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<document id="07B0CB35F87F155BBC2C4118A1ADF923" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard &amp; Janssen, Ronald" docDate="2022" docId="039487D1FF0BFF22FFBAFA9168AEFC65" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Pusionella hofmanni Harzhauser &amp; Landau &amp; Janssen 2022, nov. sp." docType="treatment" docVersion="1" lastPageNumber="139" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="138" updateTime="1709124758513" updateUser="karina">
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<mods:title id="0F4497DB6CAFD1B2E4633EFD590BAE67">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="E86AAF4362F2B9E0687D6A2D064C382B">Landau, Bernard</mods:namePart>
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<taxonomicName id="4C3D4D44FF0BFF21FFBAFA9169E4FA95" ID-CoL="8ZGQD" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[151,406,1338,1365]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="137" pageNumber="138" phylum="Mollusca" rank="species" species="hofmanni" status="sp. nov.">
<emphasis id="B949EAD5FF0BFF21FFBAFA9169E4FA95" bold="true" box="[151,406,1338,1365]" italics="true" pageId="137" pageNumber="138">Pusionella hofmanni</emphasis>
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<paragraph id="8B8236C7FF0BFF21FFBAFAF46A41FAB9" blockId="137.[151,563,1338,1407]" box="[151,563,1374,1407]" pageId="137" pageNumber="138">
<figureCitation id="13062A42FF0BFF21FFBAFAF4688EFAB9" box="[151,252,1375,1401]" captionStart="FIGURE 41" captionStartId="136.[151,250,1832,1856]" captionTargetBox="[151,1436,181,1809]" captionTargetId="figure-26@136.[151,1436,181,1809]" captionTargetPageId="136" captionText="FIGURE 41 A1A3. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0001, Várpalota (Hungary). B1B3. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). C1C3. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0002, Várpalota (Hungary). D. Pusionella palatina Strausz, 1954, lectotype, M.60.4948, Várpalota (Hungary). E1 E2. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). F1F2. Pusionella hofmanni nov. sp., NHMW 1867/0019/0142a, paratype, Coşteiu de Sus (Romania). G1G2. Pusionella hofmanni nov. sp., NHMW 1867/0019/0142b, paratype, Coşteiu de Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.6399923" httpUri="https://zenodo.org/record/6399923/files/figure.png" pageId="137" pageNumber="138">Figs 41E</figureCitation>
<figureCitation id="13062A42FF0BFF21FFD1FADA6951FAB9" box="[252,291,1375,1407]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="137" pageNumber="138">
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E
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<figureCitation id="13062A42FF0BFF21FE0EFADA6959FABF" box="[291,299,1393,1407]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="137" pageNumber="138">
<subScript id="17B93482FF0BFF21FE0EFADA6959FABF" attach="left" box="[291,299,1393,1407]" fontSize="6" pageId="137" pageNumber="138">2</subScript>
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, F
<figureCitation id="13062A42FF0BFF21FE64FADA691CFAB9" box="[329,366,1375,1407]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="137" pageNumber="138">
<subScript id="17B93482FF0BFF21FE64FADA6923FABF" attach="left" box="[329,337,1393,1407]" fontSize="6" pageId="137" pageNumber="138">1</subScript>
F
</figureCitation>
<figureCitation id="13062A42FF0BFF21FE43FADA6904FABF" box="[366,374,1393,1407]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="137" pageNumber="138">
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, G
<figureCitation id="13062A42FF0BFF21FEB4FADA69B1FAB9" box="[409,451,1375,1407]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="137" pageNumber="138">
<subScript id="17B93482FF0BFF21FEB4FADA69D3FABF" attach="right" box="[409,417,1393,1407]" fontSize="6" pageId="137" pageNumber="138">1</subScript>
G
</figureCitation>
<figureCitation id="13062A42FF0BFF21FEEEFADA69B9FABF" box="[451,459,1393,1407]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="137" pageNumber="138">
