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<document id="2FD821F6D8826832BE71177DA4D1320A" ID-CLB-Dataset="5024" ID-DOI="10.5852/ejt.2021.761.1449" ID-GBIF-Dataset="bf24ff1f-569a-4222-afe1-8527c83d8003" ID-ISSN="2118-9773" ID-Zenodo-Dep="5156259" ID-ZooBank="519E2F56-40E1-4431-BC90-8484E308D16E" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628011133532" checkinUser="felipe" docAuthor="Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C. &amp; Robillard, Tony" docDate="2021" docId="039487A4FFF2FFDCFDC7F167FEECF96A" docLanguage="en" docName="ejt-761_1-75.pdf" docOrigin="European Journal of Taxonomy 761 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.761.1449" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nisitrus danum Robillard &amp; Tan 2021, sp. nov." docType="treatment" docUuid="0DEB5BDB-6E85-4691-B5D1-E6258239370A" docUuidSource="ZooBank" docVersion="16" lastPageNumber="33" masterDocId="FFADFFDCFFECFFFCFFE0F365FFD5FFF9" masterDocTitle="Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species" masterLastPageNumber="75" masterPageNumber="1" pageNumber="31" updateTime="1699041084605" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9CBA6AD712BC5506A6FFCA65627C1ED0">Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species</mods:title>
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<mods:namePart id="5490E86E52AA9A49189F9AA62657E959">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="97A8D03E6D5262397BB48560769495E5">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national dʼhistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:namePart id="58F963524F14EA3D9C77492209735A76">Wahab, Rodzay bin Haji Abdul</mods:namePart>
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<mods:affiliation id="449799E3697F56FBDBD9565CBB392964">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam.</mods:affiliation>
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<mods:namePart id="10007BFCE3C6488008225E0509E43D1D">Japir, Razy</mods:namePart>
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<mods:affiliation id="2B907F3B88BE00F08FF1DFC1138CCFAD">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
<mods:nameIdentifier id="EED745327B40D9FA01D20C8F047E15D3" type="email">razy.Japir@sabah.gov.my</mods:nameIdentifier>
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<mods:namePart id="315431AD1DC317F0563BE2FDF9A25B1B">Chung, Authur Y. C.</mods:namePart>
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<mods:affiliation id="2E55970B566F83FEFBD4DE9160F4D85D">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
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<mods:namePart id="6EFDA10C6534786FC7B00660FDEFA16C">Robillard, Tony</mods:namePart>
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<mods:affiliation id="FED3BF9678A1D8DA812B381301B1B83A">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national dʼhistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:title id="B681193B4D85766725808B3C02021B20">European Journal of Taxonomy</mods:title>
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<mods:date id="F67B7C9BD9389E07E1CF5AF947AAE5EF">2021</mods:date>
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<treatment id="039487A4FFF2FFDCFDC7F167FEECF96A" ID-DOI="http://doi.org/10.5281/zenodo.5170001" ID-GBIF-Taxon="184010537" ID-Zenodo-Dep="5170001" LSID="urn:lsid:zoobank.org:act:0DEB5BDB-6E85-4691-B5D1-E6258239370A" httpUri="http://treatment.plazi.org/id/039487A4FFF2FFDCFDC7F167FEECF96A" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
<subSubSection id="C3276539FFF2FFE2FDC7F167FBD9FDE4" box="[551,1036,513,541]" pageId="30" pageNumber="31" type="nomenclature">
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<heading id="D0CA81DEFFF2FFE2FDC7F167FBD9FDE4" box="[551,1036,513,541]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="4C3D4D31FFF2FFE2FDC7F167FC7EFDE4" ID-CoL="8QJ6B" authority="Robillard &amp; Tan" authorityName="Robillard &amp; Tan" authorityYear="2021" box="[551,939,513,541]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="danum" status="sp. nov.">
<emphasis id="B949EAA0FFF2FFE2FDC7F167FD31FDE5" bold="true" box="[551,740,513,540]" italics="true" pageId="30" pageNumber="31">Nisitrus danum</emphasis>
Robillard &amp; Tan
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<taxonomicNameLabel id="A27A57DBFFF2FFE2FC52F166FBD9FDE4" box="[946,1036,515,541]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<figureCitation id="13062A37FFF2FFE2FE01F12FFDFEFD9C" box="[481,555,586,613]" captionStart="Fig" captionStartId="9.[189,233,1963,1989]" captionTargetBox="[187,1396,991,1924]" captionTargetId="figure-297@9.[187,1396,990,1924]" captionTargetPageId="9" captionText="Fig. 5. Map of Southeast Asia showing the distribution of species of Nisitrus Saussure, 1878. Each coloured symbol represents one sampling or observation locality listed in the material in the text." figureDoi="http://doi.org/10.5281/zenodo.5156278" httpUri="https://zenodo.org/record/5156278/files/figure.png" pageId="30" pageNumber="31">Figs 5</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FDD9F12FFD8AFD9D" box="[569,607,586,612]" captionStart="Fig" captionStartId="13.[189,233,1926,1952]" captionTargetBox="[232,1318,284,1848]" captionTargetId="figure-11@13.[199,1388,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Habitus of ♀♀ in dorsal view. A–B. Nisitrus brunnerianus Saussure, 1878. C. N. crucius Robillard &amp; Tan sp. nov. D. N. danum Robillard &amp; Tan sp. nov. E. N. hughtani Robillard &amp; Tan sp. nov. F. N. insignis Saussure, 1878. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5156286" httpUri="https://zenodo.org/record/5156286/files/figure.png" pageId="30" pageNumber="31">8D</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FD8BF12FFD4BFD9D" box="[619,670,586,612]" captionStart="Fig" captionStartId="15.[189,232,1742,1768]" captionTargetBox="[216,1388,279,1679]" captionTargetId="figure-11@15.[189,1398,263,1705]" captionTargetPageId="15" captionText="Fig. 10. Face in anterior view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156294" httpUri="https://zenodo.org/record/5156294/files/figure.png" pageId="30" pageNumber="31">10C</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FD4BF12FFD08FD9D" box="[683,733,586,612]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="30" pageNumber="31">11C</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FD09F12FFCC9FD9D" box="[745,796,586,612]" captionStart="Fig" captionStartId="17.