172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="86861C3BE1A60F668EC586610A26A585" ID-ISSN="0024-4082" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630456700959" checkinUser="felipe" docAuthor="Zaher, Hussam, Apesteguía, Sebastián & Scanferla, Carlos Agustín" docDate="2009" docId="039F87A4A57DFFFD230FF8FFFABE881A" docLanguage="en" docName="j.1096-3642.2009.00511.x.pdf" docOrigin="Zoological Journal of the Linnean Society 156" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Najash rionegrina Apesteguia & Zaher 2006" docType="treatment" docVersion="3" lastPageNumber="804" masterDocId="FFA6FFDCA57FFFFE2037FFEBFF92896D" masterDocTitle="The anatomy of the upper cretaceous snake Najash rionegrina Apesteguía & Zaher, 2006, and the evolution of limblessness in snakes" masterLastPageNumber="826" masterPageNumber="801" pageNumber="803" updateTime="1699277667141" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="B9F22484C2B54A44701C174A55436E4F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FA98E72AF85E3765E9AB5007C45C2414">
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<mods:title id="5DBFE30F97DC149FCEFB00B5B2EA3D1E">The anatomy of the upper cretaceous snake Najash rionegrina Apesteguía & Zaher, 2006, and the evolution of limblessness in snakes</mods:title>
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<mods:name id="F982312B285337764F9987320F09DD77" type="personal">
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<mods:roleTerm id="F151C28EE52BFD02BA54FB84173AC7DD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="32FFB33137A6712E19C1DA0BE4EED766">Zaher, Hussam</mods:namePart>
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</mods:name>
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<mods:roleTerm id="0522EE9731BCD3FD742937E910C09481">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B3A525A145A438E441DE3D3AF87E4D43">Apesteguía, Sebastián</mods:namePart>
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</mods:name>
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<mods:name id="E90B55FFF559F3A5C46AB2F62E6D50DF" type="personal">
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<mods:roleTerm id="1980AE0F72832589D0FEB3A0FC62D519">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6D8285A63A153636EF8A309C646A5814">Scanferla, Carlos Agustín</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DDE324882A29A56482F35EBBDA30D6EB">text</mods:typeOfResource>
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<mods:title id="F75F0A1354FEB401D2EE0C27C2679190">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="617CE5B809B36DB2E109917244AE89DB">
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<mods:date id="9DCC9260DE6BAC444F7452A0AAF8A1D8">2009</mods:date>
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<mods:detail id="1F1C899E5266684EE52A9EB28E21CDC1" type="volume">
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<mods:number id="B5BDE7DFE5B622BAB305DB9DBDC40192">156</mods:number>
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</mods:detail>
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<mods:extent id="44C0C15BE1C19E0AD5CEE3F4B335F070" unit="page">
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<mods:start id="8A489D04D798FCC99C3DECFE5D667EB3">801</mods:start>
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<mods:end id="31317B86A94B29DA294C62DD19D92BD7">826</mods:end>
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<mods:classification id="2895B9BDC37D9A90957D65CF431A58E3">journal article</mods:classification>
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<mods:identifier id="8BEDEF2BC4D1CDBA83F538BA99729714" type="ISSN">0024-4082</mods:identifier>
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</mods:mods>
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<treatment id="039F87A4A57DFFFD230FF8FFFABE881A" ID-DOI="http://doi.org/10.5281/zenodo.5492317" ID-Zenodo-Dep="5492317" LSID="urn:lsid:plazi:treatment:039F87A4A57DFFFD230FF8FFFABE881A" httpUri="http://treatment.plazi.org/id/039F87A4A57DFFFD230FF8FFFABE881A" lastPageId="3" lastPageNumber="804" pageId="2" pageNumber="803">
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<subSubSection id="C32C6539A57DFFFC230FF8FFFAE98E47" box="[824,1403,1812,1834]" pageId="2" pageNumber="803" type="nomenclature">
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<paragraph id="8B8936B2A57DFFFC230FF8FFFAE98E47" blockId="2.[824,1403,1812,1834]" box="[824,1403,1812,1834]" pageId="2" pageNumber="803">
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<heading id="D0C181DEA57DFFFC230FF8FFFAE98E47" box="[824,1403,1812,1834]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="803" reason="2">
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<taxonomicName id="4C364D31A57DFFFC230FF8FFFAE98E47" authority="APESTEGUIA & ZAHER, 2006" authorityName="Apesteguia & Zaher" authorityYear="2006" box="[824,1403,1812,1834]" class="Reptilia" family="Madtsoiidae" genus="Najash" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="821" phylum="Chordata" rank="species" species="rionegrina">
