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<document id="AA708ECAA9F47924D65FF5F55F5624DC" ID-DOI="10.5281/zenodo.184427" ID-GBIF-Dataset="0acc8d0c-ebf6-4fca-a9f6-7285ce7abb2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="184427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459979950767" checkinUser="plazi" docAuthor="Caron, Edilson, Ribeiro-Costa, Cibele S. &amp; Newton, Alfred F." docDate="2008" docId="039F87B8FF82FFADFF16FF40FA3CF34A" docLanguage="en" docName="zt01895p009.pdf" docOrigin="Zootaxa 1895" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Piestus heterocephalus Fauvel 1902" docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFA6FFC0FF80FFA8FF81FFD7FFB4F218" masterDocTitle="New position of an abdominal defensive gland complex in Staphylinidae (Coleoptera) with redescription of Piestus heterocephalus Fauvel, 1902 (Piestinae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1698231768828" updateUser="plazi">
<mods:mods id="E0E58F9D8A685E9E814EBE007B616C9C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6D8756A18DFC7D785E7B44046DCA5715">New position of an abdominal defensive gland complex in Staphylinidae (Coleoptera) with redescription of Piestus heterocephalus Fauvel, 1902 (Piestinae)</mods:title>
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<mods:namePart id="CC5831465C0197155CF4283D9D87FA8B">Caron, Edilson</mods:namePart>
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<mods:namePart id="750579021D6EB59702B5938734BE9C61">Ribeiro-Costa, Cibele S.</mods:namePart>
</mods:name>
<mods:name id="3E2A994DD6F0F033BA7F8CA3FA4943B5" type="personal">
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<mods:namePart id="30A4F3EB6B1D6225181A1E3DAEDE5167">Newton, Alfred F.</mods:namePart>
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<mods:typeOfResource id="EE7C0F73E7F3CDE5F4D7D74B9A6F6C47">text</mods:typeOfResource>
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<mods:title id="83601666DCB9300A85BA976076ECC0EA">Zootaxa</mods:title>
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<mods:part id="71A4C44166C58FEF3146BDF23D73BA2D">
<mods:date id="8979B41BBF96F2E94FC2C36702D5EFF9">2008</mods:date>
<mods:detail id="7C684AF4328E0F709B300574D5A561E2" type="volume">
<mods:number id="C1D2F6764EA5BD5E255E9B11620F4D76">1895</mods:number>
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<mods:identifier id="E3D832FA510B1EC61BE5C3A531D3ABB3" type="DOI">10.5281/zenodo.184427</mods:identifier>
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<treatment id="039F87B8FF82FFADFF16FF40FA3CF34A" ID-DOI="http://doi.org/10.5281/zenodo.6230932" ID-GBIF-Taxon="119378499" ID-Zenodo-Dep="6230932" LSID="urn:lsid:plazi:treatment:039F87B8FF82FFADFF16FF40FA3CF34A" httpUri="http://treatment.plazi.org/id/039F87B8FF82FFADFF16FF40FA3CF34A" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="C32C6525FF82FFAAFF16FF40FE92F2CC" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8B8936AEFF82FFAAFF16FF40FDE5F2A9" blockId="2.[151,593,151,212]" box="[151,593,151,177]" pageId="2" pageNumber="3">
<heading id="D0C181C2FF82FFAAFF16FF40FDE5F2A9" bold="true" box="[151,593,151,177]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="4C364D2DFF82FFAAFF16FF40FDE5F2A9" ID-CoL="4HRXT" authority="Fauvel, 1902" authorityName="Fauvel" authorityYear="1902" box="[151,593,151,177]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF82FFAAFF16FF40FDE5F2A9" bold="true" box="[151,593,151,177]" pageId="2" pageNumber="3">
<emphasis id="B942EABCFF82FFAAFF16FF40FE18F2A9" bold="true" box="[151,428,151,177]" italics="true" pageId="2" pageNumber="3">Piestus heterocephalus</emphasis>
<bibRefCitation id="EFA74B5FFF82FFAAFE32FF40FDE5F2A9" author="Fauvel" box="[435,593,151,177]" pageId="2" pageNumber="8" refString="Fauvel, A. (1902) Staphylinides exotiques nouveaux. Revue d'Entomologie, 21, 8 - 37." type="journal article" year="1902">Fauvel, 1902</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF16FF6DFE92F2CC" blockId="2.[151,593,151,212]" box="[151,294,186,212]" pageId="2" pageNumber="3">
(
<figureCitation id="130D2A2BFF82FFAAFF1EFF6DFEA9F2CC" box="[159,285,186,212]" captionStart-0="FIGURES 1 8" captionStart-1="FIGURES 9 16" captionStart-2="FIGURE 17" captionStartId-0="3.[151,269,1547,1571]" captionStartId-1="4.[151,269,1568,1592]" captionStartId-2="5.[151,255,1263,1287]" captionTargetBox-0="[171,1424,549,1522]" captionTargetBox-1="[161,1429,362,1539]" captionTargetBox-2="[160,1422,411,1229]" captionTargetId-0="figure@3.[163,1424,542,1522]" captionTargetId-1="figure@4.[151,1436,354,1544]" captionTargetId-2="figure@5.[151,1436,394,1239]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." captionText-1="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." captionText-2="FIGURE 17. Piestus heterocephalus Fauvel, 1902. Sternites V VII dorsal view, tergites and left reservoir removed. Scale: 1 mm." httpUri-0="https://zenodo.org/record/184428/files/figure.png" httpUri-1="https://zenodo.org/record/184429/files/figure.png" httpUri-2="https://zenodo.org/record/184430/files/figure.png" pageId="2" pageNumber="3">Figs. 117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C6525FF82FFAAFF16FEDDFF49F399" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="8B8936AEFF82FFAAFF16FEDDFD44F359" blockId="2.[151,1437,266,385]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB261180FF82FFAAFF16FEDDFD44F359" pageId="2" pageNumber="3">
