treatments-xml/data/03/9F/87/039F87C4FA4BFF86CBAFB983FB63FA3A.xml
2024-06-21 12:22:17 +02:00

182 lines
19 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="8D1F6C1005EDB5E0537DDCF846DEC097" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FA4BFF86CBAFB983FB63FA3A" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calliclava McLean 1971" docType="treatment" docVersion="12" lastPageNumber="47" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="47" updateTime="1698184378282" updateUser="plazi">
<mods:mods id="4D291280EEDA61645D8C7CB95EF9BB93" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CB1BFDA6531D2971AEB019369B9B86C7">
<mods:title id="E0BECB62A323BD8BA4ED0AA51D58890D">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
</mods:titleInfo>
<mods:name id="23F1408DFEC692E866658A2467507E25" type="personal">
<mods:role id="A814A2EB040CA69DC2243EC5E01D6EBE">
<mods:roleTerm id="2967D685DCA0F309BE36973C816C0ACB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E95F12F36AB32F314FAC59CAD5626716">Fallon, Phillip J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A83061353CE9C527F25EB42A5C45991F">text</mods:typeOfResource>
<mods:relatedItem id="B4F3B71E607BCAABF78FBD9C98052EC3" type="host">
<mods:titleInfo id="033A71753FC92D8F6BCACD6146950931">
<mods:title id="91FA224A7AE533A26588402EC741B525">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="350A25220EEDE41E678947784266F480">
<mods:date id="FFB92F02302BEEE242F2259DEE37CFA2">2016</mods:date>
<mods:detail id="242B1C70F2582BF78E86CA46B67A2F26" type="volume">
<mods:number id="187393F73982DC1A599A1A90A068B0E6">4090</mods:number>
</mods:detail>
<mods:detail id="37B9AFC8755F69613A9E50C97F1C2563" type="issue">
<mods:number id="E2CEC57D581C85263A53C7DEF02CE4FC">1</mods:number>
</mods:detail>
<mods:extent id="58F78F2310E726C9660636C52372F68C" unit="page">
<mods:start id="1EE16D0691D41D05046ED874A411D357">1</mods:start>
<mods:end id="CF0033EB0FCA99E182063D3734376E65">363</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9BC7FFAA3A975F6D2AFA8A82D03907B4">journal article</mods:classification>
<mods:identifier id="A184CE62A275F9EAB206BE7C18F33DD6" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
<mods:identifier id="62C93CAC5F76C20354DE24EDB8A5C2EA" type="GBIF-Dataset">e6b35f5a-435e-4473-b29e-1e4d842f84b0</mods:identifier>
<mods:identifier id="6FA3978C56BE382527F58B4AD09E140B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="45229D066879A83FB4C7EA0421779495" type="Zenodo-Dep">263299</mods:identifier>
<mods:identifier id="E7D4B09968EAEB5B9EFA29616ABC78F0" type="ZooBank">203BAC25-B542-48FE-B5AD-EBA8C0285833</mods:identifier>
</mods:mods>
<treatment id="039F87C4FA4BFF86CBAFB983FB63FA3A" ID-DOI="http://doi.org/10.5281/zenodo.6076279" ID-GBIF-Taxon="117911719" ID-Zenodo-Dep="6076279" LSID="urn:lsid:plazi:treatment:039F87C4FA4BFF86CBAFB983FB63FA3A" httpUri="http://treatment.plazi.org/id/039F87C4FA4BFF86CBAFB983FB63FA3A" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection id="C32C6559FA4BFF86CBAFB983FED5FEB9" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B8936D2FA4BFF86CBAFB983FDC9FF18" blockId="46.[151,545,151,178]" box="[151,545,151,178]" pageId="46" pageNumber="47">
<heading id="D0C181BEFA4BFF86CBAFB983FDC9FF18" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B942EAC0FA4BFF86CBAFB983FDC9FF18" bold="true" box="[151,545,151,178]" pageId="46" pageNumber="47">
Genus
<taxonomicName id="4C364D51FA4BFF86CBD6B98CFDC9FF18" ID-CoL="7NVM5" authority="McLean, 1971" authorityName="McLean" authorityYear="1971" box="[238,545,151,178]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CBD6B98CFE81FF1B" bold="true" box="[238,361,151,177]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
McLean, 1971
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8936D2FA4BFF86CBAFB9C4FED5FEB9" blockId="46.[151,1436,222,275]" pageId="46" pageNumber="47">
<typeStatus id="548D8870FA4BFF86CBAFB9C4FF21FF5E" box="[151,201,223,244]" pageId="46" pageNumber="47">Type</typeStatus>
species:
<taxonomicName id="4C364D51FA4BFF86CA17B9C5FD79FF5E" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[303,657,222,244]" class="Gastropoda" family="Drilliidae" genus="Cymatosyrinx" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="palmeri">
<emphasis id="B942EAC0FA4BFF86CA17B9C5FDFDFF5E" box="[303,533,222,244]" italics="true" pageId="46" pageNumber="47">Cymatosyrinx palmeri</emphasis>
Dall, 1919
</taxonomicName>
, by original designation (Recent, beach at the head of the Gulf, Gulf of California).