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<emphasis id="B942EAA0A57DFFFC230FF8FFFBB58E45" box="[824,1063,1812,1833]" italics="true" pageId="2" pageNumber="803">NAJASH RIONEGRINA</emphasis>
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<bibRefCitation id="EFA74B43A57DFFFC2406F8FFFAE98E47" author="Apesteguia S & Zaher H" box="[1073,1403,1812,1834]" pageId="2" pageNumber="803" pagination="1037 - 1040" refId="ref15566" refString="Apesteguia S, Zaher H. 2006. A Cretaceous terrestrial snake with robust hindlimbs and a sacrum. Nature 440: 1037 - 1040." type="journal article" year="2006">APESTEGUÍA & ZAHER, 2006</bibRefCitation>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32C6539A57DFFFD2311F8D0FABE881A" lastPageId="3" lastPageNumber="804" pageId="2" pageNumber="803" type="diagnosis">
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||
<paragraph id="8B8936B2A57DFFFC2311F8D0FC468E02" blockId="2.[806,1421,1850,1903]" pageId="2" pageNumber="803">
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<emphasis id="B942EAA0A57DFFFC2311F8D0FC0F8E3D" box="[806,925,1851,1872]" italics="true" pageId="2" pageNumber="803">Diagnosis:</emphasis>
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As for the genus, of which this is the only known species.
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</paragraph>
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<paragraph id="8B8936B2A57CFFFD2093FF2DFD968DFC" blockId="3.[164,779,198,1169]" pageId="3" pageNumber="804">
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<materialsCitation id="3B5E3CEFA57CFFFD2093FF2DFD968DFC" collectionCode="MPCA" country="Argentina" location="Museo Provincial Carlos Ameghino" municipality="Cipolletti" pageId="3" pageNumber="804" specimenCode="MPCA 390, MPCA 391, MPCA 392, MPCA 389, MPCA 393, MPCA 394, MPCA 395, MPCA 396, MPCA 397, MPCA 398, MPCA 399, MPCA 400" specimenCount="3" stateProvince="Rio Negro" typeStatus="holotype">
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||
<emphasis id="B942EAA0A57CFFFD2093FF2DFE9C89B6" box="[164,270,198,219]" italics="true" pageId="3" pageNumber="804">
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||
<typeStatus id="548D8810A57CFFFD2093FF2DFE9A89B6" box="[164,264,198,219]" pageId="3" pageNumber="804" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<location id="8EE96069A57CFFFD212EFF2DFD2589B1" LSID="urn:lsid:plazi:treatment:039F87A4A57DFFFD230FF8FFFABE881A:8EE96069A57CFFFD212EFF2DFD2589B1" box="[281,695,198,220]" country="Argentina" municipality="Cipolletti" name="Museo Provincial Carlos Ameghino" pageId="3" pageNumber="804" stateProvince="Rio Negro">Museo Provincial Carlos Ameghino</location>
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||
,
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<collectingMunicipality id="6BEDACC8A57CFFFD22F0FF2DFF478997" pageId="3" pageNumber="804">Cipolletti</collectingMunicipality>
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||
,
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<collectingRegion id="49F2F850A57CFFFD20DDFF0FFEF58997" box="[234,359,228,250]" country="Argentina" name="Rio Negro" pageId="3" pageNumber="804">Río Negro</collectingRegion>
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||
,
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<collectingCountry id="F3217622A57CFFFD214DFF0FFE7F8997" box="[378,493,228,250]" name="Argentina" pageId="3" pageNumber="804">Argentina</collectingCountry>
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(
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<collectionCode id="ED27AE77A57CFFFD2233FF0EFDC08996" box="[516,594,229,251]" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA</collectionCode>
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||
) 389–400. The
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<typeStatus id="548D8810A57CFFFD2093FEE8FE978874" box="[164,261,259,281]" pageId="3" pageNumber="804" type="holotype">holotype</typeStatus>
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consists of a series of associated materials, including a large fragment of the left dentary and anterior portion of the corresponding splenial (
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<specimenCode id="DB909EC9A57CFFFD2289FEABFF428818" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 390</specimenCode>
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), and a nearly complete and articulated postcranial skeleton, composed of 16 sections bearing a total of at least 122 articulated vertebrae (109 presacrals, two sacrals, and 11 caudals), a pelvic girdle, and hindlimbs. The
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<typeStatus id="548D8810A57CFFFD216DFE32FE298882" box="[346,443,473,495]" pageId="3" pageNumber="804" type="holotype">holotype</typeStatus>
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is represented by the following articulated and associated postcranial elements: an articulated section bearing the axis, four anterior presacral vertebrae, and the broken anterior part of the fifth vertebra (
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<specimenCode id="DB909EC9A57CFFFD21B1FDBFFD9B8B07" box="[390,521,596,618]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 391</specimenCode>
