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<document id="E884ACC778008928FF4EAEC0095BDCF6" ID-DOI="10.17109/AZH.63.2.143.2017" ID-ISSN="2064-2474" ID-Zenodo-Dep="12625425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638279146406" checkinUser="felipe" docAuthor="Makranczy, György" docDate="2017" docId="03B24C0FFFB0FFF1998AFF7DFE13F2D3" docLanguage="en" docName="ActaZoolAcadSciH.63.2.143-262.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 63 (2)" docSource="http://dx.doi.org/10.17109/azh.63.2.143.2017" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.6:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="6" docTitle="Anotylus tanator Makranczy, 2017, sp. n." docType="treatment" docUuid="DCBFCA4A-788E-4E9A-9E84-88485C129C00" docUuidSource="ZooBank" docVersion="2" lastPageNumber="257" masterDocId="FF8B3477FFC0FF839847FFC2FFBCF364" masterDocTitle="Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="262" masterPageNumber="143" pageNumber="255" updateTime="1719925380098" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="96EEF828BA582D02B61420638F07AE08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="52D5CF38FEC2F3E227FA8AFB2240ACAD">
<mods:title id="FFF380FA7A0FC2DF0C713C68955146D6">Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
</mods:titleInfo>
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<mods:role id="9B6DE2AF31AB260F8256381E02B9BE72">
<mods:roleTerm id="352CEE221BD5A2205051828736D8E125">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9B51792713C0677D209E15CD954B188">Makranczy, György</mods:namePart>
</mods:name>
<mods:typeOfResource id="65E537BC39443DB399D1199A0932A9C0">text</mods:typeOfResource>
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<mods:title id="920F247B12B263D42D34DEDC0BA1D559">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="054C2371A25D749469D72B9107D7C683">
<mods:date id="ABCAA56D4DCAACACEAA42485117675D6">2017</mods:date>
<mods:detail id="161D0E47974C797947D5EF3D2BDF9834" type="pubDate">
<mods:number id="211B804B16C3B4F55AAC342347785009">2017-06-19</mods:number>
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<mods:number id="0FBAFC64235021F8958D78253FDBFD31">63</mods:number>
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<mods:start id="66595C132C58DB2F425E65462842A701">143</mods:start>
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<mods:url id="FCBDC837E194809D4238840E94B84780">http://dx.doi.org/10.17109/azh.63.2.143.2017</mods:url>
</mods:location>
<mods:classification id="8EE62EE0E8925D753DF344FB8BEA2327">journal article</mods:classification>
<mods:identifier id="350EF2D38DC60F4D504C7FFAEC8F8882" type="DOI">10.17109/AZH.63.2.143.2017</mods:identifier>
<mods:identifier id="BDBA1FD47C2455EACE4610FB54640D6F" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="7A8F255427EC1EC7A20FC7A0AF0C976E" type="Zenodo-Dep">12625425</mods:identifier>
<mods:identifier id="30882A1E5510D065CA90213E6A602A11" type="ZooBank">671ADB82-5AA9-418A-BB5C-DB3B912D3BD2</mods:identifier>
</mods:mods>
<treatment id="03B24C0FFFB0FFF1998AFF7DFE13F2D3" LSID="urn:lsid:zoobank.org:act:DCBFCA4A-788E-4E9A-9E84-88485C129C00" httpUri="http://treatment.plazi.org/id/03B24C0FFFB0FFF1998AFF7DFE13F2D3" lastPageId="114" lastPageNumber="257" pageId="112" pageNumber="255">
<subSubSection id="C301AE92FFB0FFF3998AFF7DFD46F3BD" box="[461,762,191,217]" pageId="112" pageNumber="255" type="nomenclature">