[189,232,1645,1671]" captionTargetBox="[210,1398,290,1606]" captionTargetId="figure-11@17.[189,1398,265,1606]" captionTargetPageId="17" captionText="Fig. 12. Body in lateral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5156302" httpUri="https://zenodo.org/record/5156302/files/figure.png" pageId="30" pageNumber="31">12C</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FCC9F12FFC89FD9D" box="[809,860,586,612]" captionStart="Fig" captionStartId="19.[189,233,1894,1920]" captionTargetBox="[216,1372,287,1817]" captionTargetId="figure-11@19.[193,1394,265,1855]" captionTargetPageId="19" captionText="Fig. 14. Female tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156308" httpUri="https://zenodo.org/record/5156308/files/figure.png" pageId="30" pageNumber="31">14C</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FC88F12FFC4FFD9D" box="[872,922,586,612]" captionStart="Fig" captionStartId="22.[189,232,1809,1835]" captionTargetBox="[201,1393,282,1740]" captionTargetId="figure-14@22.[189,1398,265,1771]" captionText="Fig. 17. Female genitalia in lateral (A, C, E, F, H, J, L, N, P) and dorsal (B, D, G, I, K, M, O, Q) views. A–B. Nisitrus brunnerianus Saussure, 1878. C–D. N. crucius Robillard &amp; Tan sp. nov. E. N. danum Robillard &amp; Tan sp. nov. F–G. N. hughtani Robillard &amp; Tan sp. nov. H–I. N. insignis Saussure, 1878. J–K. N. malaya Robillard &amp; Tan sp. nov. L–M. N. musicus Ingrisch, 1987. N–O. N. rindu Robillard &amp; Tan sp. nov. P–Q. N. vittatus (Haan, 1844). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5165004" httpUri="https://zenodo.org/record/5165004/files/figure.png" pageId="30" pageNumber="31">17E</figureCitation>
, 19;
<tableCitation id="C6BF0309FFF2FFE2FC33F12FFBE2FD9D" box="[979,1079,586,612]" captionStart="Table 2" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 2. Status of original species epithets associated with Nisitrus Saussure, 1878." httpUri="http://table.plazi.org/id/DF42663AFFE6FFF6FF5DF26CFB5AFEDA" pageId="30" pageNumber="31" tableUuid="DF42663AFFE6FFF6FF5DF26CFB5AFEDA">Tables 2</tableCitation>
,
<tableCitation id="C6BF0309FFF2FFE2FBA3F12FFB87FD9C" box="[1091,1106,586,613]" captionStart="Table 5" captionStartId="32.[189,258,265,291]" captionTargetBox="[203,1373,331,470]" captionTargetPageId="32" captionText="Table 5. Measurements of Nisitrus danum Robillard &amp; Tan sp. nov. (in mm, means in brackets)." httpUri="http://table.plazi.org/id/DF42663AFFCCFFDCFF5DF26CFAF0FEDD" pageId="30" pageNumber="31" tableUuid="DF42663AFFCCFFDCFF5DF26CFAF0FEDD">5</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF2FFE2FF5DF1E9FC8CFB99" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8B8236B2FFF2FFE2FF5DF1E9FEE3FD5F" blockId="30.[189,310,652,678]" box="[189,310,652,678]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF1E9FEE3FD5F" bold="true" box="[189,310,652,678]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF1E9FEE3FD5F" bold="true" box="[189,310,652,678]" pageId="30" pageNumber="31">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF5DF1DEFC8CFB99" blockId="30.[187,1400,699,1121]" pageId="30" pageNumber="31">
This new species differs from all known congeners by face part of fastigium with a black band across eyes and antennal cavity, frons otherwise yellow (reddish on preserved specimens); pronotal lateral lobe black dorsally and yellow ventrally; longitudinal white stripe only between M and R and lateral field matted black; Sc projections black. This new species is most similar to
<taxonomicName id="4C3D4D31FFF2FFE2FC4CF04DFBE3FCB8" authority="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard, 2021" authorityName="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2021" box="[940,1078,807,833]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="hughtani" status="sp. nov.">
<emphasis id="B949EAA0FFF2FFE2FC4CF04DFBE3FCB8" box="[940,1078,807,833]" italics="true" pageId="30" pageNumber="31">N. hughtani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF2FFE2FBDEF04DFB4CFCBB" box="[1086,1177,808,834]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
from
<collectingRegion id="49F9F850FFF2FFE2FB00F042FA90FCB8" box="[1248,1349,807,833]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">Sumatra</collectingRegion>
, but differs by the following characters: smaller in size, vertex black with yellow margin around eyes (instead of cream-coloured), frons with a black band across eyes and antennal cavity (yellow in
<taxonomicName id="4C3D4D31FFF2FFE2FB09F015FAA3FC70" authority="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard, 2021" authorityName="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2021" box="[1257,1398,879,905]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="hughtani" status="sp. nov.">
<emphasis id="B949EAA0FFF2FFE2FB09F015FAA3FC70" box="[1257,1398,879,905]" italics="true" pageId="30" pageNumber="31">N. hughtani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF2FFE2FF5FF0F6FECFFC54" box="[191,282,915,941]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
); frons yellow (instead of black with a cream-coloured band ventral to eyes); mouthparts yellowish with some black spots; lateral part of head with a black band posterior of eye (instead of black with a cream-coloured ring around eye). This species is also similar to
<taxonomicName id="4C3D4D31FFF2FFE2FC5FF0B9FB97FC0C" authorityName="Ingrisch" authorityYear="1987" box="[959,1090,988,1013]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="musicus">
<emphasis id="B949EAA0FFF2FFE2FC5FF0B9FB97FC0C" box="[959,1090,988,1013]" italics="true" pageId="30" pageNumber="31">N. musicus</emphasis>
</taxonomicName>
from Kinabalu range, but differs by mouthparts yellow (instead of black), FIIIs not black dorsally, female papilla, and distribution:
<taxonomicName id="4C3D4D31FFF2FFE2FF5DF746FE95FBC5" authorityName="Ingrisch" authorityYear="1987" box="[189,320,1059,1084]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="musicus">
<emphasis id="B949EAA0FFF2FFE2FF5DF746FE95FBC5" box="[189,320,1059,1084]" italics="true" pageId="30" pageNumber="31">N. musicus</emphasis>
