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<document id="7BD185B9069BD619A1D9E7C20B588B5B" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA7AFF8ACBAFB98CFDD2FA4F" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bellaspira margaritensis McLean &amp; Poorman 1970" docType="treatment" docVersion="12" lastPageNumber="35" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="32" updateTime="1698184378282" updateUser="plazi">
<mods:mods id="2AB4995987891404879BA01E69277124" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3091B2C47C97DBE2A7BC5F8327D29055">
<mods:title id="5D9B4186FAC1975792D8783FB182452B">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
</mods:titleInfo>
<mods:name id="2C0EDC442C8D9FCF10A8FFCFAF3BABB5" type="personal">
<mods:role id="5CF87912061A06F43A51DAC65F0E342F">
<mods:roleTerm id="3B67F7BDE4F6551360ADF92B1233D3A1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="756A912318CBF6AFEB90536556C29C2D">Fallon, Phillip J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2FA66AA8293638CF288D68015BE4D2A0">text</mods:typeOfResource>
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<mods:title id="226A7A104BF099188ABEBA627FCFFB5E">Zootaxa</mods:title>
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<mods:part id="17F33A873B518B3CDF5328C56618FA6A">
<mods:date id="BB06170DD6B1FBCAFFBC22453C87CC19">2016</mods:date>
<mods:detail id="91855C929EA57CCCFACAE37897133FEC" type="volume">
<mods:number id="F4803AA44F0A29773763572B1D74151A">4090</mods:number>
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<mods:number id="5F48DB49CEFEA0D7458B049A9CEBF899">1</mods:number>
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<mods:classification id="ABCCC8D45C7F32B5EF959EA90F9E6AC4">journal article</mods:classification>
<mods:identifier id="36655EBFCA0F418FD6D7BEEB605A8785" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
<mods:identifier id="6EDD504CFE56AE656233FA469DC6233E" type="GBIF-Dataset">e6b35f5a-435e-4473-b29e-1e4d842f84b0</mods:identifier>
<mods:identifier id="C0A3A641FB4EFE3A042B82199E069335" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039F87C4FA7AFF8ACBAFB98CFDD2FA4F" ID-DOI="http://doi.org/10.5281/zenodo.6076263" ID-GBIF-Taxon="117911593" ID-Zenodo-Dep="6076263" LSID="urn:lsid:plazi:treatment:039F87C4FA7AFF8ACBAFB98CFDD2FA4F" httpUri="http://treatment.plazi.org/id/039F87C4FA7AFF8ACBAFB98CFDD2FA4F" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32">
<subSubSection id="C32C6559FA7AFFB7CBAFB98CFCF2FEFA" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B8936D2FA7AFFB7CBAFB98CFCFCFF18" blockId="31.[151,788,151,211]" box="[151,788,151,178]" pageId="31" pageNumber="32">
<heading id="D0C181BEFA7AFFB7CBAFB98CFCFCFF18" bold="true" box="[151,788,151,178]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="4C364D51FA7AFFB7CBAFB98CFCFCFF18" ID-CoL="5WGMX" authority="McLean &amp; Poorman, 1970" authorityName="McLean &amp; Poorman" authorityYear="1970" box="[151,788,151,178]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7CBAFB98CFCFCFF18" bold="true" box="[151,788,151,178]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBAFB98CFE28FF1B" bold="true" box="[151,448,151,177]" italics="true" pageId="31" pageNumber="32">Bellaspira margaritensis</emphasis>
McLean &amp; Poorman, 1970
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBAFB9A1FEF5FF79" blockId="31.[151,788,151,211]" box="[151,285,186,211]" pageId="31" pageNumber="32">
(
<figureCitation id="130D2A57FA7AFFB7CBA7B9A1FF11FF79" box="[159,249,186,211]" captionStart="PLATE 8" captionStartId="32.[151,234,1834,1856]" captionTargetBox="[246,1335,195,1796]" captionTargetId="figure@32.[246,1335,195,1796]" captionTargetPageId="32" captionText="PLATE 8. Bellaspira margaritensis McLean &amp; Poorman, 1970. Specimens from Margarita and nearby islands, Venezuela. Upper panel. Fig. 1: paratype, N of Margarita I. (USNM 679564): ventral, lateral &amp; dorsal views. Fig. 2: enlarged view of protoconch of specimen in Fig. 5 (not to scale). Fig. 3: apical view of specimen in Fig. 11, V = varix, L = edge of outer lip. Lower panel. Figs. 4 – 7, 16: W side of Punta Charagato, Cubagua I. (ANSP 464955); Figs. 8 – 10, 13: W side of Punta Charagato, Cubagua I. (USNM 1291319); Figs. 11, 15: N side of Coche I. (author’s coll.); Fig. 12: Coche I. (author’s coll.); Fig. 14: Pampatar, Margarita I. (author’s coll.)." httpUri="https://zenodo.org/record/263307/files/figure.png" pageId="31" pageNumber="32">Plates 8</figureCitation>
,
<figureCitation id="130D2A57FA7AFFB7CA3EB9A1FEFCFF79" box="[262,276,186,211]" captionStart="PLATE 9" captionStartId="33.[151,234,1790,1812]" captionTargetBox="[263,1321,195,1750]" captionTargetId="figure@33.[263,1321,195,1751]" captionTargetPageId="33" captionText="PLATE 9. Bellaspira margaritensis McLean &amp; Poorman, 1970. Specimens from Trinidad &amp; Tobago, coastal Venezuela, and Colombia. Figs. 1 – 3: Chimana Grande I., Puerto La Cruz, Venezuela (EFG 27065); Fig. 4: Chimana Grande I., Venezuela (author’s coll.); Figs. 5 – 8: Chimana Grande I., Puerto La Cruz, Venezuela. Figs. 5, 8: USNM 90008; Figs. 6 – 7: EFG 27065; Fig. 9: first island, Puerto La Cruz, Venezuela (author’s coll.); Figs. 10 – 11, 14 – 15: Scotland Bay, Trinidad I., Trinidad &amp; Tobago (author’s coll.); Figs. 12, 16: Chacachacare I., Trinidad I., Trinidad &amp; Tobago (author’s coll.); Fig. 13: Chaguaramas Bay, Chaguaramas, Trinidad I. (author’s coll.); Fig. 17: Store Bay, Tobago I., Trinidad &amp; Tobago (author’s coll.); Figs. 18 – 20: Saunders Bay, Chacachacare I., Trinidad &amp; Tobago (author’s coll.); Figs. 21 – 23: Tayrona National Park, Colombia (author’s coll.)." httpUri="https://zenodo.org/record/263308/files/figure.png" pageId="31" pageNumber="32">9</figureCitation>
)
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBAFB9E6FCF2FEFA" blockId="31.[151,1436,253,336]" pageId="31" pageNumber="32">
