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<document id="F635EC52C0CD5341A914848D6C71C8C9" ID-DOI="10.11646/zootaxa.3986.4.6" ID-GBIF-Dataset="5e3f44de-9f03-4e9c-975a-a0a7c4fb347a" ID-ISSN="1175-5326" ID-Zenodo-Dep="241332" ID-ZooBank="B9A793F1-A67F-4856-AB27-EB2999FF7321" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264548758" checkinUser="plazi" docAuthor="Anlaş, Sinan" docDate="2015" docId="039487FAFFC9AA3DFF38FC3CF94EFE24" docLanguage="en" docName="zt03986p482.pdf" docOrigin="Zootaxa 3986 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astenus (Eurysunius) gusarovi Anlaş, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="475" masterDocId="FFADFF82FFC8AA3EFFAFFF8DFC0CFFD0" masterDocTitle="Four new species of Astenus (Eurysunius) Dejean, 1833 from Western Anatolia, Tur k ey (Coleoptera: Staphylinidae, Paederinae)" masterLastPageNumber="482" masterPageNumber="472" pageNumber="473" updateTime="1698610202434" updateUser="plazi">
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<mods:title id="92D94783536A7A6F1D2DF8EDD7731329">Four new species of Astenus (Eurysunius) Dejean, 1833 from Western Anatolia, Tur k ey (Coleoptera: Staphylinidae, Paederinae)</mods:title>
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<mods:namePart id="47E263498E037C09700996091D6AB153">Anlaş, Sinan</mods:namePart>
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<mods:date id="B35D1346B8A9FE67E4E14DB3C5FE2FC2">2015</mods:date>
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<treatment id="039487FAFFC9AA3DFF38FC3CF94EFE24" ID-DOI="http://doi.org/10.5281/zenodo.6096574" ID-GBIF-Taxon="119637576" ID-Zenodo-Dep="6096574" LSID="urn:lsid:plazi:treatment:039487FAFFC9AA3DFF38FC3CF94EFE24" httpUri="http://treatment.plazi.org/id/039487FAFFC9AA3DFF38FC3CF94EFE24" lastPageId="3" lastPageNumber="475" pageId="1" pageNumber="473">
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<heading id="D0CA8180FFC9AA3FFF38FC3CFE5BFC1B" bold="true" box="[151,599,945,971]" fontSize="11" level="1" pageId="1" pageNumber="473" reason="1">
<emphasis id="B949EAFEFFC9AA3FFF38FC3CFE5BFC1B" bold="true" box="[151,599,945,971]" pageId="1" pageNumber="473">
<taxonomicName id="4C3D4D6FFFC9AA3FFF38FC3CFE01FC1B" ID-CoL="HP3W" box="[151,525,945,971]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="473" phylum="Arthropoda" rank="species" species="gusarovi" status="sp. nov." subGenus="Eurysunius">
<emphasis id="B949EAFEFFC9AA3FFF38FC3CFCFBFC1B" bold="true" box="[151,247,945,971]" italics="true" pageId="1" pageNumber="473">Astenus</emphasis>
(
<emphasis id="B949EAFEFFC9AA3FFEA7FC3CFD98FC1B" bold="true" box="[264,404,945,971]" italics="true" pageId="1" pageNumber="473">Eurysunius</emphasis>
)
<emphasis id="B949EAFEFFC9AA3FFE0AFC3CFE01FC1B" bold="true" box="[421,525,945,971]" italics="true" pageId="1" pageNumber="473">gusarovi</emphasis>
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<taxonomicNameLabel id="A27A5785FFC9AA3FFDBBFC3CFE5BFC1B" box="[532,599,945,971]" pageId="1" pageNumber="473" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF38FC59FD9FFC3C" blockId="1.[151,599,945,1005]" box="[151,403,980,1005]" pageId="1" pageNumber="473">