</taxonomicName>
is restricted to the highlands in the Kinabalu range, while
<taxonomicName id="4C3D4D31FFF2FFE2FC1EF746FBA7FBC4" authority="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard, 2021" authorityName="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2021" box="[1022,1138,1059,1085]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="danum" status="sp. nov.">
<emphasis id="B949EAA0FFF2FFE2FC1EF746FBA7FBC4" box="[1022,1138,1059,1085]" italics="true" pageId="30" pageNumber="31">N. danum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF2FFE2FB9BF746FB02FBC4" box="[1147,1239,1059,1085]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
occurs in the Danum Valley and Tawau Hill areas at the east of
<collectingRegion id="49F9F850FFF2FFE2FCE9F723FC81FB99" box="[777,852,1094,1120]" country="Malaysia" name="Sabah" pageId="30" pageNumber="31">Sabah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF2FFE2FF5DF7ECFAF1FB2B" box="[189,1316,1161,1235]" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8B8236B2FFF2FFE2FF5DF7ECFE91FB5A" blockId="30.[189,324,1161,1187]" box="[189,324,1161,1187]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF7ECFE91FB5A" bold="true" box="[189,324,1161,1187]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF7ECFE91FB5A" bold="true" box="[189,324,1161,1187]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF5DF7DDFAF1FB2B" blockId="30.[189,1316,1208,1235]" box="[189,1316,1208,1235]" pageId="30" pageNumber="31">
The specific name is a noun in apposition taken from the
<typeStatus id="54868810FFF2FFE2FCBDF7DCFC45FB2A" box="[861,912,1209,1235]" pageId="30" pageNumber="31">type</typeStatus>
locality, Danum Valley,
<collectingCountry id="F32A7622FFF2FFE2FB50F7DDFACAFB2B" box="[1200,1311,1208,1234]" name="Malaysia" pageId="30" pageNumber="31">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF2FFE2FF5DF79FFCCEF93D" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B8236B2FFF2FFE2FF5DF79FFE7FFAED" blockId="30.[189,426,1274,1300]" box="[189,426,1274,1300]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF79FFE7FFAED" bold="true" box="[189,426,1274,1300]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF79FFE7FFAED" bold="true" box="[189,426,1274,1300]" pageId="30" pageNumber="31">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF0AF64FFB93FA75" blockId="30.[189,1398,1322,1420]" pageId="30" pageNumber="31">
<materialsCitation id="3B553CEFFFF2FFE2FF0AF64FFB93FA75" ID-GBIF-Occurrence="3333037672" collectingDate="1987-05-26" collectingDateMax="1987-06-20" collectingDateMin="1987-05-26" collectionCode="RMNH" collectorName="C. v. Achterberg &amp; Kennedy" country="Malaysia" county="Danum Valley Field Centre" elevation="150" latitude="4.9647083" location="Danum Valley Field Centre" longLatPrecision="1" longitude="117.80507" municipality="Danum Valley" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B949EAA0FFF2FFE2FF0AF64FFE89FABD" bold="true" box="[234,348,1322,1348]" pageId="30" pageNumber="31">
<typeStatus id="54868810FFF2FFE2FF0AF64FFE89FABD" box="[234,348,1322,1348]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F32A7622FFF2FFE2FF5DF628FE88FA9E" box="[189,349,1357,1383]" name="Malaysia" pageId="30" pageNumber="31">MALAYSIA</collectingCountry>
–
<emphasis id="B949EAA0FFF2FFE2FE97F628FE10FA9E" bold="true" box="[375,453,1357,1383]" pageId="30" pageNumber="31">
<collectingRegion id="49F9F850FFF2FFE2FE97F628FE10FA9E" box="[375,453,1357,1383]" country="Malaysia" name="Sabah" pageId="30" pageNumber="31">Sabah</collectingRegion>
</emphasis>
•
<specimenCount id="9D3BFD3BFFF2FFE2FE3CF62BFE26FA91" box="[476,499,1358,1384]" pageId="30" pageNumber="31" type="female">♀</specimenCount>
;
<collectingCounty id="62E34E3EFFF2FFE2FDE0F62BFC95FA9E" box="[512,832,1357,1384]" pageId="30" pageNumber="31">Danum Valley Field Centre</collectingCounty>
;
<geoCoordinate id="EE095075FFF2FFE2FCA9F629FC26FA91" box="[841,1011,1356,1384]" degrees="4" direction="north" minutes="57" orientation="latitude" pageId="30" pageNumber="31" precision="1" seconds="52.95" value="4.9647083">4°57′52.95″ N</geoCoordinate>
,
<geoCoordinate id="EE095075FFF2FFE2FC1DF629FB6AFA91" box="[1021,1215,1356,1384]" degrees="117" direction="east" minutes="48" orientation="longitude" pageId="30" pageNumber="31" precision="1" seconds="18.25" value="117.80507">117°48′18.25″ E</geoCoordinate>
; ca
<elevation id="0010D181FFF2FFE2FB0AF628FAA5FA9E" box="[1258,1392,1357,1384]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="m" value="150.0">
<quantity id="4CC59B57FFF2FFE2FB0AF628FAE6FA91" box="[1258,1331,1357,1384]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="m" value="150.0">150 m</quantity>
a.s.l.
</elevation>
;
<date id="FF831072FFF2FFE2FF5DF614FE15FA72" box="[189,448,1393,1420]" pageId="30" pageNumber="31" value="1987-05-26" valueMax="1987-06-20" valueMin="1987-05-26">
<collectingDate id="EFC7E99AFFF2FFE2FF5DF614FE15FA72" box="[189,448,1393,1420]" pageId="30" pageNumber="31" value="1987-05-26" valueMax="1987-06-20" valueMin="1987-05-26">26 May–20 Jun. 1987</collectingDate>
</date>
;
<collectorName id="26C85364FFF2FFE2FE2CF614FD5EFA72" box="[460,651,1393,1420]" pageId="30" pageNumber="31">C. v. Achterberg</collectorName>
and
<collectorName id="26C85364FFF2FFE2FD25F614FCFAFA72" box="[709,815,1393,1419]" pageId="30" pageNumber="31">Kennedy</collectorName>
leg.; PII: N54;
<collectionCode id="ED2CAE77FFF2FFE2FC06F617FB97FA75" box="[998,1090,1394,1420]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="31" type="Museum">RMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF0AF6D1FD66F9EF" blockId="30.[189,1399,1460,1558]" pageId="30" pageNumber="31">
<materialsCitation id="3B553CEFFFF2FFE2FF0AF6D1FBE4FA0B" ID-GBIF-Occurrence="3333037337" collectingDate="1987-05-26" collectingDateMax="1987-06-20" collectingDateMin="1987-05-26" collectionCode="RMNH" collectorName="C. v. Achterberg &amp; Kennedy" country="Malaysia" county="Danum Valley Field Centre" elevation="150" latitude="4.9647083" location="Sabah" longLatPrecision="1" longitude="117.80507" municipality="Danum Valley" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B949EAA0FFF2FFE2FF0AF6D1FEB2FA37" bold="true" box="[234,359,1460,1486]" pageId="30" pageNumber="31">