<taxonomicName id="4C364D2DFF82FFAAFF16FEDDFDE7F339" ID-CoL="4HRXT" authority="Fauvel, 1902: 22" authorityName="Fauvel" authorityPageNumber="22" authorityYear="1902" box="[151,595,266,289]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF82FFAAFF16FEDDFE24F339" box="[151,400,266,289]" italics="true" pageId="2" pageNumber="3">Piestus heterocephalus</emphasis>
<treatmentCitation id="0A9710BFFF82FFAAFE18FEDDFDE7F339" author="Fauvel" box="[409,595,266,289]" page="22" pageId="2" pageNumber="3" year="1902">
<bibRefCitation id="EFA74B5FFF82FFAAFE18FEDDFD9FF339" author="Fauvel" box="[409,555,266,289]" pageId="2" pageNumber="8" refString="Fauvel, A. (1902) Staphylinides exotiques nouveaux. Revue d'Entomologie, 21, 8 - 37." type="journal article" year="1902">Fauvel, 1902</bibRefCitation>
: 22
</treatmentCitation>
</taxonomicName>
(original description);
<treatmentCitation id="0A9710BFFF82FFAAFCD3FEDDFBE3F339" author="Blackwelder" box="[850,1111,266,289]" page="100" pageId="2" pageNumber="3" year="1944">
<bibRefCitation id="EFA74B5FFF82FFAAFCD3FEDDFB95F339" author="Blackwelder" box="[850,1057,266,289]" pageId="2" pageNumber="8" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. United States National Museum Bulletin, 185, xii + 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 100
</treatmentCitation>
(Neotropical checklist; error: Fauvel 1865);
<treatmentCitation id="0A9710BFFF82FFAAFEE3FEFDFD8AF359" author="Herman" box="[354,574,298,321]" page="1791" pageId="2" pageNumber="3" year="2001">
<bibRefCitation id="EFA74B5FFF82FFAAFEE3FEFDFE4AF359" author="Herman" box="[354,510,298,321]" pageId="2" pageNumber="9" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium, parts I - VII. Bulletin of the American Museum of Natural History, 167, 1 - 4218." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 1791
</treatmentCitation>
(world catalog).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF16FE9DFF49F399" blockId="2.[151,1437,266,385]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AB261180FF82FFAAFF16FE9DFF49F399" pageId="2" pageNumber="3">
<taxonomicName id="4C364D2DFF82FFAAFF16FE9DFCE6F379" ID-CoL="4HRXT" authority="Bernhauer &amp; Schubert, 1910: 7" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="7" authorityYear="1910" box="[151,850,330,353]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="heterocephalus" subGenus="Piestus">
<emphasis id="B942EABCFF82FFAAFF16FE9DFF50F379" box="[151,228,330,353]" italics="true" pageId="2" pageNumber="3">Piestus</emphasis>
(
<emphasis id="B942EABCFF82FFAAFF75FE9DFEF5F379" box="[244,321,330,353]" italics="true" pageId="2" pageNumber="3">Piestus</emphasis>
)
<emphasis id="B942EABCFF82FFAAFED0FE9DFE47F379" box="[337,499,330,353]" italics="true" pageId="2" pageNumber="3">heterocephalus</emphasis>
:
<treatmentCitation id="0A9710BFFF82FFAAFD83FE9DFCE6F379" author="Bernhauer" box="[514,850,330,353]" page="7" pageId="2" pageNumber="3" year="1910">
<bibRefCitation id="EFA74B5FFF82FFAAFD83FE9DFC8EF379" author="Bernhauer" box="[514,826,330,353]" pageId="2" pageNumber="8" refString="Bernhauer, M., &amp; Schubert, K. (1910) Staphylinidae I. In: S. Schenkling (Ed.), Coleopterorum Catalogus, Vol. 5. Junk, Berlin, pp. 1 - 86." type="book" year="1910">Bernhauer &amp; Schubert, 1910</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
(world catalog);
<treatmentCitation id="0A9710BFFF82FFAAFB8CFE9DFAB7F379" author="Scheerpeltz" box="[1037,1283,330,353]" page="285" pageId="2" pageNumber="3" year="1952">
<bibRefCitation id="EFA74B5FFF82FFAAFB8CFE9DFB64F379" author="Scheerpeltz" box="[1037,1232,330,353]" pageId="2" pageNumber="9" refString="Scheerpeltz, O. (1952) Revision der Gattung Piestus Gravh. (Coleoptera Staphylinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952</bibRefCitation>
: 285
</treatmentCitation>
(characters in key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32C6525FF82FFABFF16FE65FBE8F56F" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8B8936AEFF82FFAAFF16FE65FCA8F004" blockId="2.[151,1437,434,2020]" pageId="2" pageNumber="3">
<emphasis id="B942EABCFF82FFAAFF16FE65FEF8F3D4" bold="true" box="[151,332,434,460]" pageId="2" pageNumber="3">
<typeStatus id="548D880CFF82FFAAFF16FE65FF66F3D4" box="[151,210,434,460]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548D880CFF82FFAAFED6FE65FE70F3D4" box="[343,452,434,460]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
deposited at
<collectionCode id="ED27AE6BFF82FFAAFDE4FE65FD0AF3D4" box="[613,702,434,460]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="3">IRSNB</collectionCode>
, sex undetermined, labels: 1) “Merida /
<collectingCountry id="F321763EFF82FFAAFB21FE65FAA8F3D4" box="[1184,1308,434,460]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
” [Fauvels handwritten]; 2) “
<taxonomicName id="4C364D2DFF82FFAAFEE6FE0DFDADF3EB" box="[359,537,474,499]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF82FFAAFEE6FE0DFDADF3EB" box="[359,537,474,499]" italics="true" pageId="2" pageNumber="3">heterocephalus</emphasis>
</taxonomicName>
/ Fvl.” [Fauvels handwritten]; 3) “R.I.Sc.N.B. 17.479 /
<taxonomicName id="4C364D2DFF82FFAAFB2BFE0DFB4AF3EB" box="[1194,1278,474,499]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B942EABCFF82FFAAFB2BFE0DFB4AF3EB" box="[1194,1278,474,499]" italics="true" pageId="2" pageNumber="3">Piestus</emphasis>
</taxonomicName>
/ Coll. et det. A. Fauvel”; 4) “Ex-Typis” [white label, written in red].