(
<figureCitation id="13062A69FFC9AA3FFF30FC59FD07FC3C" box="[159,267,980,1005]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Figs. 18</figureCitation>
,
<figureCitation id="13062A69FFC9AA3FFEB9FC59FD39FC3C" box="[278,309,980,1004]" captionStart="FIGURES 33 38" captionStartId="8.[151,264,1565,1588]" captionTargetBox="[338,1272,215,1544]" captionTargetId="figure@8.[302,1284,193,1545]" captionTargetPageId="8" captionText="FIGURES 33 38. Aedeagi, lateral views. 33 — Astenus (Eurysunius) gusarovi sp. n.; 34 — A. (E.) orgeli sp. n.; 35 — A. (E.) kumlutasi sp. n.; 36 — A. (E.) honazicus sp. n.; 37 — A. (E.) occiduus Assing; 38 — A. (E.) sultanicus Assing. Scale bar: 50 µm (33 38)." httpUri="https://zenodo.org/record/241337/files/figure.png" pageId="1" pageNumber="473">33</figureCitation>
; Map 1)
</paragraph>
</subSubSection>
<subSubSection id="C3276567FFC9AA3FFF38FB98F89AFB49" pageId="1" pageNumber="473" type="materials_examined">
<paragraph id="8B8236ECFFC9AA3FFF38FB98F89AFB49" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
<emphasis id="B949EAFEFFC9AA3FFF38FB98FE51FBFE" bold="true" box="[151,605,1045,1070]" pageId="1" pageNumber="473">
<typeStatus id="5486884EFFC9AA3FFF38FB98FCDEFBFE" box="[151,210,1045,1070]" pageId="1" pageNumber="473">Type</typeStatus>
material.
<typeStatus id="5486884EFFC9AA3FFEF5FB98FDC7FBFE" box="[346,459,1045,1070]" pageId="1" pageNumber="473" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F32A767CFFC9AA3FFE71FB98FE5BFBFE" box="[478,599,1045,1070]" name="Turkey" pageId="1" pageNumber="473">TURKEY</collectingCountry>
:
</emphasis>
♂, “TR—Denizli province, Çal, Küçük Çökelez Mountain,
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,
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,
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,
<date id="FF83102CFFC9AA3FFE6DFBB4FE4DFB82" box="[450,577,1081,1106]" pageId="1" pageNumber="473" value="2014-04-15">15.IV.2014</date>
, leg. Anlaş /
<typeStatus id="5486884EFFC9AA3FFD45FBB4FF53FB82" box="[746,863,1081,1106]" pageId="1" pageNumber="473" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C3D4D6FFFC9AA3FFC22FBB7F85EFB81" box="[909,1106,1082,1105]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="473" phylum="Arthropoda" rank="species" species="gusarovi" status="sp. nov.">
<emphasis id="B949EAFEFFC9AA3FFC22FBB7F85EFB81" box="[909,1106,1082,1105]" italics="true" pageId="1" pageNumber="473">Astenus gusarovi</emphasis>
</taxonomicName>
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<emphasis id="B949EAFEFFC9AA3FFBF0FBB4F8A8FB82" bold="true" box="[1119,1188,1081,1106]" pageId="1" pageNumber="473">sp. n.</emphasis>
</taxonomicNameLabel>
det.
<bibRefCitation id="EFAC4B1DFFC9AA3FFB49FBB4F99DFB81" author="Anlas" box="[1254,1425,1081,1106]" pageId="1" pageNumber="473" refString="Anlas, S. (2014) A new species of Astenus (Eurysunius) Dejean, from Turkey (Coleoptera: Staphylinidae, Paederinae). Turkish Journal of Entomology, 38 (3), 239 - 243." type="journal article" year="2014">S. Anlaş 2014</bibRefCitation>
” (
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).