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA4BFF86CBAFB824FE0DFE42" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8B8936D2FA4BFF86CBAFB824FE0DFE42" blockId="46.[151,1437,319,560]" pageId="46" pageNumber="47">
<emphasis id="B942EAC0FA4BFF86CBAFB824FEF9FEF2" bold="true" box="[151,273,319,344]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
According to McLean (1971: 117), shell small to medium sized, surface glossy, body whorl short in comparison with the spire. Protoconch large, of two carinate whorls. Axials from suture-to-suture, but weak in sulcus. Spirals of incised grooves. Anal sinus on shoulder deep, U-shaped, bordered by a curved parietal callus. Outer lip nearly straight with a moderately deep stromboid notch; a varix is present ¼-turn from the edge of the lip. Anterior canal short, notched.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA4BFF86CBFFB8EEFEF1FD15" pageId="46" pageNumber="47" type="key">
<paragraph id="8B8936D2FA4BFF86CBFFB8EEFD26FD9A" blockId="46.[151,1437,319,560]" pageId="46" pageNumber="47">
<emphasis id="B942EAC0FA4BFF86CBFFB8EEFE54FDA7" bold="true" box="[199,444,500,525]" pageId="46" pageNumber="47">Key characteristics.</emphasis>
The combination of the following characteristics is diagnostic of
<taxonomicName id="4C364D51FA4BFF86CFD7B8EFFA8CFDA7" ID-CoL="7NVM5" box="[1263,1380,500,525]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CFD7B8EFFA8CFDA7" box="[1263,1380,500,525]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
and separates the genus from all other TWA
<taxonomicName id="4C364D51FA4BFF86C963BB0CFD23FD9A" ID-CoL="7NHDF" box="[603,715,535,560]" class="Gastropoda" family="Drilliidae" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="family">Drilliidae</taxonomicName>
:
</paragraph>
<paragraph id="8B8936D2FA4BFF86CBAFBB7BFEF1FD15" blockId="46.[151,1437,607,813]" pageId="46" pageNumber="47">1. Axial ribs numerous, extend from suture-to-suture, depressed and recurved in the sulcus, and run to anterior fasciole on body whorl; ribs overridden by spiral grooves that are shallow on crests, deeper in intercostal spaces;</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA4BFF86CBAFBBD7FD8CFAC6" pageId="46" pageNumber="47" type="description">
<paragraph id="8B8936D2FA4BFF86CBAFBBD7FC5BFD4F" blockId="46.[151,1437,607,813]" box="[151,947,716,741]" pageId="46" pageNumber="47">2. Protoconch of about 22¼ smooth, translucent, keeled whorls; and</paragraph>
<paragraph id="8B8936D2FA4BFF86CBAFBBEBFEE5FC87" blockId="46.[151,1437,607,813]" pageId="46" pageNumber="47">3. Varix broad and hump-like positioned about ⅓-turn from the edge of the outer lip; marked with a dark color patch.</paragraph>
<paragraph id="8B8936D2FA4BFF86CBFFBA46FD20FC4A" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
<emphasis id="B942EAC0FA4BFF86CBFFBA46FE2EFCDE" bold="true" box="[199,454,861,884]" pageId="46" pageNumber="47">Nomenclatural notes.</emphasis>
This genus was erected by McLean for species that otherwise resembled
<taxonomicName id="4C364D51FA4BFF86CFC1BA47FA7DFCDF" box="[1273,1429,860,885]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CFC1BA47FA7DFCDF" box="[1273,1429,860,885]" italics="true" pageId="46" pageNumber="47">Clathrodrillia</emphasis>
