east" orientation="longitude" pageId="61" pageNumber="70" precision="1" value="98.66651">98.66651°E</geoCoordinate>
,
<quantity box="[876,969,343,369]" metricMagnitude="3" metricUnit="m" metricValue="2.4699999999999998" pageId="61" pageNumber="70" unit="m" value="2470.0">
<elevation box="[876,969,343,369]" metricMagnitude="3" metricUnit="m" metricValue="2.4699999999999998" pageId="61" pageNumber="70" unit="m" value="2470.0">2470 m</elevation>
</quantity>
,
<date pageId="61" pageNumber="70" value="2005-05-13" valueMax="2005-05-15" valueMin="2005-05-13">
<collectingDate pageId="61" pageNumber="70" value="2005-05-13" valueMax="2005-05-15" valueMin="2005-05-13">13-15 May 2005</collectingDate>
</date>
, mixed broadleaf deciduous and evergreen forest, dusting small webs near ground in forest understory,
<collectorName box="[374,517,413,439]" pageId="61" pageNumber="70">C. Griswold</collectorName>
, CGY104 (CASENT 9022600,
<collectionCode box="[885,957,414,440]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="61" pageNumber="70">HNU</collectionCode>
), 1♁.
</materialsCitation>
</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
<materialsCitation ID-GBIF-Occurrence="2608939459" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="CAS" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCode="PV 0756-0758" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus box="[185,295,449,475]" pageId="61" pageNumber="70">Paratypes</typeStatus>
: [same data as holotype] (CASENT 9029339,
<specimenCode box="[847,1023,448,475]" collectionCode="PV" pageId="61" pageNumber="70">PV 0756-0758</specimenCode>
,
<collectionCode box="[1038,1095,449,475]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="70" type="Museum">CAS</collectionCode>
),
<specimenCount box="[140,170,484,510]" pageId="61" pageNumber="70" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939455" box="[193,1004,483,510]" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="CAS" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCode="PV 0734-0735" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9029340,
<specimenCode box="[722,896,483,510]" collectionCode="PV" pageId="61" pageNumber="70">PV 0734-0735</specimenCode>
,
<collectionCode box="[910,967,484,510]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="70" type="Museum">CAS</collectionCode>
), 1
</materialsCitation>
♁;
<materialsCitation ID-GBIF-Occurrence="2608939443" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="CAS" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCode="PV 0770-0778" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9029341,
<specimenCode box="[582,754,519,545]" collectionCode="PV" pageId="61" pageNumber="70">PV 0770-0778</specimenCode>
,
<collectionCode box="[766,823,519,545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="70" type="Museum">CAS</collectionCode>
),
<specimenCount box="[842,872,519,545]" pageId="61" pageNumber="70" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939453" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="HNU" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCount="6" specimenCount-female="6" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022360,
<collectionCode box="[447,519,555,581]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="61" pageNumber="70">HNU</collectionCode>
), 2 ♁,
<specimenCount box="[594,630,554,580]" pageId="61" pageNumber="70" type="female">6 ♀</specimenCount>
, 1 juv
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939446" collectingDate="2005-05-13" collectingDateMax="2005-05-15" collectingDateMin="2005-05-13" collectionCode="CAS" collectorName="C. Griswold" country="China" county="Lushui Co." elevation="2470" latitude="25.99363" location="Chang Yan He" longLatPrecision="1" longitude="98.66651" municipality="Pianma Township" pageId="61" pageNumber="70" specimenCount="6" specimenCount-female="5" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022361,
<collectionCode box="[257,314,589,615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="70" type="Museum">CAS</collectionCode>
), 3 ♁,
<specimenCount box="[393,430,589,616]" pageId="61" pageNumber="70" type="female">5 ♀</specimenCount>
,
<specimenCount box="[452,515,589,615]" pageId="61" pageNumber="70" type="generic">1 juv.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[185,644,624,651]" pageId="61" pageNumber="70" type="etymology">
<paragraph blockId="61.[140,1108,307,1637]" box="[185,644,624,651]" pageId="61" pageNumber="70">
<emphasis bold="true" box="[185,320,624,650]" pageId="61" pageNumber="70">
<emphasis box="[185,315,624,650]" inLineHeading="true" pageId="61" pageNumber="70" reason="1">Etymology</emphasis>
.
</emphasis>
Named for the
<typeStatus box="[501,549,625,651]" pageId="61" pageNumber="70">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection pageId="61" pageNumber="70" type="diagnosis">
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
<emphasis bold="true" box="[185,312,659,685]" pageId="61" pageNumber="70">
<emphasis box="[185,307,659,685]" inLineHeading="true" pageId="61" pageNumber="70" reason="1">Diagnosis</emphasis>
.
</emphasis>
Male distinguished from other
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[665,803,660,686]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis box="[665,803,660,686]" italics="true" pageId="61" pageNumber="70">Crassignatha</emphasis>
</taxonomicName>
by the embolus shape (
<figureCitation captionStart="Figure 75" captionStartId="147.[140,212,1622,1644]" captionTargetBox="[180,1073,174,1590]" captionTargetId="figure@147.[168,1081,164,1602]" captionTargetPageId="147" captionText="Figure 75. Crassignatha pianma sp. n. from Chang Yan He, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768890" httpUri="https://zenodo.org/record/3768890/files/figure.png" pageId="61" pageNumber="70">Fig. 75A</figureCitation>
), which is shorter than any other species.
</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
Female distinguished by the procurved scape (
<figureCitation box="[720,827,730,756]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76A</figureCitation>
,
<figureCitation box="[842,890,730,756]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79B</figureCitation>
) composed of two lobes of equal width (
<figureCitation box="[382,486,765,791]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76B</figureCitation>
,
<figureCitation box="[497,547,765,791]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79A</figureCitation>
) in combination with ducts that do not run anteriorly between the spermathecae; other
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[590,728,800,826]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis box="[590,728,800,826]" italics="true" pageId="61" pageNumber="70">Crassignatha</emphasis>
</taxonomicName>
have the scape oriented ventrally to posteroventrally (
<figureCitation box="[364,463,835,862]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76F</figureCitation>
,
<figureCitation box="[474,527,836,862]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79D</figureCitation>
) with the distal lobe of the scape more narrow than the basal lobe (
<figureCitation box="[310,418,871,897]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76G</figureCitation>
,
<figureCitation box="[431,482,871,897]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79C</figureCitation>
) and/or with the ducts running anteriorly between the spermathecae before turning posteriorly (
<figureCitation box="[604,700,906,932]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76I</figureCitation>
,
<figureCitation box="[712,800,906,933]" captionStart="Figure 89" captionStartId="161.[140,212,700,722]" captionTargetBox="[140,1108,165,679]" captionTargetId="figure@161.[140,1109,164,679]" captionTargetPageId="161" captionText="Figure 89. A, B, Crassignatha ertou sp. n. from Zaotang He; C, D, Crassignatha gudu sp. n. from Zaotang He; E, F, Crassignatha longtou sp. n. from Yamu He. A, C, E, epigynum, ventral view; B, D, F, cleared vulva, dorsal view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768912" httpUri="https://zenodo.org/record/3768912/files/figure.png" pageId="61" pageNumber="70">89B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="62" lastPageNumber="71" pageId="61" pageNumber="70" type="description">
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
<emphasis bold="true" box="[185,335,941,967]" pageId="61" pageNumber="70">
<emphasis box="[185,330,941,967]" inLineHeading="true" pageId="61" pageNumber="70" reason="1">Description</emphasis>
.
</emphasis>
Carapace orange-brown with small tubercles and sulci. Sternum orange-brown. Legs orange, femora I and II slightly swollen basally in female. Abdomen subspherical, light to medium gray, with numerous small sclerotized patches, some bearing long setae, male with single orange scutum laterally and posteriorly, female with small sclerite around spinnerets (
<figureCitation box="[567,652,1082,1108]" captionStart="Figure 74" captionStartId="146.[140,212,1125,1147]" captionTargetBox="[163,1101,188,1093]" captionTargetId="figure@146.[140,1109,164,1105]" captionTargetPageId="146" captionText="Figure 74. Crassignatha pianma sp. n. from Chang Yan He. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3768888" httpUri="https://zenodo.org/record/3768888/files/figure.png" pageId="61" pageNumber="70">Fig. 74</figureCitation>
).
</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
Male palp: Median apophysis with two tapered distal processes. Embolus short, thick, and rigid, making a single turn (
<figureCitation box="[577,679,1152,1179]" captionStart="Figure 75" captionStartId="147.[140,212,1622,1644]" captionTargetBox="[180,1073,174,1590]" captionTargetId="figure@147.[168,1081,164,1602]" captionTargetPageId="147" captionText="Figure 75. Crassignatha pianma sp. n. from Chang Yan He, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768890" httpUri="https://zenodo.org/record/3768890/files/figure.png" pageId="61" pageNumber="70">Fig. 75A</figureCitation>
,
<figureCitation box="[691,743,1153,1179]" captionStart="Figure 77" captionStartId="149.[140,212,1205,1227]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@149.[140,1109,164,1186]" captionTargetPageId="149" captionText="Figure 77. Crassignatha pianma sp. n. from Chang Yan He, SEM. A-E, male; F, female. A, palp, prolateral; B, palp, retrolateral; C, palp, dorsal; D, distal ventral part of tibia II showing presumed mating claspers; E, F, prosoma, lateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum; TI: tibia; PA: patella." figureDoi="http://doi.org/10.5281/zenodo.3768894" httpUri="https://zenodo.org/record/3768894/files/figure.png" pageId="61" pageNumber="70">77A</figureCitation>
).
</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">
<taxonomicName box="[185,253,1188,1214]" class="Polychaeta" family="Polychaeta (awaiting allocation)" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polychaeta (awaiting allocation)" pageId="61" pageNumber="16" phylum="Annelida" rank="genus">Vulva</taxonomicName>
: Scape slightly procurved (
<figureCitation box="[567,675,1187,1214]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76A</figureCitation>
,
<figureCitation box="[691,741,1188,1214]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79B</figureCitation>
), with distal and basal lobes of equal width (
<figureCitation box="[292,399,1223,1249]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="61" pageNumber="70">Figs 76B</figureCitation>
,
<figureCitation box="[413,465,1223,1249]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="61" pageNumber="70">79A</figureCitation>
). Spermathecae separated by their diameter. Ducts arise from posteromesal part of spermathecae, follow complex path mostly posterior to spermathecae before reaching openings near apex of scape.
</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" pageId="61" pageNumber="70">Male (CASENT 9022360): Total length 0.86, carapace 0.43 long, 0.40 wide, clypeus 0.15, sternum 0.28 long, 0.26 wide, coxa IV separated by 2.15 times their width. Leg I: patella d1, tibia d2, metatarsus slightly sinuous, with two stiff bristles prolaterally; Leg II: patella d1, tibia d2, v3, metatarsus excavated proximoventrally; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.44; TmIII: absent. Leg measurements: see Appendix A.</paragraph>
<paragraph blockId="61.[140,1108,307,1637]" lastBlockId="62.[140,1107,167,264]" lastPageId="62" lastPageNumber="71" pageId="61" pageNumber="70">Female (CASENT 9022360): Total length 1.10, carapace 0.49 long, 0.45 wide, clypeus 0.11, sternum 0.30 long, 0.27 wide, coxa IV separated by 1.41 times their width. Leg I: patella d1, tibia d2; Leg II: patella d1, tibia d2; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.46; TmIII: absent. Leg measurements: see Appendix A.</paragraph>
</subSubSection>
<subSubSection box="[185,1002,237,264]" pageId="62" pageNumber="71" type="discussion">
<paragraph blockId="62.[140,1107,167,264]" box="[185,1002,237,264]" pageId="62" pageNumber="71">
<emphasis bold="true" box="[185,381,237,263]" pageId="62" pageNumber="71">
<emphasis box="[185,277,237,263]" inLineHeading="true" pageId="62" pageNumber="71" reason="1">Natural</emphasis>
History.
</emphasis>
This species builds a horizontal orb web (
<figureCitation box="[852,935,237,263]" captionStart="Figure 72" captionStartId="144.[140,212,1112,1134]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@144.[140,1109,164,1092]" captionTargetPageId="144" captionText="Figure 72. Webs of symphytognathid spiders. A, Patu jidanweishi sp. n., female from Xiao Hei Shan Nature Reserve; B, Patu jidanweishi sp. n., male from Pianma; C, D, Crassignatha pianma sp. n., female from Chang Yan He; E, Crassignatha yinzhi sp. n., female from Xiao Hei Shan Nature Reserve." figureDoi="http://doi.org/10.5281/zenodo.3768884" httpUri="https://zenodo.org/record/3768884/files/figure.png" pageId="62" pageNumber="71">Fig. 72</figureCitation>
C-D).
</paragraph>
</subSubSection>
</treatment>
</document>
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