<emphasis id="B949EAFEFFC9AA3FFEA0FBD1FE1AFBA5" bold="true" box="[271,534,1116,1141]" pageId="1" pageNumber="473">
<typeStatus id="5486884EFFC9AA3FFEA0FBD1FD87FBA5" box="[271,395,1116,1141]" pageId="1" pageNumber="473" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F32A767CFFC9AA3FFE37FBD1FE1DFBA5" box="[408,529,1116,1141]" name="Turkey" pageId="1" pageNumber="473">TURKEY</collectingCountry>
:
</emphasis>
<specimenCount id="9D3BFD65FFC9AA3FFDB1FBD0FE48FBA5" box="[542,580,1117,1141]" pageId="1" pageNumber="473" type="generic">2♂</specimenCount>
, 1♀, same data as
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;
<specimenCount id="9D3BFD65FFC9AA3FFC25FBD0FFA3FBA5" box="[906,943,1117,1141]" pageId="1" pageNumber="473" type="generic">2♂</specimenCount>
, 3♀, Denizli province, Çal, Büyük Çökelez Mountain,
<geoCoordinate id="EE09502BFFC9AA3FFEBDFB0CFD9BFB49" box="[274,407,1153,1177]" direction="north" orientation="latitude" pageId="1" pageNumber="473" precision="15" value="38.025276">38°01'31&quot;N</geoCoordinate>
,
<geoCoordinate id="EE09502BFFC9AA3FFE0BFB0DFE2BFB49" box="[420,551,1152,1177]" direction="east" orientation="longitude" pageId="1" pageNumber="473" precision="15" value="29.362778">29°21'46&quot;E</geoCoordinate>
,
<quantity id="4CC59B09FFC9AA3FFD9DFB0DFE85FB48" box="[562,649,1152,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.627" pageId="1" pageNumber="473" unit="m" value="1627.0">1627 m</quantity>
,
<date id="FF83102CFFC9AA3FFD3AFB0DFF18FB49" box="[661,788,1152,1177]" pageId="1" pageNumber="473" value="2014-04-15">15.IV.2014</date>
, leg. Anlaş &amp; Örgel (all
<collectionCode id="ED2CAE29FFC9AA3FFB81FB0CF886FB49" box="[1070,1162,1153,1177]" pageId="1" pageNumber="473">AZMM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276567FFC9AA3FFF68FB29F90CFB6D" box="[199,1280,1188,1213]" pageId="1" pageNumber="473" type="etymology">
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<emphasis id="B949EAFEFFC9AA3FFF68FB29FD42FB6D" bold="true" box="[199,334,1188,1213]" pageId="1" pageNumber="473">Etymology.</emphasis>
The species is dedicated to Dr. Vladimir Gusarov, Oslo, a specialist on staphylinids.
</paragraph>
</subSubSection>
<subSubSection id="C3276567FFC9AA3CFF68FB4AFE92F91C" lastPageId="2" lastPageNumber="474" pageId="1" pageNumber="473" type="description">
<paragraph id="8B8236ECFFC9AA3FFF68FB4AFEDBFA9C" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
<emphasis id="B949EAFEFFC9AA3FFF68FB4AFD55FB30" bold="true" box="[199,345,1223,1248]" pageId="1" pageNumber="473">Description.</emphasis>
Measurements (in mm) and ratios (range, n=9): AL 0.920.98; HL 0.550.64; HW 0.670.73; PW 0.650.72; PL 0.560.60; EL 0.430.46; EW 0.700.75; AW 0.780.82; TiL 0.470.51; TaL 0.290.31; ML 0.580.61 (n=5); TL 3.94.5 HL/HW 0.820.88; PW/HW 0.970.99; PW/PL 1.161.20; EL/PL 0.77; EW/PW 1.041.08; AW/EW 1.091.12; TiL/TaL 1.621.65.
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF68FADBFD78FA67" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
Habitus as in
<figureCitation id="13062A69FFC9AA3FFEC7FADBFDA1FABF" box="[360,429,1366,1391]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Fig. 1</figureCitation>
. Coloration: head and pronotum reddish brown, elytra yellowish brown, with area near scutellum, anterior margin, and anterior part of lateral margin infuscate, abdomen dark brown, antennae rufous, legs reddish brown.
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF68FA4FFECBF996" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
Head transverse (see
<figureCitation id="13062A69FFC9AA3FFE6CFA4FFE3EFA0A" box="[451,562,1474,1499]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Figs. 12</figureCitation>
); dorsal surface distinctly convex; with very dense and coarse punctation; microsculpture indistinct; pubescence short and yellowish. Eyes relatively small and weakly prominent, in dorsal view temples longer than eyes. Antennae moderately slender, antennomere III approximately 2.1 times as long as wide; antennomeres VX slightly oblong (
<figureCitation id="13062A69FFC9AA3FFDDBF9A0FEB4F996" box="[628,696,1581,1606]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF68F9DFF868F928" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
Pronotum transverse and slightly narrower than head (see
<figureCitation id="13062A69FFC9AA3FFCD7F9DCFFEBF9B9" box="[888,999,1617,1642]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Figs. 12</figureCitation>
); widest at anterior angles, distinctly narrowed posteriorly; anterior and posterior angles each with a long seta of little more than half length of lateral margin of pronotum; posterior margin convex; dorsal surface with slightly pronounced impressions; microsculpture almost absent; punctation similar to that of head, but denser, surface somewhat glossier than that of head; pubescence of similar length as that of head, but less fine and more conspicuous.
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF68F88EFFD7F857" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
Elytra transversely rectangular and short, slightly wider than pronotum (see ratios and
<figureCitation id="13062A69FFC9AA3FFAB8F88EF981F8CC" box="[1303,1421,1795,1820]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Figs. 12</figureCitation>
); microsculpture absent; punctation very dense and distinctly granulose, interstices narrower than diameter of punctures; pubescence yellowish, more distinct than that of head and pronotum; posterior margin with several dark brown setae, the longest near posterior angles.
<collectingCountry id="F32A767CFFC9AA3FFD0BF8E3FED0F857" box="[676,732,1902,1927]" name="India" pageId="1" pageNumber="473">Hind</collectingCountry>
wings totally reduced.
</paragraph>
<paragraph id="8B8236ECFFC9AA3FFF68F81FF89EF823" blockId="1.[151,1437,1045,2035]" pageId="1" pageNumber="473">
Abdomen slightly wider than elytra (see
<figureCitation id="13062A69FFC9AA3FFD3BF81FFED9F87B" box="[660,725,1938,1963]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="1" pageNumber="473">Fig. 1</figureCitation>
), widest at segment V, segments IIIVI moderately transverse, tergites IIIV approximately twice as wide as long; punctation very dense and very fine; interstices with distinct microsculpture; pubescence yellowish; posterior margin of tergite VII with palisade fringe.
</paragraph>
<paragraph id="8B8236ECFFCAAA3CFF68FF1AFEE0FF28" blockId="2.[151,1436,151,248]" pageId="2" pageNumber="474">
Male: sternite VII in posterior median area slightly depressed and with some modified dark stout setae, posterior margin weakly concave (
<figureCitation id="13062A69FFCAAA3CFDB4FF31FE6DFF04" box="[539,609,188,213]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="2" pageNumber="474">Fig. 4</figureCitation>
); sternite VIII deeply and acutely incised at posterior margin, pubescence unmodified (
<figureCitation id="13062A69FFCAAA3CFE88FF52FD67FF28" box="[295,363,223,248]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="2" pageNumber="474">Fig. 3</figureCitation>
); aedeagus as in
<figureCitation id="13062A69FFCAAA3CFD85FF52FE9EFF28" box="[554,658,223,248]" captionStart="FIGURES 1 8" captionStartId="2.[151,264,1520,1543]" captionTargetBox="[369,1214,301,1477]" captionTargetId="figure@2.[350,1236,301,1499]" captionTargetPageId="2" captionText="FIGURES 1 8. Details of A stenus (Eurysunius) gusarovi sp. n. 1 — habitus; 2 — forebody; 3 — male sternite VIII; 4 — male sternite VII; 5 — antenna; 6 and 7 — aedeagus, lateral view; 8 — aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (3 8)." httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="2" pageNumber="474">Figs. 68</figureCitation>
and
<figureCitation id="13062A69FFCAAA3CFD66FF6DFEE4FF28" box="[713,744,224,248]" captionStart="FIGURES 33 38" captionStartId="8.[151,264,1565,1588]" captionTargetBox="[338,1272,215,1544]" captionTargetId="figure@8.[302,1284,193,1545]" captionTargetPageId="8" captionText="FIGURES 33 38. Aedeagi, lateral views. 33 — Astenus (Eurysunius) gusarovi sp. n.; 34 — A. (E.) orgeli sp. n.; 35 — A. (E.) kumlutasi sp. n.; 36 — A. (E.) honazicus sp. n.; 37 — A. (E.) occiduus Assing; 38 — A. (E.) sultanicus Assing. Scale bar: 50 µm (33 38)." httpUri="https://zenodo.org/record/241337/files/figure.png" pageId="2" pageNumber="474">33</figureCitation>
.
</paragraph>
<caption id="DF426664FFCAAA3CFF38FA7DFD0EF994" httpUri="https://zenodo.org/record/241333/files/figure.png" pageId="2" pageNumber="474" targetBox="[369,1214,301,1477]" targetPageId="2">
<paragraph id="8B8236ECFFCAAA3CFF38FA7DFD0EF994" blockId="2.[151,1436,1520,1604]" pageId="2" pageNumber="474">
<emphasis id="B949EAFEFFCAAA3CFF38FA7DFD31F9D5" bold="true" box="[151,317,1520,1543]" pageId="2" pageNumber="474">FIGURES 18.</emphasis>
Details of A
<emphasis id="B949EAFEFFCAAA3CFE66FA7FFE04F9D7" box="[457,520,1522,1543]" italics="true" pageId="2" pageNumber="474">stenus</emphasis>
<taxonomicName id="4C3D4D6FFFCAAA3CFDBEFA7DFEF9F9D6" box="[529,757,1520,1542]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="gusarovi" status="sp. nov." subGenus="Eurysunius">
(
<emphasis id="B949EAFEFFCAAA3CFDB5FA7CFE80F9D6" box="[538,652,1521,1542]" italics="true" pageId="2" pageNumber="474">Eurysunius</emphasis>
)
<emphasis id="B949EAFEFFCAAA3CFD32FA7CFEF9F9D6" box="[669,757,1521,1542]" italics="true" pageId="2" pageNumber="474">gusarovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A5785FFCAAA3CFD51FA7CFF36F9D6" box="[766,826,1521,1542]" pageId="2" pageNumber="474" rank="species">
<emphasis id="B949EAFEFFCAAA3CFD51FA7CFF36F9D6" bold="true" box="[766,826,1521,1542]" pageId="2" pageNumber="474">sp. n.</emphasis>
</taxonomicNameLabel>
1—habitus; 2—forebody; 3—male sternite VIII; 4—male sternite VII; 5—antenna; 6 and 7—aedeagus, lateral view; 8—aedeagus, ventral view. Scale bar: 1.0 mm (1); 0.5 mm (2); 0.2 mm (38).
</paragraph>
</caption>
<paragraph id="8B8236ECFFCAAA3CFF68F9E6FE92F91C" blockId="2.[151,1436,1643,2028]" pageId="2" pageNumber="474">
<emphasis id="B949EAFEFFCAAA3CFF68F9E6FDB8F954" bold="true" box="[199,436,1643,1668]" pageId="2" pageNumber="474">Comparative notes.</emphasis>
The species is distinguished from all its consubgeners by the male sexual characters, especially by the morphology of the aedeagus. In addition, the new species differs from the seven Turkish species described previously by a different coloration:
</paragraph>
</subSubSection>
<subSubSection id="C3276567FFCAAA3DFF68F955F878FEB4" lastPageId="3" lastPageNumber="475" pageId="2" pageNumber="474" type="reference_group">
<paragraph id="8B8236ECFFCAAA3DFF68F955F878FEB4" blockId="2.[151,1436,1643,2028]" lastBlockId="3.[151,1436,151,501]" lastPageId="3" lastPageNumber="475" pageId="2" pageNumber="474">
In
<taxonomicName id="4C3D4D6FFFCAAA3CFF44F955FE98F920" ID-CoL="5W6LM" authority="Assing, 2002" authorityName="Assing" authorityYear="2002" box="[235,660,1751,1776]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="bicoloratus">
<emphasis id="B949EAFEFFCAAA3CFF44F955FD9CF920" box="[235,400,1751,1776]" italics="true" pageId="2" pageNumber="474">A. bicoloratus</emphasis>
<bibRefCitation id="EFAC4B1DFFCAAA3CFE32F95AFE3BF920" author="Assing" box="[413,567,1751,1776]" pageId="2" pageNumber="474" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing, 2002</bibRefCitation>
(Ordu)
</taxonomicName>
: head and pronotum dark brown, anterior 2/3 of elytra blackish, posterior 1/3 of elytra yellow, abdomen blackish with posterior margins of anterior tergites and apex lighter (
<bibRefCitation id="EFAC4B1DFFCAAA3CFF31F892FD3AF8E8" author="Assing" box="[158,310,1823,1848]" pageId="2" pageNumber="474" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing 2002</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCAAA3CFEC1F8ADFEFAF8E8" ID-CoL="HP3F" authority="Assing, 2011" authorityName="Assing" authorityYear="2011" box="[366,758,1823,1848]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="brachati">
<emphasis id="B949EAFEFFCAAA3CFEC1F8ADFDFDF8E8" box="[366,497,1823,1848]" italics="true" pageId="2" pageNumber="474">A. brachati</emphasis>
<bibRefCitation id="EFAC4B1DFFCAAA3CFE53F892FE9FF8E8" author="Assing" box="[508,659,1823,1848]" pageId="2" pageNumber="474" refString="Assing, V. (2011) On the Staphylinidae of Turkey VIII. Eleven new species, two new synonymies, a new combination, and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 81, 179 - 227." type="journal article" year="2011">Assing, 2011</bibRefCitation>
(Bursa)
</taxonomicName>
: body blackish, with posterior margins of elytra narrowly yellowish (
<bibRefCitation id="EFAC4B1DFFCAAA3CFEBDF8CEFDAFF88C" author="Assing" box="[274,419,1859,1884]" pageId="2" pageNumber="474" refString="Assing, V. (2011) On the Staphylinidae of Turkey VIII. Eleven new species, two new synonymies, a new combination, and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 81, 179 - 227." type="journal article" year="2011">Assing 2011</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCAAA3CFE7CF8C9FF53F88C" ID-CoL="HP4F" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[467,863,1859,1884]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="occiduus">
<emphasis id="B949EAFEFFCAAA3CFE7CF8C9FE5AF88C" box="[467,598,1859,1884]" italics="true" pageId="2" pageNumber="474">A. occiduus</emphasis>
<bibRefCitation id="EFAC4B1DFFCAAA3CFDF3F8CEFEE3F88C" author="Assing" box="[604,751,1859,1884]" pageId="2" pageNumber="474" refString="Assing, V. (2007) New species and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 700, 1 - 64." type="journal article" year="2007">Assing, 2007</bibRefCitation>
(Denizli)
</taxonomicName>
: forebody blackish, with posterior 1/41/3 of elytra yellow, abdomen blackish with narrow posterior margins of tergites and apex somewhat paler (
<bibRefCitation id="EFAC4B1DFFCAAA3CFB6FF8EAF958F850" author="Assing" box="[1216,1364,1895,1920]" pageId="2" pageNumber="474" refString="Assing, V. (2007) New species and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 700, 1 - 64." type="journal article" year="2007">Assing 2007</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCAAA3CFA2BF8E5FDDBF874" ID-CoL="HP4L" authority="Assing, 2002" authorityName="Assing" authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="paphlagonicus">
<emphasis id="B949EAFEFFCAAA3CFA2BF8E5FD30F874" italics="true" pageId="2" pageNumber="474">A. paphlagonicus</emphasis>
<bibRefCitation id="EFAC4B1DFFCAAA3CFEECF806FDDBF874" author="Assing" box="[323,471,1931,1956]" pageId="2" pageNumber="474" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing, 2002</bibRefCitation>
</taxonomicName>
(Kastamonu, Bolu): head, pronotum, and anterior half of elytra dark brown, posterior half of elytra yellow, abdomen dark brown with posterior margins of tergites and apex slightly lighter (
<bibRefCitation id="EFAC4B1DFFCAAA3CFAE2F822FCD8F83C" author="Assing" pageId="2" pageNumber="474" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing 2002</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCAAA3CFEA4F859FEB0F83C" ID-CoL="HP4X" authority="Assing, 2002" authorityName="Assing" authorityYear="2002" box="[267,700,2003,2028]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="474" phylum="Arthropoda" rank="species" species="sexsetosus">
<emphasis id="B949EAFEFFCAAA3CFEA4F859FDAEF83B" box="[267,418,2004,2027]" italics="true" pageId="2" pageNumber="474">A. sexsetosus</emphasis>
<bibRefCitation id="EFAC4B1DFFCAAA3CFE03F85EFE4FF83C" author="Assing" box="[428,579,2003,2028]" pageId="2" pageNumber="474" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing, 2002</bibRefCitation>
(Kayseri)
</taxonomicName>
: head, pronotum, and abdomen, except for the lighter posterior tergal margins, blackish brown, elytra yellow, with area near scutellum, anterior margin, and anterior part of lateral margin more or less infuscate (
<bibRefCitation id="EFAC4B1DFFCBAA3DFDA1FF31FEABFF04" author="Assing" box="[526,679,188,213]" pageId="3" pageNumber="475" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing 2002</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCBAA3DFD4CFF31F82CFF04" ID-CoL="HP52" authority="Assing, 2010" authorityName="Assing" authorityYear="2010" box="[739,1056,188,213]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="species" species="sultanicus">
<emphasis id="B949EAFEFFCBAA3DFD4CFF31FF76FF05" box="[739,890,188,213]" italics="true" pageId="3" pageNumber="475">A. sultanicus</emphasis>
<bibRefCitation id="EFAC4B1DFFCBAA3DFC29FF31F82CFF04" author="Assing" box="[902,1056,188,213]" pageId="3" pageNumber="475" refString="Assing, V. (2010) On the Staphylinidae of Turkey. VII. Five new species and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 80, 71 - 102." type="journal article" year="2010">Assing, 2010</bibRefCitation>
</taxonomicName>
(Afyonkarahisar, Konya): head, pronotum, and abdomen blackish, with posterior margins of abdominal segments narrowly reddish; elytra yellowish, with anterior margin and scutellar area narrowly, diffusely, and weakly infuscate (
<bibRefCitation id="EFAC4B1DFFCBAA3DFB78FE89F963FECC" author="Assing" box="[1239,1391,260,285]" pageId="3" pageNumber="475" refString="Assing, V. (2010) On the Staphylinidae of Turkey. VII. Five new species and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 80, 71 - 102." type="journal article" year="2010">Assing 2010</bibRefCitation>
); in
<taxonomicName id="4C3D4D6FFFCBAA3DFF38FEA5FEDBFE90" ID-CoL="HP4R" authority="Anlas, 2014" authorityName="Anlas" authorityYear="2014" box="[151,727,295,320]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="species" species="sandiklicus">
<emphasis id="B949EAFEFFCBAA3DFF38FEA5FD79FE90" box="[151,373,295,320]" italics="true" pageId="3" pageNumber="475">Astenus sandiklicus</emphasis>
<bibRefCitation id="EFAC4B1DFFCBAA3DFED1FEAAFE04FE90" author="Anlas" box="[382,520,295,320]" pageId="3" pageNumber="475" refString="Anlas, S. (2014) A new species of Astenus (Eurysunius) Dejean, from Turkey (Coleoptera: Staphylinidae, Paederinae). Turkish Journal of Entomology, 38 (3), 239 - 243." type="journal article" year="2014">Anlaş, 2014</bibRefCitation>
(Afyonkarahisar)
</taxonomicName>
: head, pronotum and elytra reddish brown, abdomen blackish brown with narrow posterior margins of tergites and apex somewhat paler (
<bibRefCitation id="EFAC4B1DFFCBAA3DFC4EFEC1F864FEB4" author="Anlas" box="[993,1128,332,357]" pageId="3" pageNumber="475" refString="Anlas, S. (2014) A new species of Astenus (Eurysunius) Dejean, from Turkey (Coleoptera: Staphylinidae, Paederinae). Turkish Journal of Entomology, 38 (3), 239 - 243." type="journal article" year="2014">Anlaş 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276567FFCBAA3DFF68FEE2F94EFE24" pageId="3" pageNumber="475" type="distribution">
<paragraph id="8B8236ECFFCBAA3DFF68FEE2F94EFE24" blockId="3.[151,1436,151,501]" pageId="3" pageNumber="475">
<emphasis id="B949EAFEFFCBAA3DFF68FEE2FE17FE58" bold="true" box="[199,539,367,392]" pageId="3" pageNumber="475">Distribution and bionomics.</emphasis>
The new species was collected in two localities in the Çökelez Mountain, in Denizli province, in grassland at an altitude of about
<quantity id="4CC59B09FFCBAA3DFC86FE19FF84FE7C" box="[809,904,404,429]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="475" unit="m" value="1600.0">1600 m</quantity>
. The specimens were found in the nests of
<taxonomicName id="4C3D4D6FFFCBAA3DFF38FE34FD24FE00" box="[151,296,441,464]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAFEFFCBAA3DFF38FE34FD24FE00" box="[151,296,441,464]" italics="true" pageId="3" pageNumber="475">Tetramorium</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C3D4D6FFFCBAA3DFED3FE35FE17FE00" box="[380,539,440,464]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
:
<taxonomicName id="4C3D4D6FFFCBAA3DFD9AFE3AFEB5FE00" box="[565,697,439,464]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
: Myrmicinae), which confirms previous observations that myrmecophilous
<taxonomicName id="4C3D4D6FFFCBAA3DFEF5FE50FDD4FE24" box="[346,472,477,500]" class="Insecta" family="Staphylinidae" genus="Eurysunius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAFEFFCBAA3DFEF5FE50FDD4FE24" box="[346,472,477,500]" italics="true" pageId="3" pageNumber="475">Eurysunius</emphasis>
</taxonomicName>
is exclusively associated with
<taxonomicName id="4C3D4D6FFFCBAA3DFC9DFE50FFCFFE24" box="[818,963,477,500]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAFEFFCBAA3DFC9DFE50FFCFFE24" box="[818,963,477,500]" italics="true" pageId="3" pageNumber="475">Tetramorium</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAC4B1DFFCBAA3DFC7DFE51F86FFE24" author="Assing" box="[978,1123,476,501]" pageId="3" pageNumber="475" refString="Assing, V. (2002) On the Turkish and Caucasian species of Eurysunius, subgenus of Astenus Dejean, with an appendix on A. breuili Jarrige (Coleoptera: Staphylinidae, Paederinae). Linzer biologische Beitrage, 34 (1), 265 - 274." type="journal article" year="2002">Assing 2002</bibRefCitation>
,
<bibRefCitation id="EFAC4B1DFFCBAA3DFBC1FE51F8A4FE24" author="Assing" box="[1134,1192,476,500]" pageId="3" pageNumber="475" refString="Assing, V. (2007) New species and additional records of Staphylinidae from Turkey V (Coleoptera). Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 700, 1 - 64." type="journal article" year="2007">2007</bibRefCitation>
,
<bibRefCitation id="EFAC4B1DFFCBAA3DFB1BFE51F8E3FE24" author="Assing" box="[1204,1263,476,500]" pageId="3" pageNumber="475" refString="Assing, V. (2010) On the Staphylinidae of Turkey. VII. Five new species and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 80, 71 - 102." type="journal article" year="2010">2010</bibRefCitation>
,
<bibRefCitation id="EFAC4B1DFFCBAA3DFB55FE51F93AFE24" author="Assing" box="[1274,1334,476,500]" pageId="3" pageNumber="475" refString="Assing, V. (2011) On the Staphylinidae of Turkey VIII. Eleven new species, two new synonymies, a new combination, and additional records (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 81, 179 - 227." type="journal article" year="2011">2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>