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF44FDFDFC16F1E4" blockId="2.[151,1437,434,2020]" pageId="2" pageNumber="3">
<emphasis id="B942EABCFF82FFAAFF44FDFDFE08F05C" bold="true" box="[197,444,554,580]" pageId="2" pageNumber="3">Additional material</emphasis>
(7ɗɗ, 6ΨΨ, 14 sex undetermined).
<emphasis id="B942EABCFF82FFAAFCE2FDFDFB52F05C" bold="true" box="[867,1254,554,580]" pageId="2" pageNumber="3">
<collectingCountry id="F321763EFF82FFAAFCE2FDFDFBBAF05C" box="[867,1038,554,580]" name="Colombia" pageId="2" pageNumber="3">COLOMBIA</collectingCountry>
: Valle del Cauca:
</emphasis>
Ψ,
<specimenCount id="9D30FD27FF82FFAAFA91FDFDFA23F05C" box="[1296,1431,554,580]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Cali, no date, Fassl coll. (1
<collectionCode id="ED27AE6BFF82FFAAFE66FD85FDF7F074" LSID="urn:lsid:biocol.org:col:34925" box="[487,579,594,620]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
, 1Ψ
<collectionCode id="ED27AE6BFF82FFAAFDFCFD85FD79F074" box="[637,717,594,620]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFD65FD85FCDAF074" box="[740,878,594,620]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, ditto but
<date id="FF88106EFF82FFAAFC6BFD85FBC7F074" box="[1002,1139,594,620]" pageId="2" pageNumber="3" value="1938-06-30">30.VI.1938</date>
, Dybas coll. (
<collectionCode id="ED27AE6BFF82FFAAFA9AFD85FAC2F074" LSID="urn:lsid:biocol.org:col:34795" box="[1307,1398,594,620]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFA0CFD85FEBDF08C" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Cali, S. Antonio,
<quantity id="4CCE9B4BFF82FFAAFE7CFDADFDD4F08C" box="[509,608,634,660]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="m" value="2000.0">2000 m</quantity>
,
<date id="FF88106EFF82FFAAFDF3FDADFD6BF08C" box="[626,735,634,660]" pageId="2" pageNumber="3" value="1908-07">VII.1908</date>
, Fass coll. (
<collectionCode id="ED27AE6BFF82FFAAFCFFFDADFC6DF08C" LSID="urn:lsid:biocol.org:col:34795" box="[894,985,634,660]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFC75FDADFB37F08C" box="[1012,1155,634,660]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, ditto but
<date id="FF88106EFF82FFAAFA8CFDADFA28F08C" box="[1293,1436,634,660]" pageId="2" pageNumber="3" value="1908-07-21">21.VII.1908</date>
(
<collectionCode id="ED27AE6BFF82FFAAFF1EFD75FF45F0A4" box="[159,241,674,700]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFE89FD75FE27F0A4" box="[264,403,674,700]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, ditto but
<date id="FF88106EFF82FFAAFD95FD75FD17F0A4" box="[532,675,674,700]" pageId="2" pageNumber="3" value="1908-07-27">27.VII.1908</date>
(
<collectionCode id="ED27AE6BFF82FFAAFD37FD75FCBCF0A4" box="[694,776,674,700]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFC9EFD75FC1FF0A4" box="[799,939,674,700]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, ditto but
<date id="FF88106EFF82FFAAFBAAFD75FB3DF0A4" box="[1067,1161,674,700]" pageId="2" pageNumber="3" value="1909-03">III.1909</date>
(
<collectionCode id="ED27AE6BFF82FFAAFB1CFD75FB4CF0A4" LSID="urn:lsid:biocol.org:col:34795" box="[1181,1272,674,700]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
); ɗ, ditto but
<date id="FF88106EFF82FFAAFF16FD1DFF56F0FC" box="[151,226,714,740]" pageId="2" pageNumber="3" value="1909-01">I.1909</date>
(
<collectionCode id="ED27AE6BFF82FFAAFF70FD1DFEF8F0FC" LSID="urn:lsid:biocol.org:col:34795" box="[241,332,714,740]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFEE1FD1DFE5CF0FC" box="[352,488,714,740]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Cali, Villa Elvira,
<quantity id="4CCE9B4BFF82FFAAFD46FD1DFC96F0FC" box="[711,802,714,740]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FF88106EFF82FFAAFCACFD1DFC04F0FC" box="[813,944,714,740]" pageId="2" pageNumber="3" value="1908-07-06">6.VII.1908</date>
, Fass coll. (
<collectionCode id="ED27AE6BFF82FFAAFBB8FD1DFB20F0FC" LSID="urn:lsid:biocol.org:col:34795" box="[1081,1172,714,740]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFB29FD1DFA9BF0FC" box="[1192,1327,714,740]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, ditto but
<date id="FF88106EFF82FFAAFF16FD25FEA8F114" box="[151,284,754,780]" pageId="2" pageNumber="3" value="1908-09-12">12.IX.1908</date>
(
<collectionCode id="ED27AE6BFF82FFAAFEACFD25FECBF114" box="[301,383,754,780]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFE15FD25FDA9F114" box="[404,541,754,780]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Cali, Alto de las ances,
<date id="FF88106EFF82FFAAFCC5FD25FC1CF114" box="[836,936,754,780]" pageId="2" pageNumber="3" value="1908-11">XI.1908</date>
, Fass coll. (
<collectionCode id="ED27AE6BFF82FFAAFBB4FD25FB33F114" box="[1077,1159,754,780]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3">NMW</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFB1AFD25FA90F114" box="[1179,1316,754,780]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Cali, Rio Vitaco,