<typeStatus id="54868810FFF2FFE2FF0AF6D1FEB2FA37" box="[234,359,1460,1486]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F32A7622FFF2FFE2FF5DF6B2FE8BFA08" box="[189,350,1495,1521]" name="Malaysia" pageId="30" pageNumber="31">MALAYSIA</collectingCountry>
–
<emphasis id="B949EAA0FFF2FFE2FE9DF6B2FE1EFA08" bold="true" box="[381,459,1495,1521]" pageId="30" pageNumber="31">
<collectingRegion id="49F9F850FFF2FFE2FE9DF6B2FE1EFA08" box="[381,459,1495,1521]" country="Malaysia" name="Sabah" pageId="30" pageNumber="31">Sabah</collectingRegion>
</emphasis>
•
<specimenCount id="9D3BFD3BFFF2FFE2FE05F6B2FDC7FA0B" box="[485,530,1495,1522]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED2CAE77FFF2FFE2FC39F6BDFBE4FA0B" box="[985,1073,1496,1522]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="31" type="Museum">RMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B553CEFFFF2FFE2FBAAF6B2FD65F9EF" ID-GBIF-Occurrence="3333037366" collectingDate="1987-05-26" collectingDateMax="1987-06-20" collectingDateMin="1987-05-26" collectionCode="RMNH" collectorName="C. v. Achterberg &amp; Kennedy" country="Malaysia" county="Danum Valley Field Centre" elevation="150" latitude="4.9647083" location="Valley Field Centre" longLatPrecision="1" longitude="117.80507" municipality="Danum Valley" pageId="30" pageNumber="31" specimenCode="MNHN-EO-ENSIF10945" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF2FFE2FBAAF6B2FBADFA0B" box="[1098,1144,1495,1522]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DB9B9EC9FFF2FFE2FE9CF69EFD65F9EF" box="[380,688,1531,1558]" pageId="30" pageNumber="31">MNHN-EO-ENSIF10945</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF0AF558FE23F9AE" blockId="30.[189,1398,1597,1732]" box="[234,502,1597,1623]" pageId="30" pageNumber="31">
<emphasis id="B949EAA0FFF2FFE2FF0AF558FE23F9AE" bold="true" box="[234,502,1597,1623]" pageId="30" pageNumber="31">Additional specimens</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF5DF504FCCEF93D" blockId="30.[189,1398,1597,1732]" pageId="30" pageNumber="31">
<materialsCitation id="3B553CEFFFF2FFE2FF5DF504FD55F959" ID-GBIF-Occurrence="3333037333" collectingDate="2018-03-07" collectorName="C. Elder" country="Malaysia" latitude="4.4" location="Tawau Hill Park" longLatPrecision="1" longitude="117.89" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<collectingCountry id="F32A7622FFF2FFE2FF5DF504FE8BF982" box="[189,350,1633,1659]" name="Malaysia" pageId="30" pageNumber="31">MALAYSIA</collectingCountry>
–
<emphasis id="B949EAA0FFF2FFE2FE9AF504FE1DF982" bold="true" box="[378,456,1633,1659]" pageId="30" pageNumber="31">
<collectingRegion id="49F9F850FFF2FFE2FE9AF504FE1DF982" box="[378,456,1633,1659]" country="Malaysia" name="Sabah" pageId="30" pageNumber="31">Sabah</collectingRegion>
</emphasis>
•
<specimenCount id="9D3BFD3BFFF2FFE2FE00F504FDD8F985" box="[480,525,1633,1660]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
;
<location id="8EE26069FFF2FFE2FDFAF507FD09F982" LSID="urn:lsid:plazi:treatment:039487A4FFF2FFDCFDC7F167FEECF96A:8EE26069FFF2FFE2FDFAF507FD09F982" box="[538,732,1633,1660]" country="Malaysia" latitude="4.4" longLatPrecision="1" longitude="117.89" name="Tawau Hill Park" pageId="30" pageNumber="31" stateProvince="Sabah">Tawau Hill Park</location>
;
<geoCoordinate id="EE095075FFF2FFE2FD07F505FCA1F985" box="[743,884,1632,1660]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="30" pageNumber="31" precision="1" seconds="0.0" value="4.4">4°24′0.0″ N</geoCoordinate>
,
<geoCoordinate id="EE095075FFF2FFE2FC9FF505FBE0F985" box="[895,1077,1632,1660]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="30" pageNumber="31" precision="1" seconds="24.0" value="117.89">117°53′24.0″ E</geoCoordinate>
;
<date id="FF831072FFF2FFE2FBA0F507FB05F982" box="[1088,1232,1633,1660]" pageId="30" pageNumber="31" value="2018-03-07">
<collectingDate id="EFC7E99AFFF2FFE2FBA0F507FB05F982" box="[1088,1232,1633,1660]" pageId="30" pageNumber="31" value="2018-03-07">7 Mar. 2018</collectingDate>
</date>
;
<collectorName id="26C85364FFF2FFE2FB3CF504FAE8F982" box="[1244,1341,1633,1659]" pageId="30" pageNumber="31">C. Elder</collectorName>
leg.; iNaturalist.org/observations/19511712
</materialsCitation>
•
<materialsCitation id="3B553CEFFFF2FFE2FD75F5E0FCC1F93D" ID-GBIF-Occurrence="3333037581" collectingDate="2020-10-02" collectorName="L. Day" country="Malaysia" latitude="4.961833" location="Danum Valley" longLatPrecision="1" longitude="117.6895" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D3BFD3BFFF2FFE2FD75F5E0FD14F959" box="[661,705,1669,1696]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
;
<location id="8EE26069FFF2FFE2FD2EF5E3FCADF966" LSID="urn:lsid:plazi:treatment:039487A4FFF2FFDCFDC7F167FEECF96A:8EE26069FFF2FFE2FD2EF5E3FCADF966" box="[718,888,1669,1696]" country="Malaysia" latitude="4.961833" longLatPrecision="1" longitude="117.6895" name="Danum Valley" pageId="30" pageNumber="31" stateProvince="Sabah">Danum Valley</location>
;
<geoCoordinate id="EE095075FFF2FFE2FC62F5E1FBCBF959" box="[898,1054,1668,1696]" degrees="4" direction="north" minutes="57" orientation="latitude" pageId="30" pageNumber="31" precision="1" seconds="42.6" value="4.961833">4°57′42.6″ N</geoCoordinate>
,
<geoCoordinate id="EE095075FFF2FFE2FBC8F5E1FB08F959" box="[1064,1245,1668,1696]" degrees="117" direction="east" minutes="41" orientation="longitude" pageId="30" pageNumber="31" precision="1" seconds="22.2" value="117.6895">117°41′22.2″ E</geoCoordinate>
;
<date id="FF831072FFF2FFE2FB08F5E0FAA7F966" box="[1256,1394,1669,1695]" pageId="30" pageNumber="31" value="2020-10-02">
<collectingDate id="EFC7E99AFFF2FFE2FB08F5E0FAA7F966" box="[1256,1394,1669,1695]" pageId="30" pageNumber="31" value="2020-10-02">2 Oct. 2020</collectingDate>
</date>
;
<collectorName id="26C85364FFF2FFE2FF5DF5CFFEDAF93D" box="[189,271,1706,1732]" pageId="30" pageNumber="31">L. Day</collectorName>
leg.; iNaturalist.org/observations/21010925
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF2FFDCFF5DF58EFE99FB2A" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="8B8236B2FFF2FFE2FF5DF58EFE9BF8FC" blockId="30.[189,334,1771,1797]" box="[189,334,1771,1797]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF58EFE9BF8FC" bold="true" box="[189,334,1771,1797]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF58EFE9BF8FC" bold="true" box="[189,334,1771,1797]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF2FFE3FF5DF47EFAA0FE49" blockId="30.[187,1399,1819,2025]" lastBlockId="31.[187,1399,265,432]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