</taxonomicName>
, but differ in possessing a protoconch that is carinate from the tip of the first whorl, and a radula with a compressed, not elongate, lateral tooth. Dall erected separate genera,
<taxonomicName id="4C364D51FA4BFF86C825BABFFBF6FC16" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[797,1054,932,957]" class="Gastropoda" family="Drilliidae" genus="Elaeocyma" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C825BABFFC72FC17" box="[797,922,932,957]" italics="true" pageId="46" pageNumber="47">Elaeocyma</emphasis>
Dall, 1918
</taxonomicName>
; and
<taxonomicName id="4C364D51FA4BFF86CF65BABFFAF2FC16" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[1117,1306,932,957]" class="Gastropoda" family="Drilliidae" genus="Kylix" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CF65BABFFB71FC17" box="[1117,1177,932,957]" italics="true" pageId="46" pageNumber="47">Kylix</emphasis>
Dall, 1919
</taxonomicName>
for eastern Pacific drilliids with partially keeled protoconchs.
</paragraph>
<paragraph id="8B8936D2FA4BFF86CBFFBAF7FAA6FBDA" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
TWA species of
<taxonomicName id="4C364D51FA4BFF86CAB9BAF7FE1EFBAF" box="[385,502,1004,1029]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CAB9BAF7FE1EFBAF" box="[385,502,1004,1029]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
are similar in appearance to
<taxonomicName id="4C364D51FA4BFF86C802BAF7FB77FBAF" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[826,1183,1004,1029]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="palmeri">
<emphasis id="B942EAC0FA4BFF86C802BAF7FBE5FBAF" box="[826,1037,1004,1029]" italics="true" pageId="46" pageNumber="47">Calliclava palmeri</emphasis>
(Dall, 1919)
</taxonomicName>
, the
<typeStatus id="548D8870FA4BFF86CFEDBAF6FAEDFBAE" box="[1237,1285,1005,1028]" pageId="46" pageNumber="47">type</typeStatus>
of the genus. Another eastern Pacific species,
<taxonomicName id="4C364D51FA4BFF86C92DBD14FBC5FB82" authority="McLean &amp; Poorman, 1971" authorityName="McLean &amp; Poorman" authorityYear="1971" box="[533,1069,1039,1064]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="subtilis">
<emphasis id="B942EAC0FA4BFF86C92DBD14FD0EFB82" box="[533,742,1039,1064]" italics="true" pageId="46" pageNumber="47">Calliclava subtilis</emphasis>
McLean &amp; Poorman, 1971
</taxonomicName>
, is also similar. Ten species of
<taxonomicName id="4C364D51FA4BFF86CBAFBD2FFEE4FBE7" box="[151,268,1076,1101]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CBAFBD2FFEE4FBE7" box="[151,268,1076,1101]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
are listed in the on-line database, World Registry of Marine Species as of the date of this research, all from the eastern Pacific (WoRMS, 2016). The
<taxonomicName id="4C364D51FA4BFF86C99BBD4CFCF0FBDA" box="[675,792,1111,1136]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C99BBD4CFCF0FBDA" box="[675,792,1111,1136]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
herein are the first to be described from the TWA.