<taxonomicName id="4C364D51FA7AFFB7CBAFB9E6FD3BFEB9" authority="McLean &amp; Poorman, 1970: 5" authorityName="McLean &amp; Poorman" authorityPageNumber="5" authorityYear="1970" box="[151,723,253,275]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7CBAFB9E6FE7DFEB9" box="[151,405,253,275]" italics="true" pageId="31" pageNumber="32">Bellaspira margaritensis</emphasis>
McLean &amp; Poorman, 1970: 5
</taxonomicName>
, fig. 6: Abbott (1974: 276, no. 3130); Díaz
<emphasis id="B942EAC0FA7AFFB7CF9CB9E5FB33FEB9" box="[1188,1243,253,275]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
(1990: 196); Díaz (1994: 40); Díaz &amp; Puyana (1994: 225, pl. 67, fig. 892); Williams (2005; 2006; 2009: 1510); Massemin
<emphasis id="B942EAC0FA7AFFB7CFD5B806FAC9FE98" box="[1261,1313,284,306]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
(2009: 204, text photo); Daccarett &amp; Bossio (2011: 131–132, fig. 809).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA7AFFB7CBAFB862FBB3FE40" pageId="31" pageNumber="32" type="description">
<paragraph id="8B8936D2FA7AFFB7CBAFB862FD24FE24" blockId="31.[151,1436,376,490]" box="[151,716,376,398]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBAFB862FD24FE24" bold="true" box="[151,716,376,398]" pageId="31" pageNumber="32">Not this species or unconfirmed extralimital reports.</emphasis>
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBAFB88DFBB3FE40" blockId="31.[151,1436,376,490]" pageId="31" pageNumber="32">
<taxonomicName id="4C364D51FA7AFFB7CBAFB88DFD42FE06" authority="McLean &amp; Poorman, 1970" authorityName="McLean &amp; Poorman" authorityYear="1970" box="[151,682,406,428]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7CBAFB88DFE7AFE06" box="[151,402,406,428]" italics="true" pageId="31" pageNumber="32">Bellaspira margaritensis</emphasis>
McLean &amp; Poorman, 1970
</taxonomicName>
: Díaz &amp; Puyana (1994: 225 [reference to
<collectingCountry id="F3217642FA7AFFB7CF76B88DFB4DFE06" box="[1102,1189,406,428]" name="Curaçao" pageId="31" pageNumber="32">Curaçao</collectingCountry>
,
<collectingCountry id="F3217642FA7AFFB7CF97B88DFAEAFE06" box="[1199,1282,406,428]" name="Bonaire, Sint Eustatius and Saba" pageId="31" pageNumber="32">Bonaire</collectingCountry>
, and Sotavento Is.]); Sander &amp; Lalli (1982: 317, Table 4 [off
<collectingCountry id="F3217642FA7AFFB7C996B8AEFCF9FE61" box="[686,785,437,459]" name="Barbados" pageId="31" pageNumber="32">Barbados</collectingCountry>
; =
<taxonomicName id="4C364D51FA7AFFB7C815B8AEFBF4FE61" box="[813,1052,437,459]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="barbadensis" status="sp. nov.">
<emphasis id="B942EAC0FA7AFFB7C815B8AEFBF4FE61" box="[813,1052,437,459]" italics="true" pageId="31" pageNumber="32">Bellaspira barbadensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA7AFFB7CF13B8ADFB46FE61" box="[1067,1198,437,459]" pageId="31" pageNumber="32" rank="species">new species</taxonomicNameLabel>
]); Odé (1991c: 20 [off Galveston-Freeport and the Louisiana coast; on Stetson Bank and in the mudlump fauna]).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA7AFFB7CBAFBB09FE82F958" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8936D2FA7AFFB7CBAFBB09FBCAFDE4" blockId="31.[151,1437,529,2030]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBAFBB09FEAEFD80" bold="true" box="[151,326,529,554]" pageId="31" pageNumber="32">
<typeStatus id="548D8870FA7AFFB7CBAFBB09FF3AFD83" box="[151,210,530,553]" pageId="31" pageNumber="32">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548D8870FA7AFFB7CA77BB0AFE5FFD80" box="[335,439,529,554]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
15.0 x
<quantity id="4CCE9B37FA7AFFB7C928BB09FD8FFD83" box="[528,615,530,553]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="mm" value="5.5">5.5 mm</quantity>
(LACM-AHF 1337);
<typeStatus id="548D8870FA7AFFB7C85EBB09FC3EFD83" box="[870,982,530,553]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
: 2 spec. (LACM-AHF 1338) &amp; 1 spec., 13.1 x
<quantity id="4CCE9B37FA7AFFB7CBDCBB2DFED2FDE7" box="[228,314,566,590]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="31" pageNumber="32" unit="mm" value="5.2">5.2 mm</quantity>
(USNM 679564). Only the
<typeStatus id="548D8870FA7AFFB7C94DBB2DFD3DFDE7" box="[629,725,566,589]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
in the USNM was examined.
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBFFBB41FE46FD3C" blockId="31.[151,1437,529,2030]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBFFBB41FE8EFDD8" bold="true" box="[199,358,601,626]" pageId="31" pageNumber="32">
<typeStatus id="548D8870FA7AFFB7CBFFBB41FEEAFDDB" box="[199,258,602,625]" pageId="31" pageNumber="32">Type</typeStatus>
locality.
</emphasis>
R/V
<emphasis id="B942EAC0FA7AFFB7CA9ABB42FDE4FDDB" box="[418,524,601,626]" italics="true" pageId="31" pageNumber="32">Velero III</emphasis>
Sta. A42-39, 4–
<quantity id="4CCE9B37FA7AFFB7C9FEBB42FD1FFDD8" box="[710,759,601,626]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="31" pageNumber="32" unit="mi" value="7.0">7 mi</quantity>
[
<quantity id="4CCE9B37FA7AFFB7C83EBB42FC85FDD8" box="[774,877,601,626]" metricMagnitude="3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="31" pageNumber="32" unit="km" value="8.5" valueMax="11.0" valueMin="6.0">6–11 km</quantity>
] N of Margarita I.,
<collectingCountry id="F3217642FA7AFFB7CF70BB42FB56FDD8" box="[1096,1214,601,626]" name="Venezuela" pageId="31" pageNumber="32">Venezuela</collectingCountry>
,
<geoCoordinate id="EE025015FA7AFFB7CFF1BB42FAC0FDDB" box="[1225,1320,601,625]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="11.133333">11°08′N</geoCoordinate>
,
<geoCoordinate id="EE025015FA7AFFB7CE0CBB42FA7FFDDB" box="[1332,1431,601,625]" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="-64.28333">64°17′W</geoCoordinate>
, in 17–21 fms [
<quantity id="4CCE9B37FA7AFFB7CA01BB65FE4BFD3C" box="[313,419,638,662]" metricMagnitude="1" metricUnit="m" metricValue="3.45" metricValueMax="3.8" metricValueMin="3.1" pageId="31" pageNumber="32" unit="m" value="34.5" valueMax="38.0" valueMin="31.0">31–38 m</quantity>
].