Slender and relatively large habitus (
<figureCitation id="13062A37FFF2FFE2FDBAF47EFD6CF8CC" box="[602,697,1819,1845]" captionStart="Fig" captionStartId="13.[189,233,1926,1952]" captionTargetBox="[232,1318,284,1848]" captionTargetId="figure-11@13.[199,1388,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Habitus of ♀♀ in dorsal view. A–B. Nisitrus brunnerianus Saussure, 1878. C. N. crucius Robillard &amp; Tan sp. nov. D. N. danum Robillard &amp; Tan sp. nov. E. N. hughtani Robillard &amp; Tan sp. nov. F. N. insignis Saussure, 1878. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5156286" httpUri="https://zenodo.org/record/5156286/files/figure.png" pageId="30" pageNumber="31">Figs 8D</figureCitation>
,
<figureCitation id="13062A37FFF2FFE2FD23F47EFD36F8CC" box="[707,739,1819,1845]" captionStart="Fig" captionStartId="31.[189,232,1963,1989]" captionTargetBox="[189,1398,667,1924]" captionTargetId="figure-96@31.[189,1398,667,1924]" captionTargetPageId="31" captionText="Fig. 19. Nisitrus danum Robillard &amp; Tan sp. nov., adult ♀♀ in their natural environment. Photograph source and credits: iNaturalist; Arthur Anker (A), Charlie Elder (B)." figureDoi="http://doi.org/10.5281/zenodo.5156322" httpUri="https://zenodo.org/record/5156322/files/figure.png" pageId="30" pageNumber="31">19</figureCitation>
). Vertex black with yellow margin around eyes (
<figureCitation id="13062A37FFF2FFE2FAE3F47EFABFF8CC" box="[1283,1386,1819,1845]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="30" pageNumber="31">Fig. 11C</figureCitation>
). Fastigium velvety black with yellow margins. Scapes dark red brown to black (particularly on the dorsoinner margins). Antennae black with wide whitish rings. Face part of fastigium with a black band across eyes and antennal cavity, otherwise yellow (reddish in preserved specimens); frons almost entirely yellow, including mouthparts, with some black spots (
<figureCitation id="13062A37FFF2FFE2FD2DF4CFFCE2F83C" box="[717,823,1962,1989]" captionStart="Fig" captionStartId="15.[189,232,1742,1768]" captionTargetBox="[216,1388,279,1679]" captionTargetId="figure-11@15.[189,1398,263,1705]" captionTargetPageId="15" captionText="Fig. 10. Face in anterior view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156294" httpUri="https://zenodo.org/record/5156294/files/figure.png" pageId="30" pageNumber="31">Fig. 10C</figureCitation>
). Maxillary palpi cream-coloured; apical segment black apically (
<figureCitation id="13062A37FFF2FFE2FE90F4ABFE0BF811" box="[368,478,1998,2024]" captionStart="Fig" captionStartId="15.[189,232,1742,1768]" captionTargetBox="[216,1388,279,1679]" captionTargetId="figure-11@15.[189,1398,263,1705]" captionTargetPageId="15" captionText="Fig. 10. Face in anterior view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156294" httpUri="https://zenodo.org/record/5156294/files/figure.png" pageId="30" pageNumber="31">Fig. 10C</figureCitation>
). Lateral part of head with a black band posterior to eye (
<figureCitation id="13062A37FFF2FFE2FB62F4ABFB3BF811" box="[1154,1262,1998,2024]" captionStart="Fig" captionStartId="17.[189,232,1645,1671]" captionTargetBox="[210,1398,290,1606]" captionTargetId="figure-11@17.[189,1398,265,1606]" captionTargetPageId="17" captionText="Fig. 12. Body in lateral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5156302" httpUri="https://zenodo.org/record/5156302/files/figure.png" pageId="30" pageNumber="31">Fig. 12C</figureCitation>
). Pronotum covered with white setae; dorsal disk dark brown to black (
<figureCitation id="13062A37FFF3FFE3FC95F26CFC0AFEDA" box="[885,991,265,291]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="31" pageNumber="32">Fig. 11C</figureCitation>
). Lateral lobes of pronotum dorsal third black, vivid yellow ventrally (cream-coloured in preserved specimens) (
<figureCitation id="13062A37FFF3FFE3FBA9F249FB60FEBF" box="[1097,1205,300,326]" captionStart="Fig" captionStartId="17.[189,232,1645,1671]" captionTargetBox="[210,1398,290,1606]" captionTargetId="figure-11@17.[189,1398,265,1606]" captionTargetPageId="17" captionText="Fig. 12. Body in lateral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5156302" httpUri="https://zenodo.org/record/5156302/files/figure.png" pageId="31" pageNumber="32">Fig. 12C</figureCitation>
). Legs generally red brown, sometimes dorsally darker. FIIIs brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Tergites red brown, with a pale band laterally; sternites pale, with red brown median area. Cerci red brown, short and conical.
</paragraph>
<paragraph id="8B8236B2FFF3FFE3FF0AF2B2FEFFFE08" blockId="31.[189,313,471,532]" box="[234,298,471,497]" pageId="31" pageNumber="32">
<emphasis id="B949EAA0FFF3FFE3FF0AF2B2FEFFFE08" bold="true" box="[234,298,471,497]" pageId="31" pageNumber="32">Male</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF3FFE3FF5DF29FFEECFDED" blockId="31.[189,313,471,532]" box="[189,313,506,532]" pageId="31" pageNumber="32">Unknown.</paragraph>
<caption id="DF42663AFFF3FFE3FF5DF4CEFC0CF811" ID-DOI="http://doi.org/10.5281/zenodo.5156322" ID-Zenodo-Dep="5156322" httpUri="https://zenodo.org/record/5156322/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1963,1989]" targetBox="[189,1398,667,1924]" targetPageId="31">
<paragraph id="8B8236B2FFF3FFE3FF5DF4CEFC0CF811" blockId="31.[189,1398,1963,2025]" pageId="31" pageNumber="32">
<emphasis id="B949EAA0FFF3FFE3FF5DF4CEFECEF83C" bold="true" box="[189,283,1963,1989]" pageId="31" pageNumber="32">Fig. 19.</emphasis>
<taxonomicName id="4C3D4D31FFF3FFE3FEC3F4C9FD70F83F" authority="Robillard &amp; Tan" authorityName="Robillard &amp; Tan" authorityYear="2021" box="[291,677,1963,1990]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="danum" status="sp. nov.">
<emphasis id="B949EAA0FFF3FFE3FEC3F4C9FE0CF83C" box="[291,473,1963,1989]" italics="true" pageId="31" pageNumber="32">Nisitrus danum</emphasis>
Robillard &amp; Tan
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF3FFE3FD4DF4C9FCD9F83F" box="[685,780,1964,1990]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, adult ♀♀ in their natural environment. Photograph source and credits: iNaturalist; Arthur Anker (A), Charlie Elder (B).