<quantity id="4CCE9B4BFF82FFAAFF72FCCDFEE4F12C" box="[243,336,794,820]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="3" unit="m" value="2200.0">2200 m</quantity>
,
<date id="FF88106EFF82FFAAFEDCFCCDFE7CF12C" box="[349,456,794,820]" pageId="2" pageNumber="3" value="1908-12">XII.1908</date>
Fassl coll. (
<collectionCode id="ED27AE6BFF82FFAAFDD9FCCDFD07F12C" LSID="urn:lsid:biocol.org:col:34795" box="[600,691,794,820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
); Ψ, Pischindé, no date, Rosemberg coll. (
<collectionCode id="ED27AE6BFF82FFAAFB27FCCDFAB7F12C" box="[1190,1283,794,820]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="3">IRSNB</collectionCode>
);
<emphasis id="B942EABCFF82FFAAFA99FCCDFEFEF144" bold="true" pageId="2" pageNumber="3">VENEZU- ELA: Mérida:</emphasis>
<specimenCount id="9D30FD27FF82FFAAFED0FC95FE6CF144" box="[337,472,834,860]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Mérida, no date and collector [“Ex-Typis”, see remarks below] (
<collectionCode id="ED27AE6BFF82FFAAFB4EFC95FA98F144" box="[1231,1324,834,860]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="3">IRSNB</collectionCode>
);
<emphasis id="B942EABCFF82FFAAFABEFC95FEDAF19C" bold="true" pageId="2" pageNumber="3">
<collectionCode id="ED27AE6BFF82FFAAFABEFC95FA28F144" box="[1343,1436,834,860]" pageId="2" pageNumber="3">ECUA-</collectionCode>
<collectionCode id="ED27AE6BFF82FFAAFF16FCBDFF69F19C" LSID="urn:lsid:biocol.org:col:12953" box="[151,221,874,900]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12953" name="Dorset County Museum" pageId="2" pageNumber="3">DOR</collectionCode>
: Pichincha:
</emphasis>
<specimenCount id="9D30FD27FF82FFAAFEF4FCBDFE49F19C" box="[373,509,874,900]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Calicali,
<quantity id="4CCE9B4BFF82FFAAFDF3FCBDFD79F19C" box="[626,717,874,900]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FF88106EFF82FFAAFD59FCBDFCDEF19C" box="[728,874,874,900]" pageId="2" pageNumber="3" value="1987-12-19">19.XII.1987</date>
, Huybensz coll. (
<collectionCode id="ED27AE6BFF82FFAAFBB4FCBDFB24F19C" LSID="urn:lsid:biocol.org:col:34795" box="[1077,1168,874,900]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<specimenCount id="9D30FD27FF82FFAAFB25FCBDFA98F19C" box="[1188,1324,874,900]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
, Tandapi,
<quantity id="4CCE9B4BFF82FFAAFF16FC45FE8FF1B4" box="[151,315,914,940]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="2" pageNumber="3" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">13001500 m</quantity>
,
<date id="FF88106EFF82FFAAFEC9FC45FE49F1B4" box="[328,509,914,940]" pageId="2" pageNumber="3" value="1965-06-15" valueMax="1965-06-20" valueMin="1965-06-15">1520.VI.1965</date>
, Pena coll. (
<collectionCode id="ED27AE6BFF82FFAAFD0DFC45FD5CF1B4" LSID="urn:lsid:biocol.org:col:34795" box="[652,744,914,940]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
);
<emphasis id="B942EABCFF82FFAAFD7DFC45FCCCF1B4" bold="true" box="[764,888,914,940]" pageId="2" pageNumber="3">Cotopaxi:</emphasis>
6ɗɗ, 4ΨΨ, Sigchos, Las Pampas Otonga Natural Reserve,
<date id="FF88106EFF82FFAAFEBAFC6DFE4EF1CC" box="[315,506,954,980]" pageId="2" pageNumber="3" value="2005-07-25" valueMax="2005-07-28" valueMin="2005-07-25">2528.VII.2005</date>
, W. Rossi coll. (
<collectionCode id="ED27AE6BFF82FFAAFD3CFC6DFCA4F1CC" box="[701,784,954,980]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="2" pageNumber="3">DZUP</collectionCode>
);
<emphasis id="B942EABCFF82FFAAFCA7FC6DFC32F1CC" bold="true" box="[806,902,954,980]" pageId="2" pageNumber="3">El Oro:</emphasis>
<specimenCount id="9D30FD27FF82FFAAFC0EFC6DFBA3F1CC" box="[911,1047,954,980]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
,
<quantity id="4CCE9B4BFF82FFAAFBA5FC6DFBDBF1CC" box="[1060,1135,954,980]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="km" value="12.0">12 km</quantity>
NW Atahualpa,
<quantity id="4CCE9B4BFF82FFAAFABAFC6DFA2CF1CC" box="[1339,1432,954,980]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="2" pageNumber="3" unit="m" value="1780.0">1780 m</quantity>
, wet forest,
<date id="FF88106EFF82FFAAFE9BFC35FE27F1E4" box="[282,403,994,1020]" pageId="2" pageNumber="3" value="1987-11-06">6.XI.1987</date>
, Young, Davidson &amp; Rawlins coll. (
<collectionCode id="ED27AE6BFF82FFAAFCBBFC35FC22F1E4" LSID="urn:lsid:biocol.org:col:34795" box="[826,918,994,1020]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF44FBDDFCEBF63C" blockId="2.[151,1437,434,2020]" box="[197,863,1034,1060]" pageId="2" pageNumber="3">
<emphasis id="B942EABCFF82FFAAFF44FBDDFECFF63C" bold="true" box="[197,379,1034,1060]" pageId="2" pageNumber="3">Redescription.</emphasis>
BL: 7.259.92 mm, BW: 1.802.40 mm.