<subScript id="17B93482FF0BFF21FEEEFADA69B9FABF" attach="left" box="[451,459,1393,1407]" fontSize="6" pageId="137" pageNumber="138">2</subScript>
</figureCitation>
,
<figureCitation id="13062A42FF0BFF21FEF4FAF4698CFAB9" box="[473,510,1375,1401]" captionStart="FIGURE 4" captionStartId="14.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@14.[151,1436,181,1595]" captionTargetPageId="14" captionText="FIGURE 4A. Clavatula justinae (Hoernes &amp; Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. Clavatula mariae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. Clavatula olgae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. Clavatula orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. Clavatula juliae (Hoernes &amp; Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. Clavatula apolloniae (Hoernes &amp; Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács &amp; Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes &amp; Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes &amp; Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes &amp; Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes &amp; Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes &amp; Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes &amp; Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes &amp; Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács &amp; Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes &amp; Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae." figureDoi="http://doi.org/10.5281/zenodo.6399837" httpUri="https://zenodo.org/record/6399837/files/figure.png" pageId="137" pageNumber="138">4K</figureCitation>
,
<figureCitation id="13062A42FF0BFF21FD24FAF56A6BFAB9" box="[521,537,1374,1401]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="137" pageNumber="138">5</figureCitation>
,
<figureCitation id="13062A42FF0BFF21FD08FAF46A41FAB9" box="[549,563,1375,1401]" captionStart="FIGURE 7" captionStartId="17.[151,250,1938,1962]" captionTargetBox="[151,1436,345,1915]" captionTargetId="figure-109@17.[151,1436,345,1915]" captionTargetPageId="17" captionText="FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, C. = Clavatula, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana)." figureDoi="http://doi.org/10.5281/zenodo.6399845" httpUri="https://zenodo.org/record/6399845/files/figure.png" pageId="137" pageNumber="138">7</figureCitation>
</paragraph>
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<paragraph id="8B8236C7FF0BFF21FFBAFA0C6B3AF9EE" blockId="137.[151,1437,1446,2013]" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFBAFA0C6937FA00" bold="true" box="[151,325,1446,1473]" pageId="137" pageNumber="138">Type material.</emphasis>
<materialsCitation id="3B553C9AFF0BFF21FE60FA0C694BFA25" collectionCode="MD" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="137" pageNumber="138" specimenCode="NHMW 1867" specimenCount="1" typeStatus="holotype">
<typeStatus id="54868865FF0BFF21FE60FA0C69C8FA01" box="[333,442,1447,1473]" pageId="137" pageNumber="138" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB9B9EBCFF0BFF21FEE9FA0C6A2FFA01" box="[452,605,1447,1473]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="137" pageNumber="138" type="Museum">NHMW 1867</specimenCode>
/0019/0136, SL: 44.0 mm,
<collectionCode id="ED2CAE02FF0BFF21FCA2FA0C6BB2FA01" box="[911,960,1447,1473]" country="Germany" name="Museum Donaueschingen" pageId="137" pageNumber="138">MD</collectionCode>
:
<quantity id="4CC59B22FF0BFF21FCE6FA0C6C40FA01" box="[971,1074,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="137" pageNumber="138" unit="mm" value="14.1">14.1 mm</quantity>
,
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Coşteiu
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(
<collectingCountry id="F32A7657FF0BFF21FBD9FA0C6D2DFA02" box="[1268,1375,1447,1474]" name="Romania" pageId="137" pageNumber="138">Romania</collectingCountry>
), figs 401E
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E
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, 4K
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.