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBFFBBBAFEACFA5C" blockId="31.[151,1437,529,2030]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBFFBBBAFE16FD10" bold="true" box="[199,510,673,698]" pageId="31" pageNumber="32">Other material examined.</emphasis>
An additional
<specimenCount id="9D30FD5BFA7AFFB7C991BBB9FCABFD10" box="[681,835,673,698]" pageId="31" pageNumber="32" type="generic">55 specimens</specimenCount>
have been examined:
<emphasis id="B942EAC0FA7AFFB7CF7ABBBAFB53FD10" bold="true" box="[1090,1211,673,698]" italics="true" pageId="31" pageNumber="32">
<collectingCountry id="F3217642FA7AFFB7CF7ABBBAFB5CFD10" box="[1090,1204,673,698]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
:
</emphasis>
3 spec., 13.4 x 5.2, 13.2 x 5.3 &amp; 12.5 x
<quantity id="4CCE9B37FA7AFFB7CAA8BBDDFE04FD77" box="[400,492,710,734]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="31" pageNumber="32" unit="mm" value="5.1">5.1 mm</quantity>
, dredged in
<quantity id="4CCE9B37FA7AFFB7C9BABBDDFD29FD77" box="[642,705,710,734]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="32" unit="m" value="14.0">14 m</quantity>
, on coralline sand, Tayrona Park, Magdalena Dept.,
<collectingCountry id="F3217642FA7AFFB7CE16BBDEFA74FD74" box="[1326,1436,709,734]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
(author’s coll.).
<emphasis id="B942EAC0FA7AFFB7CA69BBF2FE27FCA8" bold="true" box="[337,463,745,770]" italics="true" pageId="31" pageNumber="32">
<collectingCountry id="F3217642FA7AFFB7CA69BBF2FE20FCA8" box="[337,456,745,770]" name="Venezuela" pageId="31" pageNumber="32">Venezuela</collectingCountry>
:
</emphasis>
1 spec., 10.7 x
<quantity id="4CCE9B37FA7AFFB7C9B6BBF1FD00FCAB" box="[654,744,746,770]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="4.5">4.5 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C82FBBF1FCBDFCAB" box="[791,853,746,770]" metricMagnitude="1" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="32" unit="m" value="21.0">21 m</quantity>
, first island, Puerto La Cruz, Anzoategui State, G. Mackintosh!
<date id="FF881012FA7AFFB7CA14BA15FE28FC8C" box="[300,448,782,806]" pageId="31" pageNumber="32" value="1993-08-15">15 Aug 1993</date>
(author’s coll.); 1 spec., 11.3 x
<quantity id="4CCE9B37FA7AFFB7C811BA15FC6AFC8F" box="[809,898,782,806]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="31" pageNumber="32" unit="mm" value="4.3">4.3 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C894BA15FC1EFC8F" box="[940,1014,782,806]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="31" pageNumber="32" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
, Puerto La Cruz, Chimana Grande I., Anzoategui State, G. Mackintosh!
<date id="FF881012FA7AFFB7C922BA29FD62FCE0" box="[538,650,818,842]" pageId="31" pageNumber="32" value="1993-08">Aug 1993</date>
(P. Stahlschmidt coll.); 8 spec., 11.7 x
<quantity id="4CCE9B37FA7AFFB7CF7BBA29FB72FCE3" box="[1091,1178,818,842]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="4.5">4.5 mm</quantity>
(author’s coll.), 11.2 x 4.2, &amp; 10.0 x 4.0 mm (USNM 900088), 10.6 x 4.2, 10.4 x 4.4, 10.0 x 4.1, 9.1 x 3.8 &amp; 8.3 x
<quantity id="4CCE9B37FA7AFFB7CF9BBA4DFB11FCC7" box="[1187,1273,854,878]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="32" unit="mm" value="3.5">3.5 mm</quantity>
(EFG 27065), in
<quantity id="4CCE9B37FA7AFFB7CB8CBA61FF1DFC3B" box="[180,245,890,914]" metricMagnitude="0" metricUnit="m" metricValue="3.7" pageId="31" pageNumber="32" unit="m" value="3.7">3.7 m</quantity>
, Chimana Grande I., G. Mackintosh!
<date id="FF881012FA7AFFB7C9A0BA61FCC5FC38" box="[664,813,890,914]" pageId="31" pageNumber="32" value="1993-08-29">29 Aug 1993</date>
; 1 spec., 12.2 x
<quantity id="4CCE9B37FA7AFFB7C8D9BA61FBD1FC3B" box="[993,1081,890,914]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="31" pageNumber="32" unit="mm" value="4.4">4.4 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7CF59BA61FB66FC3B" box="[1121,1166,890,914]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="m" value="6.0">6 m</quantity>
, Pampatar, Margarita I., G. Mackintosh!
<date id="FF881012FA7AFFB7CA71BA85FE39FC1C" box="[329,465,926,950]" pageId="31" pageNumber="32" value="1993-06-15">15 Jun 1993</date>
(author’s coll.); 1 spec., 14.9 x
<quantity id="4CCE9B37FA7AFFB7C80ABA85FC62FC1F" box="[818,906,926,950]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="31" pageNumber="32" unit="mm" value="5.8">5.8 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C889BA85FC02FC1F" box="[945,1002,926,950]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="31" pageNumber="32" unit="m" value="11.0">11 m</quantity>
, N side of Coche I., G. Mackintosh!
<date id="FF881012FA7AFFB7CEB8BA85FEECFC70" pageId="31" pageNumber="32" value="1999-09-23">23 Sep 1999</date>
(author’s coll.); 1 spec., 13.0 x
<quantity id="4CCE9B37FA7AFFB7C9BABAD9FD35FC73" box="[642,733,962,986]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="31" pageNumber="32" unit="mm" value="4.9">4.9 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C835BAD9FCA4FC73" box="[781,844,962,986]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="15.0">15 m</quantity>
, N side of Coche I., G. Mackintosh!