</paragraph>
</caption>
<caption id="DF42663AFFCCFFDCFF5DF26CFAF0FEDD" ID-Table-UUID="DF42663AFFCCFFDCFF5DF26CFAF0FEDD" box="[189,1317,265,292]" httpUri="http://table.plazi.org/id/DF42663AFFCCFFDCFF5DF26CFAF0FEDD" pageId="32" pageNumber="33" startId="32.[189,258,265,291]" targetBox="[203,1373,331,470]" targetIsTable="true" targetPageId="32">
<paragraph id="8B8236B2FFCCFFDCFF5DF26CFAF0FEDD" blockId="32.[189,1317,265,292]" box="[189,1317,265,292]" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFF5DF26CFECAFEDA" bold="true" box="[189,287,265,291]" pageId="32" pageNumber="33">Table 5.</emphasis>
Measurements of
<taxonomicName id="4C3D4D31FFCCFFDCFE19F26FFCA1FEDD" authority="Robillard &amp; Tan" authorityName="Robillard &amp; Tan" authorityYear="2021" box="[505,884,265,292]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="danum" status="sp. nov.">
<emphasis id="B949EAA0FFCCFFDCFE19F26FFD78FEDA" box="[505,685,265,291]" italics="true" pageId="32" pageNumber="33">Nisitrus danum</emphasis>
Robillard &amp; Tan
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFCCFFDCFC9BF26FFC03FEDD" box="[891,982,266,292]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
(in mm, means in brackets).
</paragraph>
</caption>
<paragraph id="8B8236B2FFCCFFDCFE4CF22EFA8EFE2F" pageId="32" pageNumber="33">
<table id="F93DC412FFCC0003FF2BF22EFA88FE2F" box="[203,1373,331,470]" gridcols="9" gridrows="3" pageId="32" pageNumber="33" rowsContinueIn="32.[203,1212,486,667]">
<tr id="350D34F0FFCC0003FF2BF22EFA88FE9A" box="[203,1373,331,355]" gridrow="0" pageId="32" pageNumber="33" rowspan-0="1">
<th id="76DC5D8CFFCC0003FE48F22EFE2DFE9A" box="[424,504,331,355]" gridcol="1" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFE4CF22EFE23FE9A" bold="true" box="[428,502,331,355]" pageId="32" pageNumber="33">PronL</emphasis>
</th>
<th id="76DC5D8CFFCC0003FDC2F22EFDA1FE9A" box="[546,628,331,355]" gridcol="2" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFDC2F22EFDA1FE9A" bold="true" box="[546,628,331,355]" pageId="32" pageNumber="33">PronW</emphasis>
</th>
<th id="76DC5D8CFFCC0003FD79F22EFCD6FE9A" box="[665,771,331,355]" gridcol="3" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFD51F22EFD3BFE9A" bold="true" box="[689,750,331,355]" pageId="32" pageNumber="33">FWL</emphasis>
</th>
<th id="76DC5D8CFFCC0003FCC7F22EFCA2FE9A" box="[807,887,331,355]" gridcol="4" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFCCCF22EFCA7FE9A" bold="true" box="[812,882,331,355]" pageId="32" pageNumber="33">FWW</emphasis>
</th>
<th id="76DC5D8CFFCC0003FC43F22EFC26FE9A" box="[931,1011,331,355]" gridcol="5" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFC4AF22EFC3EFE9A" bold="true" box="[938,1003,331,355]" pageId="32" pageNumber="33">HWT</emphasis>
</th>
<th id="76DC5D8CFFCC0003FBCDF22EFBBBFE9A" box="[1069,1134,331,355]" gridcol="6" gridrow="0" pageId="32" pageNumber="33">FIIIL</th>
<th id="76DC5D8CFFCC0003FB42F22EFB39FE9A" box="[1186,1260,331,355]" gridcol="7" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFB42F22EFB39FE9A" bold="true" box="[1186,1260,331,355]" pageId="32" pageNumber="33">FIIIW</emphasis>
</th>
<th id="76DC5D8CFFCC0003FAFAF22EFA88FE9A" box="[1306,1373,331,355]" gridcol="8" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFAFAF22EFA88FE9A" bold="true" box="[1306,1373,331,355]" pageId="32" pageNumber="33">TIIIL</emphasis>
</th>
</tr>
<tr id="350D34F0FFCC0003FF2BF211FA88FE74" box="[203,1373,372,397]" gridrow="1" pageId="32" pageNumber="33">
<th id="76DC5D8CFFCC0003FF2BF211FEA8FE74" box="[203,381,372,397]" gridcol="0" gridrow="1" pageId="32" pageNumber="33">Female holotype</th>
<td id="76DC5D8CFFCC0003FE48F211FE2DFE74" box="[424,504,372,397]" gridcol="1" gridrow="1" pageId="32" pageNumber="33">2.0</td>
<td id="76DC5D8CFFCC0003FDC2F211FDA1FE74" box="[546,628,372,397]" gridcol="2" gridrow="1" pageId="32" pageNumber="33">3.2</td>
<td id="76DC5D8CFFCC0003FD79F211FCD6FE74" box="[665,771,372,397]" gridcol="3" gridrow="1" pageId="32" pageNumber="33">10.8</td>
<td id="76DC5D8CFFCC0003FCC7F211FCA2FE74" box="[807,887,372,397]" gridcol="4" gridrow="1" pageId="32" pageNumber="33">3.5</td>
<td id="76DC5D8CFFCC0003FC43F211FC26FE74" box="[931,1011,372,397]" gridcol="5" gridrow="1" pageId="32" pageNumber="33">6.4</td>
<td id="76DC5D8CFFCC0003FBCDF211FBBBFE74" box="[1069,1134,372,397]" gridcol="6" gridrow="1" pageId="32" pageNumber="33">15.3</td>
<td id="76DC5D8CFFCC0003FB42F211FB39FE74" box="[1186,1260,372,397]" gridcol="7" gridrow="1" pageId="32" pageNumber="33">2.7</td>
<td id="76DC5D8CFFCC0003FAFAF211FA88FE74" box="[1306,1373,372,397]" gridcol="8" gridrow="1" pageId="32" pageNumber="33">15.5</td>
</tr>
<tr id="350D34F0FFCC0003FF2BF2FBFA88FE2F" box="[203,1373,414,470]" gridrow="2" pageId="32" pageNumber="33">
<th id="76DC5D8CFFCC0003FF2BF2FBFEA8FE2F" box="[203,381,414,470]" gridcol="0" gridrow="2" pageId="32" pageNumber="33">Females (n = 3)</th>
<td id="76DC5D8CFFCC0003FE48F2FBFE2DFE2F" box="[424,504,414,470]" gridcol="1" gridrow="2" pageId="32" pageNumber="33">2.0–2.3 (2.2)</td>
<td id="76DC5D8CFFCC0003FDC2F2FBFDA1FE2F" box="[546,628,414,470]" gridcol="2" gridrow="2" pageId="32" pageNumber="33">3.2–3.5 (3.4)</td>
<td id="76DC5D8CFFCC0003FD79F2FBFCD6FE2F" box="[665,771,414,470]" gridcol="3" gridrow="2" pageId="32" pageNumber="33">10.8–11.7 (11.4)</td>
<td id="76DC5D8CFFCC0003FCC7F2FBFCA2FE2F" box="[807,887,414,470]" gridcol="4" gridrow="2" pageId="32" pageNumber="33">3.5–4.2 (4.0)</td>
<td id="76DC5D8CFFCC0003FC43F2FBFC26FE2F" box="[931,1011,414,470]" gridcol="5" gridrow="2" pageId="32" pageNumber="33">6.4–7.7 (7.2)</td>
<td id="76DC5D8CFFCC0003FBCDF2FBFBBBFE2F" box="[1069,1134,414,470]" gridcol="6" gridrow="2" pageId="32" pageNumber="33">15.3 (15.3)</td>
<td id="76DC5D8CFFCC0003FB42F2FBFB39FE2F" box="[1186,1260,414,470]" gridcol="7" gridrow="2" pageId="32" pageNumber="33">2.7 (2.7)</td>
<td id="76DC5D8CFFCC0003FAFAF2FBFA88FE2F" box="[1306,1373,414,470]" gridcol="8" gridrow="2" pageId="32" pageNumber="33">15.5 (15.5)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFE5FF175FB73FD62" pageId="32" pageNumber="33">