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF44FBE5FD2DF684" blockId="2.[151,1437,434,2020]" pageId="2" pageNumber="3">
Body reddish dark brown to black with golden setae, apical half of abdominal segment VII and segments VIIIX yellowish (
<figureCitation id="130D2A2BFF82FFAAFEFCFB8DFE41F66C" box="[381,501,1114,1140]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
); antenna with apex of antennomere 11 slightly lighter than the others; tarsi, maxillary and labial palpi lighter than body.
</paragraph>
<paragraph id="8B8936AEFF82FFAAFF44FB7DFAF6F6F4" blockId="2.[151,1437,434,2020]" pageId="2" pageNumber="3">
Integument with undulate microstriae (
<figureCitation id="130D2A2BFF82FFAAFD11FB7DFD63F6DC" box="[656,727,1194,1220]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
), except on elytra; finely punctuate; larger punctures on lateral regions of metaventrite, on anterolateral regions of tergites and uniformly distributed on sternites.
</paragraph>
<paragraph id="8B8936AEFF82FFABFF44FB2DFD9DF3EA" blockId="2.[151,1437,434,2020]" lastBlockId="3.[151,1436,152,498]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Male. Head transverse (
<figureCitation id="130D2A2BFF82FFAAFE5AFB2DFD95F70C" box="[475,545,1274,1300]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
); front with two lateral long frontal processes, each almost reaching the dorsal tooth of mandibles (
<figureCitation id="130D2A2BFF82FFAAFE30FAF5FE4FF724" box="[433,507,1314,1340]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
), ventral face coarsely serrate, basal distance between the processes 2.0 times the width of each one (
<figureCitation id="130D2A2BFF82FFAAFE31FA9DFE4DF77C" box="[432,505,1354,1380]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
); vertex with anterior angles curved and prominent (
<figureCitation id="130D2A2BFF82FFAAFBF6FA9DFB75F77C" box="[1143,1217,1354,1380]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
); V-shaped sulcus incomplete with curved arms not joined medially (
<figureCitation id="130D2A2BFF82FFAAFD6FFAA5FC82F794" box="[750,822,1394,1420]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
); two moderate punctures with long setae on dorsal margin of the salient eyes, basal seta a little longer than apical seta (
<figureCitation id="130D2A2BFF82FFAAFC48FA4DFBA0F7AC" box="[969,1044,1434,1460]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Antennae somewhat geniculate and long, reaching the apex of elytra (
<figureCitation id="130D2A2BFF82FFAAFDD3FA15FD76F7C4" box="[594,706,1474,1500]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Figs. 1, 2</figureCitation>
); scape about 2.0 times the length of pedicel and with a tuft of five long setae on apical half of dorsal face. Labrum transverse (
<figureCitation id="130D2A2BFF82FFAAFC18FA3DFC57F41C" box="[921,995,1514,1540]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
), anterior margin slightly emarginate with six setae medially, equal in length; on lateral 1/3, the antero-internal seta with same length as posteroexternal seta. Epipharynx visible dorsally, about 2.0 times the median length of labrum (
<figureCitation id="130D2A2BFF82FFAAFB22F9EDFB5FF44C" box="[1187,1259,1594,1620]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
), deeply emarginate medially, with long fringes on internal margin and short fringes on external margin. Mandibles projected (
<figureCitation id="130D2A2BFF82FFAAFF6CF95DFE81F4BC" box="[237,309,1674,1700]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
), longer than the scape; slightly asymmetric (
<figureCitation id="130D2A2BFF82FFAAFCCBF95DFC15F4BC" box="[842,929,1674,1700]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
), each with bifurcate apex, the dorsal tooth shorter than the ventral; the internal border with one acute tooth at the middle and, anterior to it, in the right mandible with an area that resembles a tooth, absent in the left mandible. Mentum about 2.0 times wider than long. Pronotum somewhat wider than long (PW/PL=1.30) (
<figureCitation id="130D2A2BFF82FFAAFCCDF8D5FC3BF504" box="[844,911,1794,1820]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
), slightly convex (
<figureCitation id="130D2A2BFF82FFAAFBEAF8D5FB06F504" box="[1131,1202,1794,1820]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
), with parallel sides and abrupt constriction at basal 1/4 (
<figureCitation id="130D2A2BFF82FFAAFDBEF8FDFD37F55C" box="[575,643,1834,1860]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
); conspicuous longitudinal median sulcus not reaching anterior and posterior margins. Elytra truncate (
<figureCitation id="130D2A2BFF82FFAAFDBCF885FD30F574" box="[573,644,1874,1900]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