<date id="FF881012FA7AFFB7CE3ABAD9FA74FC70" box="[1282,1436,962,986]" pageId="31" pageNumber="32" value="1999-08-30">30 Aug 1999</date>
(author’s coll.); 1 spec., 10.0 x
<quantity id="4CCE9B37FA7AFFB7CACABAFDFDA1FC57" box="[498,585,998,1022]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="31" pageNumber="32" unit="mm" value="3.7">3.7 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C949BAFDFD43FC57" box="[625,683,998,1022]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="15.0">15 m</quantity>
, Coche I., G. Mackintosh!
<date id="FF881012FA7AFFB7C8EDBAFDFBBFFC54" box="[981,1111,997,1022]" pageId="31" pageNumber="32" value="1993-07-16">16 Jul 1993</date>
(author’s coll.); 1 spec., 11.2 x
<quantity id="4CCE9B37FA7AFFB7CB93BD11FEEBFB8B" box="[171,259,1034,1058]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="31" pageNumber="32" unit="mm" value="4.3">4.3 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7CA11BD11FE8CFB8B" box="[297,356,1034,1058]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="15.0">15 m</quantity>
, N side of Coche I., G. Mackintosh!
<date id="FF881012FA7AFFB7C9C0BD11FC61FB88" box="[760,905,1034,1058]" pageId="31" pageNumber="32" value="1998-08-30">30 Aug 1998</date>
(author’s coll.); 23 spec., 10.4 x 4.1, 10.6 x 4.1, 10.2 x 3.7, 10.3 x 3.9, 10.1 x
<quantity id="4CCE9B37FA7AFFB7CAE5BD35FDDBFBEF" box="[477,563,1070,1094]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="mm" value="3.6">3.6 mm</quantity>
(ANSP 464955), 12.8 x 5.1, 9.6 x 4.2, 9.5 x 3.8, 9.4 x 4.0, 9.8 x 4.0, 9.3 x 3.9, 9.3 x 3.7, 9.8 x 3.2, 9.3 x 3.8, 9.5 x 3.9, 8.9 x 3.8, 8.5 x 3.6, 8.9 x 3.5, 8.9 x 3.6 &amp; 9.0 x
<quantity id="4CCE9B37FA7AFFB7CFB1BD49FB0AFBC3" box="[1161,1250,1106,1130]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="31" pageNumber="32" unit="mm" value="3.6">3.6 mm</quantity>
, (author’s coll.), 11.4 x 4.5, 11.1 x 4.1, 11.5 x
<quantity id="4CCE9B37FA7AFFB7CACFBD6DFDB8FB27" box="[503,592,1142,1166]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="4.5">4.5 mm</quantity>
(USNM 1291319) in
<quantity id="4CCE9B37FA7AFFB7C86CBD6DFC72FB27" box="[852,922,1142,1166]" metricMagnitude="0" metricUnit="m" metricValue="4.6" pageId="31" pageNumber="32" unit="m" value="4.6">4.6 m</quantity>
, W side of Punta Charagato, Cubagua I., G. Mackintosh!
<date id="FF881012FA7AFFB7CA13BD81FE28FB18" box="[299,448,1178,1202]" pageId="31" pageNumber="32" value="2001-08-31">31 Aug 2001</date>
; 1 spec., 13.6 x
<quantity id="4CCE9B37FA7AFFB7C94CBD81FD24FB1B" box="[628,716,1178,1202]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="31" pageNumber="32" unit="mm" value="4.7">4.7 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C9CCBD81FCD4FB1B" box="[756,828,1178,1202]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="31" pageNumber="32" unit="m" value="4.5" valueMax="5.0" valueMin="4.0">4–5 m</quantity>
, off Punta Charagato, Cubagua I., G. Mackintosh!
<date id="FF881012FA7AFFB7CE4ABD81FF27FB7C" pageId="31" pageNumber="32" value="1999-09">Sep 1999</date>
(P. Stahlschmidt coll.); 1 spec., 12.4 x
<quantity id="4CCE9B37FA7AFFB7C9B4BDA5FD0DFB7F" box="[652,741,1214,1238]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="31" pageNumber="32" unit="mm" value="4.9">4.9 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C837BDA5FCD5FB7F" box="[783,829,1214,1238]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="m" value="8.0">8 m</quantity>
,
<collectingCountry id="F3217642FA7AFFB7C871BDA5FC48FB7F" box="[841,928,1214,1237]" name="Haiti" pageId="31" pageNumber="32">Tortuga</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CF44BDA5FB0EFB7C" box="[1148,1254,1214,1238]" pageId="31" pageNumber="32" value="1993-09">Sep 1993</date>
(author’s coll.).
<emphasis id="B942EAC0FA7AFFB7CBAFBDFAFE96FB50" bold="true" box="[151,382,1249,1274]" italics="true" pageId="31" pageNumber="32">
<collectingCountry id="F3217642FA7AFFB7CBAFBDFAFE9FFB50" box="[151,375,1249,1274]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad &amp; Tobago</collectingCountry>
:
</emphasis>
1 spec., 12.5 x
<quantity id="4CCE9B37FA7AFFB7C90ABDF9FD63FB53" box="[562,651,1250,1274]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="31" pageNumber="32" unit="mm" value="5.1">5.1 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C98DBDF9FD19FB53" box="[693,753,1250,1274]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="31" pageNumber="32" unit="m" value="29.0">29 m</quantity>
, Store Bay,
<collectingCountry id="F3217642FA7AFFB7C842BDFAFC26FB50" box="[890,974,1249,1274]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Tobago</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CF92BDF9FAD8FB50" box="[1194,1328,1250,1274]" pageId="31" pageNumber="32" value="1997-11-02">2 Nov 1997</date>
(author’s coll.); 2 spec., 13.3 x 4.7 &amp; 11.9 x
<quantity id="4CCE9B37FA7AFFB7C924BC1DFD9CFAB7" box="[540,628,1286,1310]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="4.5">4.5 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C9A5BC1DFD30FAB7" box="[669,728,1286,1310]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="24.0">24 m</quantity>
, Chacachacare I., G. Mackintosh!
<date id="FF881012FA7AFFB7CF60BC1DFB0FFAB4" box="[1112,1255,1286,1310]" pageId="31" pageNumber="32" value="1996-12-29">29 Dec 1996</date>
(author’s coll.); 3 spec., 12.4 x 4.6, 13.1 x 4.8 &amp; 15.4 x
<quantity id="4CCE9B37FA7AFFB7C965BC31FD5EFAEB" box="[605,694,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="31" pageNumber="32" unit="mm" value="5.2">5.2 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C9D8BC31FCF5FAEB" box="[736,797,1322,1346]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="24.0">24 m</quantity>
, Saunders Bay, Chacachacare I., G. Mackintosh!