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<typeStatus id="54868865FF0BFF21FE6AFA6069CAFA25" box="[327,440,1483,1509]" pageId="137" pageNumber="138" type="paratype">Paratypes</typeStatus>
:
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/0019/0142a, SL:
<quantity id="4CC59B22FF0BFF21FC18FA606BECFA25" box="[821,926,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="137" pageNumber="138" unit="mm" value="33.8">33.8 mm</quantity>
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,
<materialsCitation id="3B553C9AFF0BFF21FC80FA606DEAFA26" box="[941,1432,1483,1510]" collectionCode="MD" country="Romania" location="Costeiu de Sus" pageId="137" pageNumber="138" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED2CAE02FF0BFF21FC80FA606BACFA25" box="[941,990,1483,1509]" country="Germany" name="Museum Donaueschingen" pageId="137" pageNumber="138">MD</collectionCode>
:
<quantity id="4CC59B22FF0BFF21FCC0FA606C24FA25" box="[1005,1110,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="137" pageNumber="138" unit="mm" value="12.1">12.1 mm</quantity>
,
<location id="8EE2601CFF0BFF21FB48FA606D60FA25" LSID="urn:lsid:plazi:treatment:039487D1FF0BFF22FFBAFA9168AEFC65:8EE2601CFF0BFF21FB48FA606D60FA25" box="[1125,1298,1483,1509]" country="Romania" name="Costeiu de Sus" pageId="137" pageNumber="138">Coşteiu de Sus</location>
(
<collectingCountry id="F32A7657FF0BFF21FA08FA606DE2FA26" box="[1317,1424,1483,1510]" name="Romania" pageId="137" pageNumber="138">Romania</collectingCountry>
)
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, figs 41F
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F
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;
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<specimenCode id="DB9B9EBCFF0BFF21FE19FA4469BCF9C9" box="[308,462,1519,1545]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="137" pageNumber="138" type="Museum">NHMW 1867</specimenCode>
/0019/0142b, SL:
<quantity id="4CC59B22FF0BFF21FDB3FA446B74F9C9" box="[670,774,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="137" pageNumber="138" unit="mm" value="30.1">30.1 mm</quantity>
</materialsCitation>
,
<materialsCitation id="3B553C9AFF0BFF21FC3EFA446C80F9CA" box="[787,1266,1519,1546]" collectionCode="MD" country="Romania" location="Costeiu de Sus" pageId="137" pageNumber="138" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED2CAE02FF0BFF21FC3EFA446B36F9C9" box="[787,836,1519,1545]" country="Germany" name="Museum Donaueschingen" pageId="137" pageNumber="138">MD</collectionCode>
: 11.0 mm,
<location id="8EE2601CFF0BFF21FCE8FA446C1CF9C9" LSID="urn:lsid:plazi:treatment:039487D1FF0BFF22FFBAFA9168AEFC65:8EE2601CFF0BFF21FCE8FA446C1CF9C9" box="[965,1134,1519,1545]" country="Romania" name="Costeiu de Sus" pageId="137" pageNumber="138">Coşteiu de Sus</location>
(
<collectingCountry id="F32A7657FF0BFF21FB52FA446C98F9CA" box="[1151,1258,1519,1546]" name="Romania" pageId="137" pageNumber="138">Romania</collectingCountry>
)
</materialsCitation>
, figs 41G
<subScript id="17B93482FF0BFF21FA4FF9AA6D18F9CF" attach="left" box="[1378,1386,1537,1551]" fontSize="6" pageId="137" pageNumber="138">1</subScript>
G
<subScript id="17B93482FF0BFF21FAA1F9AA6DE6F9CF" attach="left" box="[1420,1428,1537,1551]" fontSize="6" pageId="137" pageNumber="138">2</subScript>
;
<materialsCitation id="3B553C9AFF0BFF21FFBAF9B86B36F9EE" box="[151,836,1555,1582]" collectionCode="NHMW" country="Romania" location="Costeiu de Sus" pageId="137" pageNumber="138" specimenCode="NHMW 1867" specimenCount="4" typeStatus="paratype">
<specimenCode id="DB9B9EBCFF0BFF21FFBAF9B8695DF9ED" box="[151,303,1555,1581]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="137" pageNumber="138" type="Museum">NHMW 1867</specimenCode>
/0019/0142,
<specimenCount id="9D3BFD4EFF0BFF21FE92F9B86A62F9ED" box="[447,528,1555,1581]" pageId="137" pageNumber="138" type="generic">4 spec.</specimenCount>
,
<location id="8EE2601CFF0BFF21FD36F9B86AB3F9ED" LSID="urn:lsid:plazi:treatment:039487D1FF0BFF22FFBAFA9168AEFC65:8EE2601CFF0BFF21FD36F9B86AB3F9ED" box="[539,705,1555,1581]" country="Romania" name="Costeiu de Sus" pageId="137" pageNumber="138">Coşteiu de Sus</location>
(
<collectingCountry id="F32A7657FF0BFF21FDFDF9B86B4EF9EE" box="[720,828,1555,1582]" name="Romania" pageId="137" pageNumber="138">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236C7FF0BFF21FFEAF99C6B5EF991" blockId="137.[151,1437,1446,2013]" box="[199,812,1590,1618]" pageId="137" pageNumber="138">
<materialsCitation id="3B553C9AFF0BFF21FFEAF99C6B5AF991" box="[199,808,1590,1618]" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="137" pageNumber="138" specimenCount="1" typeStatus="holotype">
<emphasis id="B949EAD5FF0BFF21FFEAF99C6914F990" bold="true" box="[199,358,1590,1617]" pageId="137" pageNumber="138">
<typeStatus id="54868865FF0BFF21FFEAF99C6970F991" box="[199,258,1591,1617]" pageId="137" pageNumber="138">Type</typeStatus>
locality.