), together somewhat longer than wide (EL/BW=1.15); each with five longitudinal striae, sixth stria only on basal 2/3 of elytra; interstria 1 wider than the other ones. Abdominal sternite VI with very small emargination on each posterolateral area (external openings of abdominal defensive gland complex) (
<figureCitation id="130D2A2BFF82FFAAFE57F81DFDAAF5FC" box="[470,542,1994,2020]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
, arrow;
<figureCitation id="130D2A2BFF82FFAAFD01F81DFD66F5FC" box="[640,722,1994,2020]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
,
<figureCitation id="130D2A2BFF82FFAAFD5EF81DFD4AF5FC" box="[735,766,1994,2020]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="2" pageNumber="3">11</figureCitation>
); sternite VII with anterior margin broadly but shallowly projected above sternite VI (
<figureCitation id="130D2A2BFF83FFABFE75FF4FFDFEF2AA" box="[500,586,152,178]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="3" pageNumber="4">Figs. 6</figureCitation>
,
<figureCitation id="130D2A2BFF83FFABFDDAFF4FFDCDF2AA" box="[603,633,152,178]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">11</figureCitation>
), the projection immediately under the cell mass of the defensive gland (
<figureCitation id="130D2A2BFF83FFABFF69FF17FE8AF2C2" box="[232,318,192,218]" captionStart="FIGURE 17" captionStartId="5.[151,255,1263,1287]" captionTargetBox="[160,1422,411,1229]" captionTargetId="figure@5.[151,1436,394,1239]" captionTargetPageId="5" captionText="FIGURE 17. Piestus heterocephalus Fauvel, 1902. Sternites V VII dorsal view, tergites and left reservoir removed. Scale: 1 mm." httpUri="https://zenodo.org/record/184430/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
) and bearing some glandular pores (see separate description of gland complex below); apex of sternite VIII slightly curved; tergite X separating tergite IX in two equal parts, contiguous at base of tergite X; tergite IX with ventral struts short; tergite X with two pairs of setae, the apical pair the longest, about 2.0 times the length of the subapical (
<figureCitation id="130D2A2BFF83FFABFE6AFEEFFDF0F34A" box="[491,580,312,338]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
); sternite IX weakly pigmented, composed of a single median sclerite, symmetrical, 3 times longer than wide, basal margin acute, apical margin rounded, with two short setae on the apex. Aedeagus with median lobe curved in lateral view (
<figureCitation id="130D2A2BFF83FFABFCE9FE5FFC77F3BA" box="[872,963,392,418]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
), bulbous base (
<figureCitation id="130D2A2BFF83FFABFB10FE5FFB59F3BA" box="[1169,1261,392,418]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
); lateral lobes exceeding the apex of median lobe by 1/4 of its length in lateral view, with short setae apically and apex of external margin truncate (
<figureCitation id="130D2A2BFF83FFABFE44FE0FFDAFF3EA" box="[453,539,472,498]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF496626FF83FFABFF16F9DCFDCEF492" httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="3" pageNumber="4" targetBox="[171,1424,549,1522]" targetPageId="3">
<paragraph id="8B8936AEFF83FFABFF16F9DCFDCEF492" blockId="3.[151,1436,1547,1674]" pageId="3" pageNumber="4">
<emphasis id="B942EABCFF83FFABFF16F9DCFEF0F43B" bold="true" box="[151,324,1547,1571]" pageId="3" pageNumber="4">FIGURES 18.</emphasis>
<taxonomicName id="4C364D2DFF83FFABFECCF9DCFD69F43A" authority="Fauvel, 1902" authorityName="Fauvel" authorityYear="1902" box="[333,733,1547,1570]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF83FFABFECCF9DCFDF0F43A" box="[333,580,1547,1570]" italics="true" pageId="3" pageNumber="4">Piestus heterocephalus</emphasis>
Fauvel, 1902
</taxonomicName>
. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass.
</paragraph>
</caption>
<paragraph id="8B8936AEFF83FFABFF44F96AFBE8F56F" blockId="3.[151,1436,1725,2031]" pageId="3" pageNumber="4">
Female. Identical to male except for apex of sternite VIII slightly acute; tergite IX without ventral struts; area of sternite IX, which lacks a median sclerite and instead has an ovipositor consisting of a pair of very weakly pigmented hemisternites and a pair of more apical coxites with external margin somewhat rounded, and with a short seta and some microsetae on the apex; stylus absent (
<figureCitation id="130D2A2BFF83FFABFC5AF8E2FB80F557" box="[987,1076,1845,1871]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
). Spermathecal duct short and sclerotized at base (
<figureCitation id="130D2A2BFF83FFABFEFFF88AFE60F56F" box="[382,468,1885,1911]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="3" pageNumber="4">Fig. 16</figureCitation>
); spermatheca with basal half narrower than the apical.