<date id="FF881012FA7AFFB7CE73BC31FF27FACC" pageId="31" pageNumber="32" value="1996-12-29">29 Dec 1996</date>
(author’s coll.); 1 spec., 12.3 x
<quantity id="4CCE9B37FA7AFFB7C90ABC55FD62FACF" box="[562,650,1358,1382]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="31" pageNumber="32" unit="mm" value="4.8">4.8 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C989BC55FD04FACF" box="[689,748,1358,1382]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="31" pageNumber="32" unit="m" value="17.0">17 m</quantity>
, Chaguaramas Bay,
<collectingCountry id="F3217642FA7AFFB7C8E8BC56FBD8FACC" box="[976,1072,1357,1382]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CE3FBC55FA74FACC" box="[1287,1436,1358,1382]" pageId="31" pageNumber="32" value="2003-05-27">27 May 2003</date>
(author’s coll.); 1 spec., 10.2 x
<quantity id="4CCE9B37FA7AFFB7CAC4BC69FDBDFA23" box="[508,597,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="31" pageNumber="32" unit="mm" value="4.1">4.1 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C9B8BC69FD54FA23" box="[640,700,1394,1418]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="31" pageNumber="32" unit="m" value="22.0">22 m</quantity>
,
<collectingCountry id="F3217642FA7AFFB7C9F1BC6AFCC3FA20" box="[713,811,1393,1418]" name="United Kingdom" pageId="31" pageNumber="32">Scotland</collectingCountry>
Bay,
<collectingCountry id="F3217642FA7AFFB7C856BC6AFC26FA20" box="[878,974,1393,1418]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CF94BC69FAC7FA20" box="[1196,1327,1394,1418]" pageId="31" pageNumber="32" value="2003-12-04">4 Dec 2003</date>
(author’s coll.); 1 spec., 11.4 x
<quantity id="4CCE9B37FA7AFFB7CAACBC8DFE06FA07" box="[404,494,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="31" pageNumber="32" unit="mm" value="4.3">4.3 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C923BC8DFDB0FA07" box="[539,600,1430,1454]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="24.0">24 m</quantity>
,
<collectingCountry id="F3217642FA7AFFB7C95FBC8EFD21FA04" box="[615,713,1429,1454]" name="United Kingdom" pageId="31" pageNumber="32">Scotland</collectingCountry>
Bay,
<collectingCountry id="F3217642FA7AFFB7C836BC8EFC85FA04" box="[782,877,1429,1454]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CF77BC8DFB08FA04" box="[1103,1248,1429,1454]" pageId="31" pageNumber="32" value="2009-02-27">27 Feb 2009</date>
(UF 470266); 3 spec., 10.9 x 4.4, 13.1 x 4.7 &amp; 14.2 x
<quantity id="4CCE9B37FA7AFFB7C97ABCA1FD73FA7B" box="[578,667,1466,1490]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="31" pageNumber="32" unit="mm" value="5.1">5.1 mm</quantity>
, in
<quantity id="4CCE9B37FA7AFFB7C9FCBCA1FCC1FA7B" box="[708,809,1466,1490]" metricMagnitude="1" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="31" pageNumber="32" unit="m" value="16.5" valueMax="18.0" valueMin="15.0">15–18 m</quantity>
,
<collectingCountry id="F3217642FA7AFFB7C80EBCA2FC70FA78" box="[822,920,1465,1490]" name="United Kingdom" pageId="31" pageNumber="32">Scotland</collectingCountry>
Bay,
<collectingCountry id="F3217642FA7AFFB7C8E1BCA2FBD1FA78" box="[985,1081,1465,1490]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad</collectingCountry>
I., G. Mackintosh!
<date id="FF881012FA7AFFB7CE2CBCA1FA74FA78" box="[1300,1436,1466,1490]" pageId="31" pageNumber="32" value="2006-01-27">27 Jan 2006</date>
(author’s coll.).
</paragraph>
<paragraph id="8B8936D2FA7AFFB7CBFFBF19FE82F958" blockId="31.[151,1437,529,2030]" pageId="31" pageNumber="32">
<emphasis id="B942EAC0FA7AFFB7CBFFBF19FE5DF9B0" bold="true" box="[199,437,1537,1562]" pageId="31" pageNumber="32">Range and habitat.</emphasis>
Coastal areas and islands of northern South
<collectingCountry id="F3217642FA7AFFB7C8F6BF1AFBDBF9B0" box="[974,1075,1537,1562]" name="United States of America" pageId="31" pageNumber="32">America</collectingCountry>
:
<collectingCountry id="F3217642FA7AFFB7CF7ABF1AFB58F9B0" box="[1090,1200,1537,1562]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
(Magdalena Dept.);
<collectingCountry id="F3217642FA7AFFB7CBAFBF3EFEE2F994" box="[151,266,1573,1598]" name="Venezuela" pageId="31" pageNumber="32">Venezuela</collectingCountry>
(Margarita, Coche, Cubagua,
<collectingCountry id="F3217642FA7AFFB7C95ABF3DFD54F997" box="[610,700,1574,1597]" name="Haiti" pageId="31" pageNumber="32">Tortuga</collectingCountry>
, and Chimana Grande Is.);
<collectingCountry id="F3217642FA7AFFB7C8CDBF3EFB09F994" box="[1013,1249,1573,1598]" name="Trinidad and Tobago" pageId="31" pageNumber="32">Trinidad and Tobago</collectingCountry>
Is.;
<collectingCountry id="F3217642FA7AFFB7CE2CBF3DFA83F994" box="[1300,1387,1574,1598]" name="Guyana" pageId="31" pageNumber="32">Guyana</collectingCountry>
and
<collectingCountry id="F3217642FA7AFFB7CBAFBF52FEEBF9C8" box="[151,259,1609,1634]" name="Suriname" pageId="31" pageNumber="32">Suriname</collectingCountry>
(
<collectingCountry id="F3217642FA7AFFB7CA28BF52FE81F9C8" box="[272,361,1609,1634]" name="Guyana" pageId="31" pageNumber="32">Guyana</collectingCountry>
Plateau). Specimens have been reported taken from
<quantity id="4CCE9B37FA7AFFB7C88DBF51FC1DF9CB" box="[949,1013,1610,1634]" metricMagnitude="0" metricUnit="m" metricValue="4.6" pageId="31" pageNumber="32" unit="m" value="4.6">4.6 m</quantity>
(Chimana Grande I.) and from
<quantity id="4CCE9B37FA7AFFB7CE6EBF51FF45F92F" metricMagnitude="1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="31" pageNumber="32" unit="m" value="55.0" valueMax="80.0" valueMin="30.0">30–80 m</quantity>
depths elsewhere. They have been reported from on coralline sand (
<collectingCountry id="F3217642FA7AFFB7C8DABF76FBBEF92C" box="[994,1110,1645,1670]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
), and muddy sand (