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); a section bearing a continuous string of 47 articulated presacral vertebrae, with associated ribs and two isolated vertebrae (
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<specimenCode id="DB909EC9A57CFFFD209CFD5BFEA08BAB" box="[171,306,688,710]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 392</specimenCode>
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), from which the proximal head of a middle presacral rib (
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<specimenCode id="DB909EC9A57CFFFD21AAFD25FD898B89" box="[413,539,718,740]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 389</specimenCode>
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) was prepared separately; three sections bearing no more than six, seven, and nine presacral vertebrae, respectively (
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||
<specimenCode id="DB909EC9A57CFFFD2289FCE0FF428A2D" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 393</specimenCode>
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||
); two sections bearing only associated ribs (
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||
<specimenCode id="DB909EC9A57CFFFD2289FCC1FF428A32" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 394</specimenCode>
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||
); one section bearing five articulated midpresacral vertebrae, and the broken anterior part of a sixth vertebra (
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||
<specimenCode id="DB909EC9A57CFFFD2168FC6DFE768AF6" box="[351,484,901,923]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 395</specimenCode>
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||
); two articulated caudal vertebrae (
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||
<specimenCode id="DB909EC9A57CFFFD2110FC4FFE3F8AD7" box="[295,429,932,954]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 396</specimenCode>
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||
); eight fragmentary sections bearing a total of 18 articulated presacral vertebrae (
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||
<specimenCode id="DB909EC9A57CFFFD209CFC09FEB88A95" box="[171,298,994,1016]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 397</specimenCode>
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||
); five undetermined and fragmentary elements (
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||
<specimenCode id="DB909EC9A57CFFFD20C9FBEBFE108D7B" box="[254,386,1024,1046]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 398</specimenCode>
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||
); several fragmentary associated presacral vertebrae (
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||
<specimenCode id="DB909EC9A57CFFFD21A5FBF4FD838D58" box="[402,529,1055,1077]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 399</specimenCode>
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||
); a section containing the pelvic girdle and hindlimb elements, articulated with eight posterior presacral, two sacral, and nine caudal vertebrae (
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||
<specimenCode id="DB909EC9A57CFFFD2140FB90FE6A8DFC" box="[375,504,1147,1169]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 400</specimenCode>
|
||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B8936B2A57CFFFD2093FB2CFD808FE4" blockId="3.[164,779,1223,1673]" pageId="3" pageNumber="804">
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||
<emphasis id="B942EAA0A57CFFFD2093FB2CFEE78DB1" box="[164,373,1223,1244]" italics="true" pageId="3" pageNumber="804">Referred material:</emphasis>
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||
<materialsCitation id="3B5E3CEFA57CFFFD2149FB2CFE918C54" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 385" specimenCount="1">
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Five specimens: (1) a fragmentary skull comprising the posterior half of the braincase, and associated vertebrae of a small specimen (
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<specimenCode id="DB909EC9A57CFFFD2289FAEFFF428C54" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 385</specimenCode>
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||
); (2)
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||
</materialsCitation>
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||
<materialsCitation id="3B5E3CEFA57CFFFD213BFAC8FEAE8CF8" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 380" specimenCount="1" typeStatus="holotype">
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||
several associated cranial and vertebral elements of a larger individual, probably twice as large as the
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||
<typeStatus id="548D8810A57CFFFD20C3FA8BFEC58C1B" box="[244,343,1376,1398]" pageId="3" pageNumber="804" type="holotype">holotype</typeStatus>
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||
, including an incomplete left dentary (
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||