</paragraph>
</subSubSection>
<subSubSection id="C32C6525FF83FFACFF44F852FD39F5C5" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8B8936AEFF83FFACFF44F852FB0DF332" blockId="3.[151,1436,1725,2031]" lastBlockId="4.[151,1437,152,298]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B942EABCFF83FFABFF44F852FE8AF587" bold="true" box="[197,318,1925,1951]" pageId="3" pageNumber="4">Remarks.</emphasis>
In the original description of
<taxonomicName id="4C364D2DFF83FFABFD2EF851FB8FF587" authority="Fauvel (1902)" authorityName="Fauvel" authorityYear="1902" box="[687,1083,1925,1951]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF83FFABFD2EF851FC35F587" box="[687,897,1926,1951]" italics="true" pageId="3" pageNumber="4">P. heterocephalus</emphasis>
,
<bibRefCitation id="EFA74B5FFF83FFABFC12F852FB8FF587" author="Fauvel" box="[915,1083,1925,1951]" pageId="3" pageNumber="8" refString="Fauvel, A. (1902) Staphylinides exotiques nouveaux. Revue d'Entomologie, 21, 8 - 37." type="journal article" year="1902">Fauvel (1902)</bibRefCitation>
</taxonomicName>
specified only one specimen studied (“Un seul exemplaire.”) and provided only data of locality (“
<collectingCountry id="F321763EFF83FFABFC64F87AFBD2F5DF" box="[997,1126,1965,1991]" name="Venezuela" pageId="3" pageNumber="4">Venezuela</collectingCountry>
: Merida.”). However, we received from IRSNB two specimens with locality label (“Merida /
<collectingCountry id="F321763EFF83FFABFC3BF802FB81F5F7" box="[954,1077,2005,2031]" name="Venezuela" pageId="3" pageNumber="4">Venezuela</collectingCountry>
”—presumably Fauvels handwritten), an additional museum label with information about Fauvels collection, and a
<typeStatus id="548D880CFF84FFACFB57FF4FFABDF2AA" box="[1238,1289,152,178]" pageId="4" pageNumber="5">type</typeStatus>
label (“Ex- Typis”), but only one specimen has an identification label (“
<taxonomicName id="4C364D2DFF84FFACFCF9FF17FB99F2C2" box="[888,1069,192,218]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="heterocephalus">heterocephalus</taxonomicName>
/ Fvl.”—presumably Fauvels handwritten). This was the main reason why we considered one specimen as the
<typeStatus id="548D880CFF84FFACFBC2FF3FFB1DF31A" box="[1091,1193,232,258]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(with Fauvels identification label) and the other as additional material (without Fauvels identification label).
</paragraph>
<caption id="DF496626FF84FFACFF16F9F7FD23F487" httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="4" pageNumber="5" targetBox="[161,1429,362,1539]" targetPageId="4">
<paragraph id="8B8936AEFF84FFACFF16F9F7FD23F487" blockId="4.[151,1436,1568,1695]" pageId="4" pageNumber="5">
<emphasis id="B942EABCFF84FFACFF16F9F7FEE5F420" bold="true" box="[151,337,1568,1592]" pageId="4" pageNumber="5">FIGURES 916.</emphasis>
<taxonomicName id="4C364D2DFF84FFACFED9F9F7FD51F42F" authority="Fauvel, 1902" authorityName="Fauvel" authorityYear="1902" box="[344,741,1568,1591]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF84FFACFED9F9F7FDFBF42F" box="[344,591,1568,1591]" italics="true" pageId="4" pageNumber="5">Piestus heterocephalus</emphasis>
Fauvel, 1902
</taxonomicName>
. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 1314) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25mm, except Fig. 11: 1mm.
</paragraph>
</caption>
<paragraph id="8B8936AEFF84FFACFF44F904FD39F5C5" blockId="4.[151,1436,1747,2013]" pageId="4" pageNumber="5">
This species is similar to
<emphasis id="B942EABCFF84FFACFE75F904FD25F4F4" box="[500,657,1747,1772]" italics="true" pageId="4" pageNumber="5">P. l a c o rd a i re i</emphasis>
<bibRefCitation id="EFA74B5FFF84FFACFD1DF904FCF6F4F5" author="Laporte" box="[668,834,1747,1773]" pageId="4" pageNumber="9" refString="Laporte, F. L. (Castelnau) (1835) Etudes entomologiques, ou description d'insectes nouveaux, et observations sur leur synonymie. Mequignon-Marvis, Paris, 159 pp." type="book" year="1835">Laporte, 1835</bibRefCitation>
mainly by the similar distance between the frontal processes of the head (2.0 times the width of each process). However,
<taxonomicName id="4C364D2DFF84FFACFC5CF92CFB1AF50C" box="[989,1198,1787,1812]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF84FFACFC5CF92CFB1AF50C" box="[989,1198,1787,1812]" italics="true" pageId="4" pageNumber="5">P. heterocephalus</emphasis>
</taxonomicName>
differs by the black body with apical half of abdominal segment VII and all of segments VIIIX yellowish (
<figureCitation id="130D2A2BFF84FFACFB4FF8F4FAF0F525" box="[1230,1348,1827,1853]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="4" pageNumber="5">Figs. 1, 2</figureCitation>
) (body entirely light brown to dark brown in
<taxonomicName id="4C364D2DFF84FFACFDCCF89CFD58F57C" box="[589,748,1867,1892]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lacordairei">
<emphasis id="B942EABCFF84FFACFDCCF89CFD58F57C" box="[589,748,1867,1892]" italics="true" pageId="4" pageNumber="5">P. lacordairei</emphasis>
</taxonomicName>
); ventral face of frontal processes of the head coarsely serrate (
<figureCitation id="130D2A2BFF84FFACFF52F8A4FEAEF595" box="[211,282,1907,1933]" captionStart="FIGURES 1 8" captionStartId="3.[151,269,1547,1571]" captionTargetBox="[171,1424,549,1522]" captionTargetId="figure@3.[163,1424,542,1522]" captionTargetPageId="3" captionText="FIGURES 1 8. Piestus heterocephalus Fauvel, 1902. 1) habitus, dorsal view; 2) habitus, lateral view; 3) head, dorsal view; 4) head, lateral view; 5) sternites VI and VII, ventrolateral view (arrow: external opening of the defensive gland); 6) sternites VI and VII, ventral view; 7) sternites VI and VII, dorsal view, tergites removed, micrograph; 8) micrograph showing details of reservoir and of cell mass." httpUri="https://zenodo.org/record/184428/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