<collectingCountry id="F3217642FA7AFFB7CE7CBF76FA75F92C" box="[1348,1437,1645,1670]" name="Guyana" pageId="31" pageNumber="32">Guyana</collectingCountry>
Plateau). The shallow water specimens close to the Venezuelan coast are more heavily spirally ridged, probably an adaptation to a sandy, higher energy environment. The shallow water (
<quantity id="4CCE9B37FA7AFFB7C892BFAEFC03F967" box="[938,1003,1717,1742]" metricMagnitude="0" metricUnit="m" metricValue="4.6" pageId="31" pageNumber="32" unit="m" value="4.6">4.6 m</quantity>
depth) Cubagua I. specimens are from a sheltered habitat.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA7AFF8ACBFFBFE6FCBBFC0F" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
<paragraph id="8B8936D2FA7AFF8ACBFFBFE6FD51FEEA" blockId="31.[151,1437,529,2030]" lastBlockId="34.[151,1437,151,1509]" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32">
<taxonomicName id="4C364D51FA7AFFB7CBFFBFE6FE09F8BF" box="[199,481,1789,1814]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7CBFFBFE6FE09F8BF" box="[199,481,1789,1814]" italics="true" pageId="31" pageNumber="32">Bellaspira margaritensis</emphasis>
</taxonomicName>
appears to be confined to the shallow coastal shelf, its coastline and islands, and absent from the offshore islands. Specimens reported from the offshore islands of
<collectingCountry id="F3217642FA7AFFB7CF51BE39FB21F890" box="[1129,1225,1826,1850]" name="Curaçao" pageId="31" pageNumber="32">Curaçao</collectingCountry>
,
<collectingCountry id="F3217642FA7AFFB7CFE2BE3AFADDF890" box="[1242,1333,1825,1850]" name="Bonaire, Sint Eustatius and Saba" pageId="31" pageNumber="32">Bonaire</collectingCountry>
, and the Sotaveto Is. by Díaz &amp; Puyana (1994: 225, pl. 67, fig. 892) should be verified. Specimens identified as
<taxonomicName id="4C364D51FA7AFFB7CE20BE5EFA75F8F4" box="[1304,1437,1861,1886]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA7AFFB7CE20BE5EFA75F8F4" box="[1304,1437,1861,1886]" italics="true" pageId="31" pageNumber="32">Splendrillia</emphasis>
</taxonomicName>
sp. by Jong &amp; Coomans (1988: 113, pl. 43, fig. 598) but said instead to be this species by Diaz &amp; Puyana (1994: 225), are not on the basis of the specimen shown in their figure 598. No other evidence has been found that
<taxonomicName id="4C364D51FA7AFFB7CEBCBE95FEDAF863" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7CEBCBE95FEDAF863" italics="true" pageId="31" pageNumber="32">B. margaritensis</emphasis>
</taxonomicName>
occurs in
<collectingCountry id="F3217642FA7AFFB7CA9CBEAAFDBCF860" box="[420,596,1969,1994]" name="Netherlands" pageId="31" pageNumber="32">the Netherlands</collectingCountry>
Antilles, or any other offshore island not connected to the South American continental shelf. The report of the occurrence of
<taxonomicName id="4C364D51FA7AFFB7C9E0BECDFC7AF847" box="[728,914,2006,2029]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA7AFFB7C9E0BECDFC7AF847" box="[728,914,2006,2029]" italics="true" pageId="31" pageNumber="32">B. margaritensis</emphasis>
</taxonomicName>
off
<collectingCountry id="F3217642FA7AFFB7C887BECEFBC1F844" box="[959,1065,2005,2030]" name="Barbados" pageId="31" pageNumber="32">Barbados</collectingCountry>
by Sander &amp; Lalli (1982: 317) have been determined to be
<taxonomicName id="4C364D51FA47FF8ACAC9B983FD48FF1A" box="[497,672,151,176]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="barbadensis" status="sp. nov.">
<emphasis id="B942EAC0FA47FF8ACAC9B983FD48FF1A" box="[497,672,151,176]" italics="true" pageId="34" pageNumber="35">B. barbadensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA47FF8AC98CB983FCACFF1A" box="[692,836,151,176]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
, on examination of their specimens in the ANSP. Specimens have been examined from most but not the entire range in published reports of the species. The photograph in Massemin
<emphasis id="B942EAC0FA47FF8ACA8BB9FBFE04FF52" box="[435,492,223,248]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(2009) appears to be this species and if so extends its range to at least as far south as the
<collectingCountry id="F3217642FA47FF8ACBF9B81FFEF0FEB6" box="[193,280,260,284]" name="Guyana" pageId="34" pageNumber="35">Guyana</collectingCountry>
and
<collectingCountry id="F3217642FA47FF8ACA68B81FFE53FEB7" box="[336,443,260,285]" name="Suriname" pageId="34" pageNumber="35">Suriname</collectingCountry>
plateau (in
<quantity id="4CCE9B37FA47FF8AC97BB81FFD40FEB6" box="[579,680,260,284]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="34" pageNumber="35" unit="m" value="60.0" valueMax="80.0" valueMin="40.0">40–80 m</quantity>
, on muddy sand). Specimens tentatively identified as this from the Gulf of
<collectingCountry id="F3217642FA47FF8ACBD7B83CFEACFEEA" box="[239,324,295,320]" name="Mexico" pageId="34" pageNumber="35">Mexico</collectingCountry>
by Odé are unlikely this species.
</paragraph>
<caption id="DF49665AFA45FF88CBAFBE31FD9BF870" httpUri="https://zenodo.org/record/263307/files/figure.png" pageId="32" pageNumber="33" targetBox="[246,1335,195,1796]" targetPageId="32">
<paragraph id="8B8936D2FA45FF88CBAFBE31FD9BF870" blockId="32.[151,1437,1834,2010]" pageId="32" pageNumber="33">
<emphasis id="B942EAC0FA45FF88CBAFBE31FEEDF8EA" bold="true" box="[151,261,1834,1856]" pageId="32" pageNumber="33">PLATE 8.</emphasis>
<taxonomicName id="4C364D51FA45FF88CA37BE31FCDAF8EA" authority="McLean &amp; Poorman, 1970" authorityName="McLean &amp; Poorman" authorityYear="1970" box="[271,818,1834,1856]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA45FF88CA37BE31FDE5F8EA" box="[271,525,1834,1856]" italics="true" pageId="32" pageNumber="33">Bellaspira margaritensis</emphasis>
McLean &amp; Poorman, 1970
</taxonomicName>
. Specimens from Margarita and nearby islands, Venezuela.