</paragraph>
<paragraph id="8B8936D2FA4BFF86CBFFBD67FAA8FB77" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
<emphasis id="B942EAC0FA4BFF86CBFFBD67FE68FB3E" bold="true" box="[199,384,1148,1173]" pageId="46" pageNumber="47">Similar genera.</emphasis>
The genus
<taxonomicName id="4C364D51FA4BFF86C93CBD67FD91FB3F" box="[516,633,1148,1173]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C93CBD67FD91FB3F" box="[516,633,1148,1173]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
is most easily distinguished by its fully carinate protoconch. Although otherwise close in appearance, only the second protoconch whorl of
<taxonomicName id="4C364D51FA4BFF86C8FCBD84FB2FFB12" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[964,1223,1183,1208]" class="Gastropoda" family="Drilliidae" genus="Elaeocyma" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C8FCBD84FBA9FB12" box="[964,1089,1183,1208]" italics="true" pageId="46" pageNumber="47">Elaeocyma</emphasis>
Dall, 1918
</taxonomicName>
is carinate. Other drilliid genera with carinate protoconchs include
<taxonomicName id="4C364D51FA4BFF86C985BDDFFCCDFB77" box="[701,805,1220,1245]" class="Gastropoda" family="Drilliidae" genus="Imaclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C985BDDFFCCDFB77" box="[701,805,1220,1245]" italics="true" pageId="46" pageNumber="47">Imaclava</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C364D51FA4BFF86C85BBDDFFC76FB77" box="[867,926,1220,1245]" class="Gastropoda" family="Drilliidae" genus="Kylix" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C85BBDDFFC76FB77" box="[867,926,1220,1245]" italics="true" pageId="46" pageNumber="47">Kylix</emphasis>
</taxonomicName>
, but none have a carinate first whorl.
</paragraph>
<paragraph id="8B8936D2FA4BFF86CBFFBDFCFD8CFAC6" blockId="46.[151,1437,860,1424]" pageId="46" pageNumber="47">
<taxonomicName id="4C364D51FA4BFF86CBFFBDFCFE09FAAA" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[199,481,1255,1280]" class="Gastropoda" family="Drilliidae" genus="Clathrodrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CBFFBDFCFE8BFAAA" box="[199,355,1255,1280]" italics="true" pageId="46" pageNumber="47">Clathrodrillia</emphasis>
Dall, 1918
</taxonomicName>
differs in possessing a smooth, not carinate protoconch.
<taxonomicName id="4C364D51FA4BFF86CF5CBDFCFA74FAAA" authority="Woodring, 1928" authorityName="Woodring" authorityYear="1928" box="[1124,1436,1255,1280]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CF5CBDFCFB37FAAA" box="[1124,1247,1255,1280]" italics="true" pageId="46" pageNumber="47">Agladrillia</emphasis>
Woodring, 1928
</taxonomicName>
differs in possessing only a carinate second protoconch whorl, a varix positioned ½-turn from the edge of the outer lip, and axials obsolete from the varix to the outer lip.
<taxonomicName id="4C364D51FA4BFF86C834BC34FC6FFAE2" box="[780,903,1327,1352]" class="Gastropoda" family="Drilliidae" genus="Agladrillia" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86C834BC34FC6FFAE2" box="[780,903,1327,1352]" italics="true" pageId="46" pageNumber="47">Agladrillia</emphasis>
</taxonomicName>
also has a more acutely “pinched” anterior, as opposed to the tapered one of
<taxonomicName id="4C364D51FA4BFF86CAD0BC4FFDB4FAC7" box="[488,604,1364,1389]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CAD0BC4FFDB4FAC7" box="[488,604,1364,1389]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C6559FA4BFF86CBFFBC6CFB63FA3A" box="[199,1163,1399,1424]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8B8936D2FA4BFF86CBFFBC6CFB63FA3A" blockId="46.[151,1437,860,1424]" box="[199,1163,1399,1424]" pageId="46" pageNumber="47">
<emphasis id="B942EAC0FA4BFF86CBFFBC6CFE88FA3A" bold="true" box="[199,352,1399,1424]" pageId="46" pageNumber="47">Distribution.</emphasis>
<taxonomicName id="4C364D51FA4BFF86CA5FBC6CFE34FA3A" box="[359,476,1399,1424]" class="Gastropoda" family="Drilliidae" genus="Calliclava" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B942EAC0FA4BFF86CA5FBC6CFE34FA3A" box="[359,476,1399,1424]" italics="true" pageId="46" pageNumber="47">Calliclava</emphasis>
</taxonomicName>
is only known from off the northern coast of South
<collectingCountry id="F3217642FA4BFF86CF1BBC6CFB6FFA3A" box="[1059,1159,1399,1424]" name="United States of America" pageId="46" pageNumber="47">America</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>