) (only a short tooth on basal half in
<taxonomicName id="4C364D2DFF84FFACFD46F8A4FCD2F594" box="[711,870,1907,1932]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lacordairei">
<emphasis id="B942EABCFF84FFACFD46F8A4FCD2F594" box="[711,870,1907,1932]" italics="true" pageId="4" pageNumber="5">P. lacordairei</emphasis>
</taxonomicName>
); the labrum with six setae medially on anterior margin (
<figureCitation id="130D2A2BFF84FFACFF7AF84CFEF7F5AD" box="[251,323,1947,1973]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
) (four in
<taxonomicName id="4C364D2DFF84FFACFE38F84CFDEDF5AC" box="[441,601,1947,1972]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lacordairei">
<emphasis id="B942EABCFF84FFACFE38F84CFDEDF5AC" box="[441,601,1947,1972]" italics="true" pageId="4" pageNumber="5">P. lacordairei</emphasis>
</taxonomicName>
); and the lateral lobes of the aedeagus with truncate apex on external margin (
<figureCitation id="130D2A2BFF84FFACFF78F814FEFAF5C5" box="[249,334,1987,2013]" captionStart="FIGURES 9 16" captionStartId="4.[151,269,1568,1592]" captionTargetBox="[161,1429,362,1539]" captionTargetId="figure@4.[151,1436,354,1544]" captionTargetPageId="4" captionText="FIGURES 9 16. Piestus heterocephalus Fauvel, 1902. 9) labrum and epipharynx, left epiphayneal lobe and right setae removed; 10) mandibles, apex in dorsal view; 11) sternites VI and VII, ventral view, gently separated; 12) tergite X, apex, dorsal view; 13 14) aedeagus, 13) lateral view, 14) ventral view; 15) sternite IX, female, ventral view; 16) spermatheca. Scales: 0.25 mm, except Fig. 11: 1 mm." httpUri="https://zenodo.org/record/184429/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
) (curved in
<emphasis id="B942EABCFF84FFACFE5CF814FDCCF5C4" box="[477,632,1987,2012]" italics="true" pageId="4" pageNumber="5">P. l a c o rd a i re i</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C6525FF85FFADFF44FF4FFC25F332" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8B8936AEFF85FFADFF44FF4FFC25F332" blockId="5.[151,1436,152,338]" pageId="5" pageNumber="6">
<taxonomicName id="4C364D2DFF85FFADFF44FF4FFE6EF2A9" ID-CoL="4HRXT" box="[197,474,152,177]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="heterocephalus">
<emphasis id="B942EABCFF85FFADFF44FF4FFE6EF2A9" box="[197,474,152,177]" italics="true" pageId="5" pageNumber="6">Piestus heterocephalus</emphasis>
</taxonomicName>
may be confused with
<emphasis id="B942EABCFF85FFADFC8BFF4FFC3EF2A9" box="[778,906,152,177]" italics="true" pageId="5" pageNumber="6">P. b i c o r n i s</emphasis>
(
<bibRefCitation id="EFA74B5FFF85FFADFC22FF4FFBF2F2AA" author="Olivier" box="[931,1094,152,178]" pageId="5" pageNumber="9" refString="Olivier, A. G. (1811) Encyclopedie Methodique. Histoire naturelle. Insectes, 8. H. Agasse, Paris, pp. 361 - 722." type="journal article" year="1811">Olivier, 1811</bibRefCitation>
) or
<emphasis id="B942EABCFF85FFADFB02FF4FFB43F2A9" box="[1155,1271,152,177]" italics="true" pageId="5" pageNumber="6">P. v a l i d u s</emphasis>
<bibRefCitation id="EFA74B5FFF85FFADFA89FF4FFA28F2AA" author="Sharp" box="[1288,1436,152,178]" pageId="5" pageNumber="9" refString="Sharp, D. S. (1876) Contribution to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424." type="journal article" year="1876">Sharp, 1876</bibRefCitation>
because they share a similar body coloration. However, in
<emphasis id="B942EABCFF85FFADFCF6FF17FC42F2C1" box="[887,1014,192,217]" italics="true" pageId="5" pageNumber="6">P. b i c o r n i s</emphasis>
and
<emphasis id="B942EABCFF85FFADFBBFFF17FB06F2C1" box="[1086,1202,192,217]" italics="true" pageId="5" pageNumber="6">P. v a l i d u s</emphasis>
the basal distance between the frontal processes of the head is equal to the width of each process and the lateral lobes of the aedeagus are short, never exceeding the apex of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C32C6525FF85FFADFF44FEE0FA3CF34A" box="[197,1416,311,338]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8B8936AEFF85FFADFF44FEE0FA3CF34A" blockId="5.[151,1436,152,338]" box="[197,1416,311,338]" pageId="5" pageNumber="6">
<emphasis id="B942EABCFF85FFADFF44FEE0FED2F349" bold="true" box="[197,358,311,337]" pageId="5" pageNumber="6">Distribution.</emphasis>
Old records:
<collectingCountry id="F321763EFF85FFADFD87FEEFFDCAF34A" box="[518,638,312,338]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
(
<bibRefCitation id="EFA74B5FFF85FFADFD0CFEEFFC80F34A" author="Herman" box="[653,820,312,338]" pageId="5" pageNumber="9" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium, parts I - VII. Bulletin of the American Museum of Natural History, 167, 1 - 4218." type="journal article" year="2001">Herman 2001</bibRefCitation>
). New records:
<collectingCountry id="F321763EFF85FFADFC6DFEEFFBD7F34A" box="[1004,1123,312,338]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
,
<collectingCountry id="F321763EFF85FFADFBEEFEEFFB53F34A" box="[1135,1255,312,338]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
and
<collectingCountry id="F321763EFF85FFADFAA1FEEFFA30F34A" box="[1312,1412,312,338]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>