<emphasis id="B942EAC0FA45FF88CBAFBE51FECDF8F5" bold="true" box="[151,293,1866,1887]" pageId="32" pageNumber="33">Upper panel.</emphasis>
Fig. 1: paratype, N of Margarita I. (USNM 679564): ventral, lateral &amp; dorsal views. Fig. 2: enlarged view of protoconch of specimen in Fig. 5 (not to scale). Fig. 3: apical view of specimen in Fig. 11, V = varix, L = edge of outer lip.
<emphasis id="B942EAC0FA45FF88CBAFBE9CFECBF836" bold="true" box="[151,291,1927,1948]" pageId="32" pageNumber="33">Lower panel</emphasis>
. Figs. 4–7, 16: W side of Punta Charagato, Cubagua I. (ANSP 464955); Figs. 8–10, 13: W side of Punta Charagato, Cubagua I. (USNM 1291319); Figs. 11, 15: N side of Coche I. (author’s coll.); Fig. 12: Coche I. (author’s coll.); Fig. 14: Pampatar, Margarita I. (author’s coll.).
</paragraph>
</caption>
<caption id="DF49665AFA44FF89CBAFBFE5FF3BF841" httpUri="https://zenodo.org/record/263308/files/figure.png" pageId="33" pageNumber="34" targetBox="[263,1321,195,1750]" targetPageId="33">
<paragraph id="8B8936D2FA44FF89CBAFBFE5FF3BF841" blockId="33.[151,1436,1790,2027]" pageId="33" pageNumber="34">
<emphasis id="B942EAC0FA44FF89CBAFBFE5FEECF8BE" bold="true" box="[151,260,1790,1812]" pageId="33" pageNumber="34">PLATE 9.</emphasis>
<taxonomicName id="4C364D51FA44FF89CA34BFE5FCCFF8BE" authority="McLean &amp; Poorman, 1970" authorityName="McLean &amp; Poorman" authorityYear="1970" box="[268,807,1790,1812]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA44FF89CA34BFE5FDE1F8BE" box="[268,521,1790,1812]" italics="true" pageId="33" pageNumber="34">Bellaspira margaritensis</emphasis>
McLean &amp; Poorman, 1970
</taxonomicName>
. Specimens from Trinidad &amp; Tobago, coastal Venezuela, and Colombia. Figs. 1–3: Chimana Grande I., Puerto La Cruz, Venezuela (EFG 27065); Fig. 4: Chimana Grande I., Venezuela (author’s coll.); Figs. 5–8: Chimana Grande I., Puerto La Cruz, Venezuela. Figs. 5, 8: USNM 90008; Figs. 6–7: EFG 27065; Fig. 9: first island, Puerto La Cruz, Venezuela (author’s coll.); Figs. 10–11, 14–15: Scotland Bay, Trinidad I., Trinidad &amp; Tobago (author’s coll.); Figs. 12, 16: Chacachacare I., Trinidad I., Trinidad &amp; Tobago (author’s coll.); Fig. 13: Chaguaramas Bay, Chaguaramas, Trinidad I. (author’s coll.); Fig. 17: Store Bay, Tobago I., Trinidad &amp; Tobago (author’s coll.); Figs. 18–20: Saunders Bay, Chacachacare I., Trinidad &amp; Tobago (author’s coll.); Figs. 21–23: Tayrona National Park, Colombia (author’s coll.).
</paragraph>
</caption>
<paragraph id="8B8936D2FA47FF8ACBFFB857FCBBFC0F" blockId="34.[151,1437,151,1509]" pageId="34" pageNumber="35">
<emphasis id="B942EAC0FA47FF8ACBFFB857FE74FECF" bold="true" box="[199,412,332,357]" pageId="34" pageNumber="35">
Description.
<emphasis id="B942EAC0FA47FF8ACA58B857FE74FECF" bold="true" box="[352,412,332,357]" italics="true" pageId="34" pageNumber="35">Shell</emphasis>
</emphasis>
small (to
<quantity id="4CCE9B37FA47FF8AC937B857FD9BFECE" box="[527,627,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="34" pageNumber="35" unit="mm" value="15.6">15.6 mm</quantity>
in total length), fusiform, truncated anteriorly, up to 10 slightly convex to moderately convex whorls, suture undulates over rib ends, last whorl approximately 60% of total shell length.
<emphasis id="B942EAC0FA47FF8ACBAFB88FFEF5FE07" bold="true" box="[151,285,404,429]" italics="true" pageId="34" pageNumber="35">Protoconch</emphasis>
of approximately 2 smooth whorls, first not immersed in second, second large, its peripheral diameter greater than the first teleoconch whorl at suture.
<emphasis id="B942EAC0FA47FF8AC9DAB8ACFC70FE7A" bold="true" box="[738,920,439,464]" italics="true" pageId="34" pageNumber="35">Axial sculpture</emphasis>
of broad ribs but their width less than the intercostal space, relatively few in number, 6–8 on penultimate, 4–6 on last whorl to varix, tend to align whorl-towhorl. Last rib before varix sometimes incompletely formed—narrower or shorter than previous ones. Ribs run suture-to-suture on spire whorls, slightly reduced in sulcal area; crests round.
<emphasis id="B942EAC0FA47FF8AC8CEBB3FFBDAFD97" bold="true" box="[1014,1074,548,573]" italics="true" pageId="34" pageNumber="35">Varix</emphasis>
an expanded last or penultimate rib with up to 3 post-varical axial folds or axial folds, the last, on lip edge, follows the contour of the lip from the suture to stromboid notch.
<emphasis id="B942EAC0FA47FF8ACAFCBB77FD94FD2F" bold="true" box="[452,636,620,645]" italics="true" pageId="34" pageNumber="35">Spiral sculpture</emphasis>
of evenly-spaced incised lines over entire shell, closer in sulcal region, stronger and ridge-like on shell base.
<emphasis id="B942EAC0FA47FF8AC97EBB94FD7CFD02" bold="true" box="[582,660,655,680]" italics="true" pageId="34" pageNumber="35">Sulcus</emphasis>
absent, lower ribs and more closely spaced incised lines present on whorl shoulder.
<emphasis id="B942EAC0FA47FF8ACA74BBAFFE5DFD67" bold="true" box="[332,437,692,717]" italics="true" pageId="34" pageNumber="35">Outer lip</emphasis>
thin, somewhat flattened from varix to lip edge forming a low sigmoid arc from the anal sinus to the anterior canal, with a slight dip for a stromboid notch.
<emphasis id="B942EAC0FA47FF8AC8B9BBCCFC15FD5A" bold="true" box="[897,1021,727,752]" italics="true" pageId="34" pageNumber="35">Anal sinus</emphasis>
only a slight indentation of the outer lip beginning at the suture, when viewed laterally; inverted V-shaped with inner callus on both parietal wall and outer lip of mature specimens, when viewed ventrally.