<table id="F93DC412FFCC0003FF2BF283FB69FD62" box="[203,1212,486,667]" gridcols="7" gridrows="3" pageId="32" pageNumber="33" rowsContinueFrom="32.[203,1373,331,470]">
<tr id="350D34F0FFCC0003FF2BF283FB69FDD1" box="[203,1212,486,552]" gridrow="0" pageId="32" pageNumber="33" rowspan-0="1">
<th id="76DC5D8CFFCC0003FE5EF283FE30FDD1" box="[446,485,486,552]" gridcol="1" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFE5FF175FE34FDD1" bold="true" box="[447,481,528,552]" pageId="32" pageNumber="33">Ias</emphasis>
</th>
<th id="76DC5D8CFFCC0003FDF8F283FCD1FDD1" box="[536,772,486,552]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFDF8F282FCD1FE06" bold="true" box="[536,772,486,511]" pageId="32" pageNumber="33">Dorsal spines of TIII</emphasis>
<emphasis id="B949EAA0FFCCFFDCFDD9F175FD88FDD1" bold="true" box="[569,605,528,552]" pageId="32" pageNumber="33">Ibs</emphasis>
<emphasis id="B949EAA0FFCCFFDCFD58F16AFD31FDDE" bold="true" box="[696,740,527,551]" pageId="32" pageNumber="33">Oas</emphasis>
</th>
<th id="76DC5D8CFFCC0003FCD8F283FCB3FDD1" box="[824,870,486,552]" gridcol="4" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFCD8F16AFCB3FDDE" bold="true" box="[824,870,527,551]" pageId="32" pageNumber="33">Obs</emphasis>
</th>
<th id="76DC5D8CFFCC0003FC48F283FC3BFDD1" box="[936,1006,486,552]" gridcol="5" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFC48F175FC3BFDD1" bold="true" box="[936,1006,528,552]" pageId="32" pageNumber="33">TaIIIs</emphasis>
</th>
<th id="76DC5D8CFFCC0003FBB1F283FB69FDD1" box="[1105,1212,486,552]" gridcol="6" gridrow="0" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFB93F16AFB4FFDDE" bold="true" box="[1139,1178,527,551]" pageId="32" pageNumber="33">OL</emphasis>
</th>
</tr>
<tr id="350D34F0FFCC0003FF2BF15CFB69FDA8" box="[203,1212,569,593]" gridrow="1" pageId="32" pageNumber="33">
<th id="76DC5D8CFFCC0003FF2BF15CFEA8FDA8" box="[203,381,569,593]" gridcol="0" gridrow="1" pageId="32" pageNumber="33">Female holotype</th>
<td id="76DC5D8CFFCC0003FE5EF15CFE30FDA8" box="[446,485,569,593]" gridcol="1" gridrow="1" pageId="32" pageNumber="33">13</td>
<td id="76DC5D8CFFCC0003FDF8F15CFD8AFDA8" box="[536,607,569,593]" gridcol="2" gridrow="1" pageId="32" pageNumber="33">2–3</td>
<td id="76DC5D8CFFCC0003FD58F15CFCD1FDA8" box="[696,772,569,593]" gridcol="3" gridrow="1" pageId="32" pageNumber="33">24</td>
<td id="76DC5D8CFFCC0003FCD8F15CFCB3FDA8" box="[824,870,569,593]" gridcol="4" gridrow="1" pageId="32" pageNumber="33">2–3</td>
<td id="76DC5D8CFFCC0003FC48F15CFC3BFDA8" box="[936,1006,569,593]" gridcol="5" gridrow="1" pageId="32" pageNumber="33">2</td>
<td id="76DC5D8CFFCC0003FBB1F15CFB69FDA8" box="[1105,1212,569,593]" gridcol="6" gridrow="1" pageId="32" pageNumber="33">17.2</td>
</tr>
<tr id="350D34F0FFCC0003FF2BF107FB69FD62" box="[203,1212,610,667]" gridrow="2" pageId="32" pageNumber="33">
<th id="76DC5D8CFFCC0003FF2BF107FEA8FD62" box="[203,381,610,667]" gridcol="0" gridrow="2" pageId="32" pageNumber="33">Females (n = 3)</th>
<td id="76DC5D8CFFCC0003FE5EF107FE30FD62" box="[446,485,610,667]" gridcol="1" gridrow="2" pageId="32" pageNumber="33">NA</td>
<td id="76DC5D8CFFCC0003FDF8F107FD8AFD62" box="[536,607,610,667]" gridcol="2" gridrow="2" pageId="32" pageNumber="33">NA</td>
<td id="76DC5D8CFFCC0003FD58F107FCD1FD62" box="[696,772,610,667]" gridcol="3" gridrow="2" pageId="32" pageNumber="33">NA</td>
<td id="76DC5D8CFFCC0003FCD8F107FCB3FD62" box="[824,870,610,667]" gridcol="4" gridrow="2" pageId="32" pageNumber="33">NA</td>
<td id="76DC5D8CFFCC0003FC48F107FC3BFD62" box="[936,1006,610,667]" gridcol="5" gridrow="2" pageId="32" pageNumber="33">NA</td>
<td id="76DC5D8CFFCC0003FBB1F107FB69FD62" box="[1105,1212,610,667]" gridcol="6" gridrow="2" pageId="32" pageNumber="33">17.2–18.5 (17.9)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF0AF1BCFAA3FCEF" blockId="32.[189,1399,729,897]" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFF0AF1BCFE90FD0A" bold="true" box="[234,325,729,755]" pageId="32" pageNumber="33">Female</emphasis>
FOREWINGS. FW colouration (
<figureCitation id="13062A37FFCCFFDCFDC6F199FD41FCEE" box="[550,660,764,791]" captionStart="Fig" captionStartId="19.[189,233,1894,1920]" captionTargetBox="[216,1372,287,1817]" captionTargetId="figure-11@19.[193,1394,265,1855]" captionTargetPageId="19" captionText="Fig. 14. Female tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156308" httpUri="https://zenodo.org/record/5156308/files/figure.png" pageId="32" pageNumber="33">Fig. 14C</figureCitation>
): veins generally yellow; cells infumated black. Longitudinal
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5FF045FE74FC78" blockId="32.[189,1399,729,897]" pageId="32" pageNumber="33">infumated white stripe only in M and R. Lateral field matted black, Sc projections black. FW venation: 6–7 strong longitudinal veins on dorsal field, cross-veins irregular and often sinuous; lateral field with 6 projections of Sc.</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF0CDFBADFBF0" blockId="32.[189,1399,936,1033]" pageId="32" pageNumber="33">
GENITALIA. Ovipositor: distinctly longer than FIII; apex thin with both dorsal and ventral edges smooth. Copulatory papilla conical, smaller and stout; apex strongly folded (could be due to the preservation of specimen) ventrally, short, pointed; dorsal face with a sclerotized area (
<figureCitation id="13062A37FFCCFFDCFBE1F08BFBBEFBF0" box="[1025,1131,1006,1033]" captionStart="Fig" captionStartId="22.[189,232,1809,1835]" captionTargetBox="[201,1393,282,1740]" captionTargetId="figure-14@22.[189,1398,265,1771]" captionText="Fig. 17. Female genitalia in lateral (A, C, E, F, H, J, L, N, P) and dorsal (B, D, G, I, K, M, O, Q) views. A–B. Nisitrus brunnerianus Saussure, 1878. C–D. N. crucius Robillard &amp; Tan sp. nov. E. N. danum Robillard &amp; Tan sp. nov. F–G. N. hughtani Robillard &amp; Tan sp. nov. H–I. N. insignis Saussure, 1878. J–K. N. malaya Robillard &amp; Tan sp. nov. L–M. N. musicus Ingrisch, 1987. N–O. N. rindu Robillard &amp; Tan sp. nov. P–Q. N. vittatus (Haan, 1844). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5165004" httpUri="https://zenodo.org/record/5165004/files/figure.png" pageId="32" pageNumber="33">Fig. 17E</figureCitation>
).