</emphasis>
<location id="8EE2601CFF0BFF21FE41F99C6A60F991" LSID="urn:lsid:plazi:treatment:039487D1FF0BFF22FFBAFA9168AEFC65:8EE2601CFF0BFF21FE41F99C6A60F991" box="[364,530,1591,1617]" country="Romania" name="Costeiu de Sus" pageId="137" pageNumber="138">
Coşteiu
<collectorName id="26C85311FF0BFF21FEE4F99C6A60F991" box="[457,530,1591,1617]" pageId="137" pageNumber="138">de Sus</collectorName>
</location>
(
<collectingCountry id="F32A7657FF0BFF21FD0CF99C6AFEF992" box="[545,652,1591,1618]" name="Romania" pageId="137" pageNumber="138">Romania</collectingCountry>
), Făget Basin
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236C7FF0BFF21FFEAF9F06A8EF9B5" blockId="137.[151,1437,1446,2013]" box="[199,764,1627,1653]" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFEAF9F0691DF9B5" bold="true" box="[199,367,1627,1653]" pageId="137" pageNumber="138">
<typeStatus id="54868865FF0BFF21FFEAF9F06970F9B5" box="[199,258,1627,1653]" pageId="137" pageNumber="138">Type</typeStatus>
stratum.
</emphasis>
Silt and clay of the Dej Formation.
</paragraph>
<paragraph id="8B8236C7FF0BFF21FFEAF9D56B27F95A" blockId="137.[151,1437,1446,2013]" box="[199,853,1662,1690]" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFEAF9D5688FF958" bold="true" box="[199,253,1662,1688]" pageId="137" pageNumber="138">Age.</emphasis>
Middle Miocene, early/middle Badenian (Langhian).
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF0BFF21FFEAF9086A20F921" pageId="137" pageNumber="138" type="etymology">
<paragraph id="8B8236C7FF0BFF21FFEAF9086A20F921" blockId="137.[151,1437,1446,2013]" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFEAF908693CF97D" bold="true" box="[199,334,1699,1725]" pageId="137" pageNumber="138">Etymology.</emphasis>
In honor of Thomas Hofmann (Geological Survey of
<collectingCountry id="F32A7657FF0BFF21FC88F9086B88F97D" box="[933,1018,1699,1725]" name="Austria" pageId="137" pageNumber="138">Austria</collectingCountry>
,
<collectingRegion id="49F9F825FF0BFF21FB29F9086C2AF97E" box="[1028,1112,1699,1726]" country="Austria" name="Wien" pageId="137" pageNumber="138">Vienna</collectingRegion>
), geologist, great popularizer of science and friend of the first author.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF0BFF21FFEAF9416964F88D" pageId="137" pageNumber="138" type="diagnosis">
<paragraph id="8B8236C7FF0BFF21FFEAF9416964F88D" blockId="137.[151,1437,1446,2013]" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFEAF9416930F8C4" bold="true" box="[199,322,1770,1796]" pageId="137" pageNumber="138">Diagnosis.</emphasis>
Medium-sized, fusiform with high, weakly gradate spire, subcylindrical whorls with rounded shoulder and convex base. Early teleoconch whorls with deep spiral groove; siphonal canal with strongly raised spiral cord.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF0BFF22FFEAF8FD6C4AFE6D" lastPageId="138" lastPageNumber="139" pageId="137" pageNumber="138" type="description">
<paragraph id="8B8236C7FF0BFF22FFEAF8FD6C4AFE6D" blockId="137.[151,1437,1446,2013]" lastBlockId="138.[151,1437,151,933]" lastPageId="138" lastPageNumber="139" pageId="137" pageNumber="138">
<emphasis id="B949EAD5FF0BFF21FFEAF8FD692BF8B0" bold="true" box="[199,345,1878,1904]" pageId="137" pageNumber="138">Description.</emphasis>
Shell medium-sized, moderately broad fusiform with high, weakly gradate spire; apical angle ~38°. Protoconch not preserved. Teleoconch of 12 whorls. Early teleoconch whorls weakly concave, with weak, smooth subsutural spiral cord and suprasutural row of large beads, separated by deeply incised spiral groove. On third to fourth whorl suprasutural beads fade, deep groove separates narrower slightly concave subsutural cord from wider abapical portion of whorl. Spiral groove shifting slightly in abapical direction on subsequent whorls, becoming subobsolete on eighth whorl. Late whorls smooth, subcylindrical. Suture weakly impressed, linear. Last whorl 56 60% of total height, subcylindrical. Subsutural collar, ramp and shoulder not developed. Rounded and moderately constricted at base. Siphonal fasciole strongly delimited by raised ridge, flattened, twisted, with prominent growth lines. Sculpture of weak, narrow cords restricted to base and fasciole.Aperture moderately wide, pyriform. Outer lip thin, smooth within. Anal sinus very shallow, wide; siphonal canal moderately short, strongly deflected to left and dorsally, shallowly notched at tip. Columella weakly twisted, smooth. Columella excavated in upper third, strongly twisted at fasciole. Columellar and parietal callus thin, poorly delimited from base.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF08FF22FFEAFE1D6A57FC9D" pageId="138" pageNumber="139" type="discussion">
<paragraph id="8B8236C7FF08FF22FFEAFE1D6927FDFD" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
<emphasis id="B949EAD5FF08FF22FFEAFE1D693EFE10" bold="true" box="[199,332,438,464]" pageId="138" pageNumber="139">Discussion.</emphasis>
<taxonomicName id="4C3D4D44FF08FF22FE75FE1C6A37FE11" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[344,581,439,465]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="hofmanni" status="sp. nov.">
<emphasis id="B949EAD5FF08FF22FE75FE1C6A37FE11" box="[344,581,439,465]" italics="true" pageId="138" pageNumber="139">Pusionella hofmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFF08FF22FD7DFE1C6AD9FE11" box="[592,683,439,465]" pageId="138" pageNumber="139" rank="species">nov. sp.</taxonomicNameLabel>
is morphologically close to
<taxonomicName id="4C3D4D44FF08FF22FCD2FE1C6D5AFE11" authority="Strausz, 1954" authorityName="Strausz" authorityYear="1954" box="[1023,1320,438,465]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="palatina">
<emphasis id="B949EAD5FF08FF22FCD2FE1C6C0EFE11" box="[1023,1148,439,465]" italics="true" pageId="138" pageNumber="139">P. palatina</emphasis>
<bibRefCitation id="EFAC4B36FF08FF22FBA5FE1C6D5AFE11" author="Strausz, L." box="[1160,1320,438,465]" pageId="138" pageNumber="139" pagination="1 - 150" refId="ref144830" refString="Strausz, L. (1954) Varpalotai felso-mediteean csigak. Geologica Hungarica, 25, 1 - 150. [http: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954 _ 085 - 125. pdf]" type="journal article" year="1954">Strausz, 1954</bibRefCitation>
</taxonomicName>
, but in
<taxonomicName id="4C3D4D44FF08FF22FAA5FE1C6886FE35" authorityName="Strausz" authorityYear="1954" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="palatina">
<emphasis id="B949EAD5FF08FF22FAA5FE1C6886FE35" italics="true" pageId="138" pageNumber="139">P. palatina</emphasis>
</taxonomicName>
the early whorls are coeloconoid, with the beads at the abapical suture persisting to the sixth whorl. In
<taxonomicName id="4C3D4D44FF08FF22FAA5FE706973FDD9" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="hofmanni">
<emphasis id="B949EAD5FF08FF22FAA5FE706973FDD9" italics="true" pageId="138" pageNumber="139">P. hofmanni</emphasis>
</taxonomicName>
the early whorl profile is regularly conical and broader, the whorls more depressed and the beads fade by the fourth whorl.
</paragraph>
<paragraph id="8B8236C7FF08FF22FFEAFDEC6ADBFCF9" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
In addition,
<taxonomicName id="4C3D4D44FF08FF22FE60FDEC6A46FDA1" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[333,564,583,609]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="hofmanni">
<emphasis id="B949EAD5FF08FF22FE60FDEC6A46FDA1" box="[333,564,583,609]" italics="true" pageId="138" pageNumber="139">Pusionella hofmanni</emphasis>
</taxonomicName>
differs in the generally less conical, but slightly gradate spire, the deeper spiral groove on early spire whorls, and the shallowly notched and less deflected siphonal canal. The extant
<taxonomicName id="4C3D4D44FF08FF22FA09FDC06A73FD69" authority="Boyer &amp; Ryall, 2006" authorityName="Boyer &amp; Ryall" authorityYear="2006" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="ghanaensis">
<emphasis id="B949EAD5FF08FF22FA09FDC06967FD69" italics="true" pageId="138" pageNumber="139">Pusionella ghanaensis</emphasis>
<bibRefCitation id="EFAC4B36FF08FF22FE36FD246A73FD69" author="Boyer, F. &amp; Ryall, P." box="[283,513,655,681]" pageId="138" pageNumber="139" pagination="33 - 38" refId="ref130057" refString="Boyer, F. &amp; Ryall, P. (2006) Two new Clavatulinae species (Caenogastropda: Turridae) from Ghana. Iberus, 24, 33 - 38. https: // www. biodiversitylibrary. org / item / 133983 # page / 205 / mode / 1 up" type="journal article" year="2006">Boyer &amp; Ryall, 2006</bibRefCitation>
</taxonomicName>
develops a similar spiral groove on early spire whorls, but differs from
<taxonomicName id="4C3D4D44FF08FF22FA35FD246DEEFD69" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1304,1436,655,681]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="hofmanni">
<emphasis id="B949EAD5FF08FF22FA35FD246DEEFD69" box="[1304,1436,655,681]" italics="true" pageId="138" pageNumber="139">P. hofmanni</emphasis>
</taxonomicName>
in having spiral cords on the last whorl.
<taxonomicName id="4C3D4D44FF08FF22FD59FD186C08FD0D" authority="(Des Moulins, 1842)" baseAuthorityName="Des Moulins" baseAuthorityYear="1842" box="[628,1146,691,717]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="pseudofusus">
<emphasis id="B949EAD5FF08FF22FD59FD186B0DFD0D" box="[628,895,691,717]" italics="true" pageId="138" pageNumber="139">Pusionella pseudofusus</emphasis>
(
<bibRefCitation id="EFAC4B36FF08FF22FCBCFD186C01FD0D" author="Des Moulins, M. Ch." box="[913,1139,691,717]" pageId="138" pageNumber="139" pagination="109 - 185" refId="ref131318" refString="Des Moulins, M. Ch. (1842) Revision de quelques especes de Pleurotomes. Actes de la Societe linneenne de Bordeaux, 12, 109 - 185. [https: // www. biodiversitylibrary. org / item / 101660 # page / 164 / mode / 1 up]" type="journal article" year="1842">Des Moulins, 1842</bibRefCitation>
)
</taxonomicName>
, from the Burdigalian of
<collectingCountry id="F32A7657FF08FF22FFBAFD7C6894FD31" box="[151,230,727,753]" name="France" pageId="138" pageNumber="139">France</collectingCountry>
, and
<taxonomicName id="4C3D4D44FF08FF22FE08FD7C6A34FD31" authority="Sacco, 1891" authorityName="Sacco" authorityYear="1891" box="[293,582,727,753]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="tauronifat">
<emphasis id="B949EAD5FF08FF22FE08FD7C69C0FD31" box="[293,434,727,753]" italics="true" pageId="138" pageNumber="139">P. tauronifat</emphasis>
<bibRefCitation id="EFAC4B36FF08FF22FE96FD7C6A34FD31" author="Sacco, F." box="[443,582,727,753]" pageId="138" pageNumber="139" refId="ref143073" refString="Sacco, F. (1891) I Molluschi dei Terreni Terziarii del Piemonte e della Liguria. Parte 10: Cassididae (Aggiunte), Terebridae e Pusionellidae. Clausen, Torino, 68 pp. [https: // www. biodiversitylibrary. org / item / 47831 # page / 7 / mode / 1 up]" type="book" year="1891">Sacco, 1891</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3D4D44FF08FF22FDADFD7C6BB9FD31" authority="Sacco, 1891" authorityName="Sacco" authorityYear="1891" box="[640,971,727,753]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="138" pageNumber="139" phylum="Mollusca" rank="species" species="pedemontana">
<emphasis id="B949EAD5FF08FF22FDADFD7C6B47FD31" box="[640,821,727,753]" italics="true" pageId="138" pageNumber="139">P. pedemontana</emphasis>
<bibRefCitation id="EFAC4B36FF08FF22FC13FD7C6BB9FD31" author="Sacco, F." box="[830,971,727,753]" pageId="138" pageNumber="139" refId="ref143073" refString="Sacco, F. (1891) I Molluschi dei Terreni Terziarii del Piemonte e della Liguria. Parte 10: Cassididae (Aggiunte), Terebridae e Pusionellidae. Clausen, Torino, 68 pp. [https: // www. biodiversitylibrary. org / item / 47831 # page / 7 / mode / 1 up]" type="book" year="1891">Sacco, 1891</bibRefCitation>
</taxonomicName>
, from the Burdigalian of
<collectingCountry id="F32A7657FF08FF22FBDCFD7C6D57FD31" box="[1265,1317,727,753]" name="Italy" pageId="138" pageNumber="139">Italy</collectingCountry>
, all lack a spiral groove on early teleoconch whorls (see
<bibRefCitation id="EFAC4B36FF08FF22FD8BFD506B1FFCD5" author="Peyrot, A." box="[678,877,763,789]" pageId="138" pageNumber="139" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 &amp; 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot 1931: 103</bibRefCitation>
, pl. 9, figs 7476;
<bibRefCitation id="EFAC4B36FF08FF22FB6AFD506C8BFCD5" author="Sacco, F." box="[1095,1273,763,789]" pageId="138" pageNumber="139" refId="ref143133" refString="Sacco, F. (1904) I molluschi dei terreni terziari del Piemonte e della Liguria. Parte 30. Aggiunte e correzioni (con 1400 figure). Considerazioni generali. Indice generale dell'opera. C. Clausen, Torino, 203 + xxxvi pp., 31 pls. [https: // www. biodiversitylibrary. org / item / 46270 # page / 90 / mode / 1 up]" type="book" year="1904">Sacco 1904: 62</bibRefCitation>
, pl. 2, fig. 77;
<bibRefCitation id="EFAC4B36FF08FF22FFBAFCB46984FCF9" author="Ferrero Mortara, E. &amp; Montefameglio &amp; Novelli, M. &amp; Opesso, G. &amp; Pavia, G. &amp; Tampieri, R." box="[151,502,799,825]" pageId="138" pageNumber="139" pagination="1 - 489" refId="ref132014" refString="Ferrero Mortara, E., Montefameglio, Novelli, M., Opesso, G., Pavia, G. &amp; Tampieri, R. 1984 Museo Regionale di Scienze Naturali, Catalogo, 7, 1 - 489. [https: // www. biodiversitylibrary. org / item / 269689 # page / 5 / mode / 1 up]" type="journal article" year="1984">
Ferrero Mortara
<emphasis id="B949EAD5FF08FF22FE7FFCB469F8FCF9" box="[338,394,799,825]" italics="true" pageId="138" pageNumber="139">et al.</emphasis>
1984: 66
</bibRefCitation>
, pl. 56, fig. 14).
</paragraph>
<paragraph id="8B8236C7FF08FF22FFEAFCE96A57FC9D" blockId="138.[151,1437,151,933]" box="[199,549,834,861]" pageId="138" pageNumber="139">
<emphasis id="B949EAD5FF08FF22FFEAFCE969DAFC9C" bold="true" box="[199,424,834,860]" pageId="138" pageNumber="139">Paleoenvironment.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFF08FF22FFEAFCCD68AEFC65" pageId="138" pageNumber="139" type="distribution">
<paragraph id="8B8236C7FF08FF22FFEAFCCD68AEFC65" blockId="138.[151,1437,151,933]" pageId="138" pageNumber="139">
<emphasis id="B949EAD5FF08FF22FFEAFCCD6A14FC41" bold="true" box="[199,614,870,897]" pageId="138" pageNumber="139">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="B949EAD5FF08FF22FC9DFCCC6C45FC40" box="[944,1079,871,897]" italics="true" pageId="138" pageNumber="139">Făget Basin</emphasis>
: Coşteiu de Sus (
<collectingCountry id="F32A7657FF08FF22FBD8FCCC6D2CFC42" box="[1269,1374,871,898]" name="Romania" pageId="138" pageNumber="139">Romania</collectingCountry>
) (
<emphasis id="B949EAD5FF08FF22FA59FCCC68BEFC64" italics="true" pageId="138" pageNumber="139">hoc opus</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>