<emphasis id="B942EAC0FA47FF8AC822BA3BFC6CFC92" bold="true" box="[794,900,799,824]" italics="true" pageId="34" pageNumber="35">Inner lip</emphasis>
emarginated, thickened posteriorly into a low callus at its junction with the outer lip.
<emphasis id="B942EAC0FA47FF8AC951BA5FFCFDFCF7" bold="true" box="[617,789,836,861]" italics="true" pageId="34" pageNumber="35">Anterior canal</emphasis>
open, deep, and notched; anterior fasciole not swollen, bearing 5–8 spiral ridges.
<emphasis id="B942EAC0FA47FF8ACAF8BA7CFDEAFC2A" bold="true" box="[448,514,871,896]" italics="true" pageId="34" pageNumber="35">Color</emphasis>
white, pink and light brown specimens are known; also white with areas flushed with pink, or white with faint brown spots on whorl shoulders.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA47FF8ACBFFBAABFDD2FA4F" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8B8936D2FA47FF8ACBFFBAABFDD2FA4F" blockId="34.[151,1437,151,1509]" pageId="34" pageNumber="35">
<emphasis id="B942EAC0FA47FF8ACBFFBAABFE50FC6D" bold="true" box="[199,440,944,967]" pageId="34" pageNumber="35">
Remarks.
<emphasis id="B942EAC0FA47FF8ACA7ABAABFE50FC6D" bold="true" box="[322,440,944,967]" italics="true" pageId="34" pageNumber="35">Taxonomy</emphasis>
</emphasis>
. According to McLean &amp; Poorman,
<typeStatus id="548D8870FA47FF8AC86EBAB4FC50FC62" box="[854,952,943,968]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
large for genus, protoconch with 2 smooth rounded whorls, teleoconch with 8 whorls and 7 axial ribs aligned from whorl-to-whorl, evenly rounded and without a subsutural channel; spiral sculpture of well-spaced incisions, more closely spaced in the subsutural area. Parietal callus well defined; edge of lip thin with a
<taxonomicName id="4C364D51FA47FF8AC9D7BD07FCDAFB9F" box="[751,818,1052,1077]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="minor">minor</taxonomicName>
rib, penultimate rib strongest; color white-pinkish on whorl periphery. This and examined specimens all have the characteristic anal sinus, spiral incised lines, and ribs that extend from suture-to-suture with only the slightest change in the sulcal area, and ribs number 7, usually aligned along all whorls, but not always.
<emphasis id="B942EAC0FA47FF8AC950BD9CFD36FB0A" bold="true" box="[616,734,1159,1184]" italics="true" pageId="34" pageNumber="35">Variability</emphasis>
. The 54 examined specimens have an average length of
<quantity id="4CCE9B37FA47FF8ACE66BD93FF2BFB6E" metricMagnitude="-2" metricUnit="m" metricValue="1.116" pageId="34" pageNumber="35" unit="mm" value="11.16">11.16 mm</quantity>
(
<quantity id="4CCE9B37FA47FF8ACBEBBDB7FE82FB6F" box="[211,362,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.56" metricValueMin="0.83" pageId="34" pageNumber="35" unit="mm" value="11.95" valueMax="15.6" valueMin="8.3">8.3–15.6 mm</quantity>
) and average W/
<taxonomicName id="4C364D51FA47FF8AC914BDB7FD90FB6F" box="[556,632,1196,1221]" class="Gastropoda" family="Drilliidae" genus="Lissodrillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="ratio">L ratio</taxonomicName>
of 0.391. Shells vary in form, some narrower than others. Color varies with white being the commonest and pink specimens being occasionally found. A light brown specimen from an island near Puerto La Cruz,
<collectingCountry id="F3217642FA47FF8ACAE8BDEFFDABFAA7" box="[464,579,1268,1293]" name="Venezuela" pageId="34" pageNumber="35">Venezuela</collectingCountry>
has faint brown spots on rib shoulders. Specimens from Chimana Grande I.,
<collectingCountry id="F3217642FA47FF8ACBAFBC0CFEE2FA9A" box="[151,266,1303,1328]" name="Venezuela" pageId="34" pageNumber="35">Venezuela</collectingCountry>
are shades of brown in color and have stronger spirally incised lines that are heavily ridged. These differences are not believed to rise to the specific level.
<emphasis id="B942EAC0FA47FF8AC813BC27FB1CFAFE" box="[811,1268,1340,1365]" italics="true" pageId="34" pageNumber="35">
<emphasis id="B942EAC0FA47FF8AC813BC27FC38FAFF" bold="true" box="[811,976,1340,1365]" italics="true" pageId="34" pageNumber="35">Identification.</emphasis>
<taxonomicName id="4C364D51FA47FF8AC8E2BC27FB1CFAFE" box="[986,1268,1340,1365]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="margaritensis">Bellaspira margaritensis</taxonomicName>
</emphasis>
should not be confused with any of its congeners, and is easily distinguished from other drilliids of the same size and color, such as
<taxonomicName id="4C364D51FA47FF8ACB8DBC9EFE69FA37" box="[181,385,1412,1437]" class="Gastropoda" family="Drilliidae" genus="Fenimorea" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="nivalis" status="sp. nov.">
<emphasis id="B942EAC0FA47FF8ACB8DBC9EFE69FA37" box="[181,385,1412,1437]" italics="true" pageId="34" pageNumber="35">Fenimorea nivalis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA47FF8ACAB7BC9EFDFEFA37" box="[399,534,1412,1437]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
by its distinctive anal sinus.
<taxonomicName id="4C364D51FA47FF8AC863BC9FFB88FA37" box="[859,1120,1412,1437]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="barbadensis" status="sp. nov.">
<emphasis id="B942EAC0FA47FF8AC863BC9FFB88FA37" box="[859,1120,1412,1437]" italics="true" pageId="34" pageNumber="35">Bellaspira barbadensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27157BBFA47FF8ACF56BC9EFB10FA37" box="[1134,1272,1412,1437]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
, usually found at greater depths, is very similar to
<taxonomicName id="4C364D51FA47FF8AC90EBCB3FD1CFA15" box="[566,756,1448,1471]" class="Gastropoda" family="Drilliidae" genus="Bellaspira" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="margaritensis">
<emphasis id="B942EAC0FA47FF8AC90EBCB3FD1CFA15" box="[566,756,1448,1471]" italics="true" pageId="34" pageNumber="35">B. margaritensis</emphasis>
</taxonomicName>
but differs in having more inflated whorls (sulcus is more obscure) and has different coloration.
</paragraph>
</subSubSection>
</treatment>
</document>
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