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF0AF755FE87FBB3" blockId="32.[189,338,1072,1133]" box="[234,338,1072,1098]" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFF0AF755FE87FBB3" bold="true" box="[234,338,1072,1098]" pageId="32" pageNumber="33">Juvenile</emphasis>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF736FEECFB94" blockId="32.[189,338,1072,1133]" box="[189,313,1107,1133]" pageId="32" pageNumber="33">Unknown.</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF0AF7F0FE77FB56" blockId="32.[189,418,1173,1235]" box="[234,418,1173,1199]" pageId="32" pageNumber="33">
<emphasis id="B949EAA0FFCCFFDCFF0AF7F0FE77FB56" bold="true" box="[234,418,1173,1199]" pageId="32" pageNumber="33">Measurements</emphasis>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF7DDFE99FB2A" blockId="32.[189,418,1173,1235]" box="[189,332,1208,1235]" pageId="32" pageNumber="33">
See
<tableCitation id="C6BF0309FFCCFFDCFF0EF7DDFE92FB2A" box="[238,327,1208,1235]" captionStart="Table 5" captionStartId="32.[189,258,265,291]" captionTargetBox="[203,1373,331,470]" captionTargetPageId="32" captionText="Table 5. Measurements of Nisitrus danum Robillard &amp; Tan sp. nov. (in mm, means in brackets)." httpUri="http://table.plazi.org/id/DF42663AFFCCFFDCFF5DF26CFAF0FEDD" pageId="32" pageNumber="33" tableUuid="DF42663AFFCCFFDCFF5DF26CFAF0FEDD">Table 5</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFCCFFDCFF5DF79FFEECFABB" box="[189,313,1274,1346]" pageId="32" pageNumber="33" type="biology_ecology">
<paragraph id="8B8236B2FFCCFFDCFF5DF79FFEF5FAED" blockId="32.[189,288,1274,1300]" box="[189,288,1274,1300]" pageId="32" pageNumber="33">
<heading id="D0CA81DEFFCCFFDCFF5DF79FFEF5FAED" bold="true" box="[189,288,1274,1300]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B949EAA0FFCCFFDCFF5DF79FFEF5FAED" bold="true" box="[189,288,1274,1300]" pageId="32" pageNumber="33">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF64DFEECFABB" blockId="32.[189,313,1320,1346]" box="[189,313,1320,1346]" pageId="32" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3276539FFCCFFDCFF5DF60FFEABFA4A" box="[189,382,1386,1459]" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8B8236B2FFCCFFDCFF5DF60FFE83FA7D" blockId="32.[189,342,1386,1412]" box="[189,342,1386,1412]" pageId="32" pageNumber="33">
<heading id="D0CA81DEFFCCFFDCFF5DF60FFE83FA7D" bold="true" box="[189,342,1386,1412]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B949EAA0FFCCFFDCFF5DF60FFE83FA7D" bold="true" box="[189,342,1386,1412]" pageId="32" pageNumber="33">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF6FCFEABFA4A" blockId="32.[189,382,1432,1459]" box="[189,382,1432,1459]" pageId="32" pageNumber="33">
Borneo (
<collectingRegion id="49F9F850FFCCFFDCFEC3F6FDFEA4FA4A" box="[291,369,1432,1459]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFCCFFDCFF5DF6BFFE52F9DA" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8B8236B2FFCCFFDCFF5DF6BFFE88FA0D" blockId="32.[189,349,1498,1524]" box="[189,349,1498,1524]" pageId="32" pageNumber="33">
<heading id="D0CA81DEFFCCFFDCFF5DF6BFFE88FA0D" bold="true" box="[189,349,1498,1524]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B949EAA0FFCCFFDCFF5DF6BFFE88FA0D" bold="true" box="[189,349,1498,1524]" pageId="32" pageNumber="33">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF56CFE52F9DA" blockId="32.[189,391,1545,1571]" box="[189,391,1545,1571]" pageId="32" pageNumber="33">
<collectingCountry id="F32A7622FFCCFFDCFF5DF56CFEF8F9DA" box="[189,301,1545,1571]" name="Malaysia" pageId="32" pageNumber="33">Malaysia</collectingCountry>
:
<collectingRegion id="49F9F850FFCCFFDCFED8F56CFE56F9DA" box="[312,387,1545,1571]" country="Malaysia" name="Sabah" pageId="32" pageNumber="33">Sabah</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFCCFFDCFF5DF52FFEECF96A" pageId="32" pageNumber="33" type="description">
<paragraph id="8B8236B2FFCCFFDCFF5DF52FFE8CF99D" blockId="32.[189,345,1610,1636]" box="[189,345,1610,1636]" pageId="32" pageNumber="33">
<heading id="D0CA81DEFFCCFFDCFF5DF52FFE8CF99D" bold="true" box="[189,345,1610,1636]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B949EAA0FFCCFFDCFF5DF52FFE8CF99D" bold="true" box="[189,345,1610,1636]" pageId="32" pageNumber="33">Calling song</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFCCFFDCFF5DF51CFEECF96A" blockId="32.[189,313,1657,1683]" box="[189,313,1657,1683]" pageId="32" pageNumber="33">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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