<specimenCode id="DB909EC9A57CFFFD209CFA94FEA68CF8" box="[171,308,1407,1429]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 380</specimenCode>
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||
)
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B5E3CEFA57CFFFD2164FA94FE3E8CBF" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 381, MPCA 383" specimenCount="1" typeStatus="holotype">
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||
two undetermined cranial elements (possibly mandibular fragments) (
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||
<specimenCode id="DB909EC9A57CFFFD2270FA76FD5F8CDF" box="[583,717,1437,1458]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 381</specimenCode>
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||
and 382), axis (
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||
<specimenCode id="DB909EC9A57CFFFD2113FA57FE368CBF" box="[292,420,1468,1490]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 383</specimenCode>
|
||
)
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||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5E3CEFA57CFFFD218DFA57FDAA8C9D" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 384" specimenCount="1" typeStatus="holotype">
|
||
and associated presacral and caudal vertebrae (
|
||
<specimenCode id="DB909EC9A57CFFFD214BFA31FD928C9D" box="[380,512,1498,1520]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 384</specimenCode>
|
||
); (3)
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5E3CEFA57CFFFD2273FA30FF508F43" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 387" specimenCount="1" typeStatus="holotype">
|
||
a right quadrate and five associated presacral vertebrae (
|
||
<specimenCode id="DB909EC9A57CFFFD224AFA12FD6C8F62" box="[637,766,1529,1551]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 387</specimenCode>
|
||
); (4)
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5E3CEFA57CFFFD20FAF9FCFE068F06" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 386" specimenCount="1" typeStatus="holotype">
|
||
four associated sections of articulated vertebrae, and one fragment totaling 16 middle presacral vertebrae (
|
||
<specimenCode id="DB909EC9A57CFFFD20D2F9BEFEF18F06" box="[229,355,1621,1643]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 386</specimenCode>
|
||
); (5)
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5E3CEFA57CFFFD21ABF9BEFD9F8FE4" collectionCode="MPCA" pageId="3" pageNumber="804" specimenCode="MPCA 388" specimenCount="1" typeStatus="holotype">
|
||
a posterior presacral vertebra of a large individual (
|
||
<specimenCode id="DB909EC9A57CFFFD21B3F998FD978FE4" box="[388,517,1651,1673]" collectionCode="MPCA" country="Argentina" httpUri="http://grbio.org/cool/hjr9-dzvh" name="Museo Provincial &quot;Carlos Ameghino&quot;" pageId="3" pageNumber="804">MPCA 388</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8936B2A57CFFFD2093F92BFC5B8874" blockId="3.[164,779,1728,1903]" lastBlockId="3.[826,1441,198,281]" pageId="3" pageNumber="804">
|
||
<emphasis id="B942EAA0A57CFFFD2093F92BFEFC8FB8" box="[164,366,1728,1749]" italics="true" pageId="3" pageNumber="804">Stratum typicum:</emphasis>
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||
Mid to upper layers of the Candeleros Formation, basal unit of the
|
||
<collectingRegion id="49F2F850A57CFFFD2270F935FD3C8F99" box="[583,686,1758,1780]" country="Argentina" name="Neuquen" pageId="3" pageNumber="804">Neuquén</collectingRegion>
|
||
Group, early Upper Cretaceous. The bearing strata, about
|
||
<quantity id="4CCE9B57A57CFFFD2093F8F7FF4E8E5C" box="[164,220,1820,1841]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="804" unit="m" value="40.0">40 m</quantity>
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||
under the boundary with the overlying Huincul Formation, are considered as having been deposited around 92–94 Mya, during the Cenomanian. The fission track study on a tuff in the overlying Huincul Formation gave an age of 88 ± 3.9 Myr (
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||
<bibRefCitation id="EFA74B43A57CFFFD2577FF0FFC2F8874" author="Corbella H & Novas FE & Apesteguia S & Leanza HA" pageId="3" pageNumber="804" pagination="227 - 232" refId="ref15917" refString="Corbella H, Novas FE, Apesteguia S, Leanza HA 2004. First fission-track age for the dinosaur-bearing Neuquen Group (Upper Cretaceous), Neuquen basin, Argentina. Revista Museo Argentino de Ciencias Naturales n. s. 6: 227 - 232." type="book chapter" year="2004">
|
||
Corbella
|
||
<emphasis id="B942EAA0A57CFFFD230DFEEFFCE08875" box="[826,882,259,281]" italics="true" pageId="3" pageNumber="804">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8936B2A57CFFFD230DFEA8FABE881A" blockId="3.[826,1440,322,375]" pageId="3" pageNumber="804">
|
||
<emphasis id="B942EAA0A57CFFFD230DFEA8FC738835" box="[826,993,323,344]" italics="true" pageId="3" pageNumber="804">Locus typicus:</emphasis>
|
||
La Buitrera, Rentería Mesa,
|
||
<quantity id="4CCE9B57A57CFFFD256DFEA8FA328835" box="[1370,1440,322,344]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="804" unit="km" value="30.0">30 km</quantity>
|
||
north-west from the town of Cerro Policia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |