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<mods:title id="DA4FB6805857B08CA58B55D2B173F1E3">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="17551C3CE782124D4130CC77EE32076A">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="2256EEE69ACCEBA0C9F6BE7027197577">Landau, Bernard</mods:namePart>
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<subSubSection id="C327654CFFE3FFC9FFBAFE396986FE6D" box="[151,500,402,429]" pageId="97" pageNumber="98" type="distribution">
<paragraph id="8B8236C7FFE3FFC9FFBAFE396986FE6D" blockId="97.[151,500,402,429]" box="[151,500,402,429]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFBAFE396986FE6D" bold="true" box="[151,500,402,429]" pageId="97" pageNumber="98">
Genus
<taxonomicName id="4C3D4D44FFE3FFC9FFC3FE38690DFE6D" ID-CoL="8ZH4Q" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[238,383,403,429]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FFE3FFC9FFC3FE38690DFE6D" bold="true" box="[238,383,403,429]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFE3FFC9FEAAFE386986FE6D" box="[391,500,403,429]" pageId="97" pageNumber="98" rank="genus">nov. gen.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C327654CFFE3FFC9FFBAFE7069EEFDD9" pageId="97" pageNumber="98" type="type_taxon">
<paragraph id="8B8236C7FFE3FFC9FFBAFE7069EEFDD9" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<materialsCitation id="3B553C9AFFE3FFC9FFBAFE7069EAFDD9" country="Hungary" county="Central Paratethys Sea" location="Middle Miocene" municipality="Langhian" pageId="97" pageNumber="98" specimenCount="1" typeStatus="holotype">
<emphasis id="B949EAD5FFE3FFC9FFBAFE706945FE34" bold="true" box="[151,311,474,501]" pageId="97" pageNumber="98">
<typeStatus id="54868865FFE3FFC9FFBAFE7068A0FE35" box="[151,210,475,501]" pageId="97" pageNumber="98">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3D4D44FFE3FFC9FE6FFE706B23FE35" ID-CoL="8KS29" authority="Kovacs &amp; Vician, 2021" authorityName="Kovacs &amp; Vician" authorityYear="2021" box="[322,849,475,501]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="harzhauseri">
<emphasis id="B949EAD5FFE3FFC9FE6FFE706A5DFE35" box="[322,559,475,501]" italics="true" pageId="97" pageNumber="98">Perrona harzhauseri</emphasis>
<bibRefCitation id="EFAC4B36FFE3FFC9FD16FE706B23FE35" author="Kovacs, Z. &amp; Vician, Z." box="[571,849,475,501]" pageId="97" pageNumber="98" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. &amp; Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács &amp; Vicián, 2021</bibRefCitation>
</taxonomicName>
;
<location id="8EE2601CFFE3FFC9FC4CFE706C50FE35" LSID="urn:lsid:plazi:treatment:039487D1FFE3FFC9FFBAFE3969EDF8B1:8EE2601CFFE3FFC9FC4CFE706C50FE35" box="[865,1058,475,501]" country="Hungary" county="Central Paratethys Sea" municipality="Langhian" name="Middle Miocene" pageId="97" pageNumber="98">Middle Miocene</location>
,
<collectingMunicipality id="6BE6ACBDFFE3FFC9FB1CFE706CE9FE35" box="[1073,1179,475,501]" pageId="97" pageNumber="98">Langhian</collectingMunicipality>
(Badenian),
<collectingCountry id="F32A7657FFE3FFC9FA18FE706DEAFE35" box="[1333,1432,475,501]" name="Hungary" pageId="97" pageNumber="98">Hungary</collectingCountry>
,
<collectingCounty id="62E34E4BFFE3FFC9FFBAFE5469EAFDD9" box="[151,408,511,537]" pageId="97" pageNumber="98">Central Paratethys Sea</collectingCounty>
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.
</paragraph>
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<subSubSection id="C327654CFFE3FFC9FFEAFD886B0AFDFD" box="[199,888,547,573]" pageId="97" pageNumber="98" type="etymology">
<paragraph id="8B8236C7FFE3FFC9FFEAFD886B0AFDFD" blockId="97.[151,1437,474,1905]" box="[199,888,547,573]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFD88693CFDFD" bold="true" box="[199,334,547,573]" pageId="97" pageNumber="98">Etymology.</emphasis>
Combination of
<emphasis id="B949EAD5FFE3FFC9FD21FD886A4BFDFC" box="[524,569,547,572]" italics="true" pageId="97" pageNumber="98">Neo</emphasis>
- (Greek: new) and
<taxonomicName id="4C3D4D44FFE3FFC9FC38FD886B03FDFC" box="[789,881,547,572]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FC38FD886B03FDFC" box="[789,881,547,572]" italics="true" pageId="97" pageNumber="98">Perrona</emphasis>
</taxonomicName>
.
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<subSubSection id="C327654CFFE3FFC9FFEAFDED6B85FCD5" pageId="97" pageNumber="98" type="diagnosis">
<paragraph id="8B8236C7FFE3FFC9FFEAFDED6B85FCD5" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFDED6930FDA0" bold="true" box="[199,322,582,608]" pageId="97" pageNumber="98">Diagnosis.</emphasis>
Large, solid, slender fusiform shell with high conical spire and low last whorl with short to moderately long siphonal canal. Early teleoconch whorls flat-sided with tripartite sculpture of weak, faintly beaded subsutural spiral cord and beaded suprasutural cord, separated by canaliculated mid-portion. Later teleoconch whorls nearly flat-sided with well-developed subsutural collar and suprasutural cord of equal strength, separated by canaliculate concavity. Last whorl with poorly developed to tubercular shoulder, with irregular spirals below, most strongly developed over mid-whorl and base, siphonal canal short to moderate length.
</paragraph>
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<subSubSection id="C327654CFFE3FFC9FFEAFCB56C08FA01" pageId="97" pageNumber="98" type="description">
<paragraph id="8B8236C7FFE3FFC9FFEAFCB5697CFB05" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFCB5692BFCF8" bold="true" box="[199,345,798,824]" pageId="97" pageNumber="98">Description.</emphasis>
Moderately large, slender fusiform with very high spire; apical angle ~3035°. Protoconch of about two smooth convex whorls. Teleoconch of up to11 whorls. Early teleoconch whorls flat-sided, with tripartite sculpture; weak, faintly beaded subsutural spiral cord, beaded suprasutural cord, separated by concavity filled by smooth, flattened central cord. Mid-spire subsutural cord becomes swollen to form broad, smooth, sharply delimited collar, central cord remains narrow, suprasutural beads weaken leaving broad, smooth to vaguely tubercular rounded cord above suture. Suture narrowly impressed, linear. Last whorl about 5055% of total height; with broad subsutural collar. Subsutural ramp not well-defined, with smooth, narrow, low, rounded cord. Shoulder not developed, evenly rounded at periphery, moderately strongly constricted at base. Siphonal fasciole weakly swollen, narrow. culpture of irregular spirals below shoulder. Aperture moderately narrow to moderately wide, ovoid. Outer lip thin, smooth within. Anal sinus moderately deep, narrow to moderately wide, asymmetrically U-shaped. Siphonal canal moderately short to long. Columellar and parietal callus thickened, sharply delimited, forming moderately broad callus rim.
</paragraph>
<paragraph id="8B8236C7FFE3FFC9FFEAFB656A22FA96" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFB656A0BFB29" bold="true" box="[199,633,1230,1257]" pageId="97" pageNumber="98">Stratigraphic and geographic range.</emphasis>
<taxonomicName id="4C3D4D44FFE3FFC9FDADFB646BE6FB29" baseAuthorityName="Kovacs &amp; Vician" baseAuthorityYear="2021" box="[640,916,1231,1257]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="harzhauseri">
<emphasis id="B949EAD5FFE3FFC9FDADFB646BE6FB29" box="[640,916,1231,1257]" italics="true" pageId="97" pageNumber="98">Neoperrona harzhauseri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D4D44FFE3FFC9FCE7FB646C19FB29" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[970,1131,1231,1257]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="zoltanorum">
<emphasis id="B949EAD5FFE3FFC9FCE7FB646C19FB29" box="[970,1131,1231,1257]" italics="true" pageId="97" pageNumber="98">N. zoltanorum</emphasis>
</taxonomicName>
both derive from Letkés in
<collectingCountry id="F32A7657FFE3FFC9FFBAFB586888FACD" box="[151,250,1267,1293]" name="Hungary" pageId="97" pageNumber="98">Hungary</collectingCountry>
, representing middle Miocene (Badenian, Langhian) occurrences in the Central Paratethys. The distribution in the Proto-Mediterranean Sea is documented by
<taxonomicName id="4C3D4D44FFE3FFC9FD92FABC6C0CFAF1" authority="(Bellardi, 1877)" baseAuthorityName="Bellardi" baseAuthorityYear="1877" box="[703,1150,1303,1329]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="taurinensis">
<emphasis id="B949EAD5FFE3FFC9FD92FABC6BB5FAF0" box="[703,967,1303,1328]" italics="true" pageId="97" pageNumber="98">Neoperrona taurinensis</emphasis>
(
<bibRefCitation id="EFAC4B36FFE3FFC9FCF9FABC6C04FAF1" author="Bellardi, L" box="[980,1142,1303,1329]" pageId="97" pageNumber="98" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
)
</taxonomicName>
from the middle Miocene (Langhian) of the Colli Torinesi (
<collectingCountry id="F32A7657FFE3FFC9FD26FA906A36FA96" box="[523,580,1339,1366]" name="Italy" pageId="97" pageNumber="98">Italy</collectingCountry>
).
</paragraph>
<paragraph id="8B8236C7FFE3FFC9FFEAFAF46BC7FA5D" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFAF469FDFAB8" bold="true" box="[199,399,1374,1401]" pageId="97" pageNumber="98">Included species.</emphasis>
Central Paratethys Sea:
<taxonomicName id="4C3D4D44FFE3FFC9FDB7FAF46CFBFAB9" authority="Kovacs &amp; Vician, 2021" authorityName="Kovacs &amp; Vician" authorityYear="2021" box="[666,1161,1375,1401]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="harzhauseri">
<emphasis id="B949EAD5FFE3FFC9FDB7FAF46BF3FAB9" box="[666,897,1375,1401]" italics="true" pageId="97" pageNumber="98">Perrona harzhauseri</emphasis>
<bibRefCitation id="EFAC4B36FFE3FFC9FCABFAF46CFBFAB9" author="Kovacs, Z. &amp; Vician, Z." box="[902,1161,1375,1401]" pageId="97" pageNumber="98" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. &amp; Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács &amp; Vicián, 2021</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C3D4D44FFE3FFC9FBBCFAF46DEFFAB9" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1169,1437,1375,1401]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="zoltanorum">
<emphasis id="B949EAD5FFE3FFC9FBBCFAF46DEFFAB9" box="[1169,1437,1375,1401]" italics="true" pageId="97" pageNumber="98">Neoperrona zoltanorum</emphasis>
</taxonomicName>
nov. sp.. Proto-Mediterraean Sea:
<taxonomicName id="4C3D4D44FFE3FFC9FD38FA286BC3FA5D" authority="Bellardi, 1877" authorityName="Bellardi" authorityYear="1877" box="[533,945,1411,1437]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="taurinensis">
<emphasis id="B949EAD5FFE3FFC9FD38FA286B74FA5C" box="[533,774,1411,1437]" italics="true" pageId="97" pageNumber="98">Clavatula taurinensis</emphasis>
<bibRefCitation id="EFAC4B36FFE3FFC9FC23FA286BC3FA5D" author="Bellardi, L" box="[782,945,1411,1437]" pageId="97" pageNumber="98" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8236C7FFE3FFC9FFEAFA0D6C08FA01" blockId="97.[151,1437,474,1905]" box="[199,1146,1446,1473]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFA0D69DAFA00" bold="true" box="[199,424,1446,1472]" pageId="97" pageNumber="98">Paleoenvironment.</emphasis>
Inner neritic environments with corals (
<bibRefCitation id="EFAC4B36FFE3FFC9FC44FA0C6C1DFA01" author="Kovacs, Z. &amp; Vician, Z." box="[873,1135,1447,1473]" pageId="97" pageNumber="98" pagination="53 - 100" refId="ref136963" refString="Kovacs, Z. &amp; Vician, Z. (2013) Badenian (middle Miocene) Conoidean (Neogastropoda) fauna from Letkes (N Hungary). Fragmenta Palaeontologica Hungarica, 30, 53 - 100." type="journal article" year="2013">Kovács &amp; Vicián 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFE3FFC9FFEAFA6169EDF8B1" pageId="97" pageNumber="98" type="discussion">
<paragraph id="8B8236C7FFE3FFC9FFEAFA616A6CF8C5" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<emphasis id="B949EAD5FFE3FFC9FFEAFA61693EFA24" bold="true" box="[199,332,1482,1508]" pageId="97" pageNumber="98">Discussion.</emphasis>
<taxonomicName id="4C3D4D44FFE3FFC9FE78FA6069A9FA24" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[341,475,1483,1508]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B949EAD5FFE3FFC9FE78FA6069A9FA24" box="[341,475,1483,1508]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFE3FFC9FEC8FA606A3FFA25" box="[485,589,1483,1509]" pageId="97" pageNumber="98" rank="genus">nov. gen.</taxonomicNameLabel>
is superficially reminiscent of a slender
<taxonomicName id="4C3D4D44FFE3FFC9FB0FFA606D2EFA25" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[1058,1372,1483,1509]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FB0FFA606C0CFA24" box="[1058,1150,1483,1508]" italics="true" pageId="97" pageNumber="98">Perrona</emphasis>
<bibRefCitation id="EFAC4B36FFE3FFC9FBAAFA606D2EFA25" author="Schumacher, H. C. F." box="[1159,1372,1483,1509]" pageId="97" pageNumber="98" refId="ref143804" refString="Schumacher, H. C. F. (1817) Essais d'un Nuveau Systeme des Habitations des Vers Testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up]" type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
, such as
<taxonomicName id="4C3D4D44FFE3FFC9FF9BFA446AD1F9C9" authority="(Hoernes &amp; Auinger, 1891)" baseAuthorityName="R. Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[182,675,1519,1545]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="oliviae">
<emphasis id="B949EAD5FFE3FFC9FF9BFA446917F9C9" box="[182,357,1519,1545]" italics="true" pageId="97" pageNumber="98">Perrona oliviae</emphasis>
(
<bibRefCitation id="EFAC4B36FFE3FFC9FE59FA446AE8F9C9" author="Hoernes, R. &amp; Auinger, M." box="[372,666,1519,1545]" pageId="97" pageNumber="98" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. &amp; Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes &amp; Auinger, 1891</bibRefCitation>
)
</taxonomicName>
. Consequently, its
<typeStatus id="54868865FFE3FFC9FC5BFA446BD4F9C9" box="[886,934,1519,1545]" pageId="97" pageNumber="98">type</typeStatus>
species was placed in
<taxonomicName id="4C3D4D44FFE3FFC9FB88FA446D73F9C8" authorityName="Schumacher" authorityYear="1817" box="[1189,1281,1519,1544]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FB88FA446D73F9C8" box="[1189,1281,1519,1544]" italics="true" pageId="97" pageNumber="98">Perrona</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAC4B36FFE3FFC9FA06FA446945F9ED" author="Kovacs, Z. &amp; Vician, Z." pageId="97" pageNumber="98" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. &amp; Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács &amp; Vicián (2021)</bibRefCitation>
. The tripartite sculpture of early teleoconch whorls, however, differs fundamentally from
<taxonomicName id="4C3D4D44FFE3FFC9FA14F9B86DE7F9EC" authorityName="Schumacher" authorityYear="1817" box="[1337,1429,1555,1580]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FA14F9B86DE7F9EC" box="[1337,1429,1555,1580]" italics="true" pageId="97" pageNumber="98">Perrona</emphasis>
</taxonomicName>
, which has comma-shaped riblets.
<taxonomicName id="4C3D4D44FFE3FFC9FD21F99C6A08F991" box="[524,634,1591,1617]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FD21F99C6A08F991" box="[524,634,1591,1617]" italics="true" pageId="97" pageNumber="98">Clavatula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D4D44FFE3FFC9FD80F99C6B69F991" box="[685,795,1591,1617]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FD80F99C6B69F991" box="[685,795,1591,1617]" italics="true" pageId="97" pageNumber="98">Clavatula</emphasis>
</taxonomicName>
-like groups discussed above do not develop a canaliculate mid-portion like
<taxonomicName id="4C3D4D44FFE3FFC9FE71F9F06990F9B4" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[348,482,1627,1652]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FE71F9F06990F9B4" box="[348,482,1627,1652]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
. Moreover,
<taxonomicName id="4C3D4D44FFE3FFC9FD5EF9F06A93F9B5" box="[627,737,1627,1653]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FD5EF9F06A93F9B5" box="[627,737,1627,1653]" italics="true" pageId="97" pageNumber="98">Clavatula</emphasis>
</taxonomicName>
develops a swollen and more or less tuberculate subsutural collar. Finally, the slender fusiform shell and the high conical spire and very low last whorl separate
<taxonomicName id="4C3D4D44FFE3FFC9FA3BF9D46DEEF958" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1302,1436,1663,1688]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FA3BF9D46DEEF958" box="[1302,1436,1663,1688]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
from
<taxonomicName id="4C3D4D44FFE3FFC9FFF9F9086930F97D" box="[212,322,1699,1725]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FFF9F9086930F97D" box="[212,322,1699,1725]" italics="true" pageId="97" pageNumber="98">Clavatula</emphasis>
</taxonomicName>
, which is broader, and in which the last whorl accounts for more than 60% of the total height. Thus,
<taxonomicName id="4C3D4D44FFE3FFC9FFBAF96C696FF920" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[151,285,1735,1760]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FFBAF96C696FF920" box="[151,285,1735,1760]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
differs from other clavatulid genera and species-groups recognized herein by having a proportionately higher spire and shorter last whorl.
</paragraph>
<paragraph id="8B8236C7FFE3FFC9FFEAF8A469EDF8B1" blockId="97.[151,1437,474,1905]" pageId="97" pageNumber="98">
<taxonomicName id="4C3D4D44FFE3FFC9FFEAF8A4693FF8E8" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[199,333,1807,1832]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FFEAF8A4693FF8E8" box="[199,333,1807,1832]" italics="true" pageId="97" pageNumber="98">Neoperrona</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="4C3D4D44FFE3FFC9FD88F8A46B61F8E9" box="[677,787,1807,1833]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFE3FFC9FD88F8A46B61F8E9" box="[677,787,1807,1833]" italics="true" pageId="97" pageNumber="98">Clavatula</emphasis>
</taxonomicName>
<emphasis id="B949EAD5FFE3FFC9FC08F8A46B10F8E8" box="[805,866,1807,1832]" italics="true" pageId="97" pageNumber="98">taxea</emphasis>
and
<emphasis id="B949EAD5FFE3FFC9FCB2F8A46CFCF8E8" box="[927,1166,1807,1833]" italics="true" pageId="97" pageNumber="98">Clavatula tripartita</emphasis>
concerning the slender outline. The
<emphasis id="B949EAD5FFE3FFC9FE09F8986918F88C" box="[292,362,1843,1868]" italics="true" pageId="97" pageNumber="98">taxea-</emphasis>
clade, however, is characterized by early teleoconch whorls with numerous spiral cords and lack beaded adsutural cords.
</paragraph>
</subSubSection>
</treatment>
</document>