<paragraph id="8BA4FD19FFB0FFF3998AFF7DFD46F3BD" blockId="112.[461,762,191,217]" box="[461,762,191,217]" pageId="112" pageNumber="255">
<heading id="D0EC4A75FFB0FFF3998AFF7DFD46F3BD" box="[461,762,191,217]" centered="true" fontSize="11" level="2" pageId="112" pageNumber="255" reason="2">
<taxonomicName id="4C1B869AFFB0FFF3998AFF7DFD13F3BD" ID-CoL="66XTT" authorityName="Makranczy" authorityYear="2017" box="[461,687,191,217]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="255" phylum="Arthropoda" rank="species" species="tanator" status="sp. nov.">
<emphasis id="B96F210BFFB0FFF3998AFF7DFD13F3BD" bold="true" box="[461,687,191,217]" pageId="112" pageNumber="255">Anotylus tanator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C9C70FFB0FFF39AF2FF7DFD46F3BD" box="[693,762,191,217]" pageId="112" pageNumber="255" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C301AE92FFB0FFF19992FF20FE13F2D3" lastPageId="114" lastPageNumber="257" pageId="112" pageNumber="255" type="description">
<paragraph id="8BA4FD19FFB0FFF39992FF20FD4EF398" blockId="112.[469,754,226,253]" box="[469,754,226,253]" pageId="112" pageNumber="255">
<heading id="D0EC4A75FFB0FFF39992FF20FD4EF398" box="[469,754,226,253]" centered="true" fontSize="11" level="3" pageId="112" pageNumber="255" reason="3">
(
<figureCitation id="1320E19CFFB0FFF3999AFF20FD80F398" box="[477,572,226,253]" captionStart="Figs 96–97" captionStartId="51.[216,261,1553,1575]" captionTargetBox="[174,1124,514,1511]" captionTargetId="figure-152@51.[173,1125,506,1517]" captionTargetPageId="51" captionText="Figs 96–97. Habitus of Anotylus species. 96 = A. rurukan sp. n., 97 = A. tanator sp. n." figureDoi="http://doi.org/10.5281/zenodo.12625481" httpUri="https://zenodo.org/record/12625481/files/figure.png" pageId="112" pageNumber="255">Figs 97</figureCitation>
,
<figureCitation id="1320E19CFFB0FFF39A0FFF20FD0DF399" box="[584,689,226,253]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">251–257</figureCitation>
,
<figureCitation id="1320E19CFFB0FFF39AFAFF20FD57F398" box="[701,747,226,252]" captionStart="Figs 265–278" captionStartId="116.[136,181,1475,1497]" captionTargetBox="[138,1088,607,1439]" captionTargetId="figure-539@116.[137,1089,605,1454]" captionTargetPageId="116" captionText="Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali" figureDoi="http://doi.org/10.5281/zenodo.12625539" httpUri="https://zenodo.org/record/12625539/files/figure.png" pageId="112" pageNumber="255">277</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BA4FD19FFB0FFF39883FED8FED6F108" blockId="112.[136,1089,281,620]" pageId="112" pageNumber="255">
<emphasis id="B96F210BFFB0FFF39883FED8FEF6F24B" box="[196,330,282,303]" italics="true" pageId="112" pageNumber="255">Typematerial</emphasis>
–
<materialsCitation id="3B73F744FFB0FFF39924FED8FEBEF20E" collectingDate="1990-08-13" collectingDateMax="1990-08-16" collectingDateMin="1990-08-13" collectionCode="SMNS" collectorName="A. Riedel" country="Indonesia" elevation="1700" latitude="-2.8128333" location="Pulu Pulu" longLatPrecision="1" longitude="119.7936" municipality="TanahToraja" pageId="112" pageNumber="255" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<typeStatus id="54A043BBFFB0FFF39924FED8FE7BF24B" box="[355,455,282,304]" pageId="112" pageNumber="255" type="holotype">Holotype</typeStatus>
(m):
<collectingCountry id="F30CBD89FFB0FFF399BBFED8FD3AF24B" box="[508,646,282,303]" name="Indonesia" pageId="112" pageNumber="255">INDONESIA</collectingCountry>
/
<collectingRegion id="49DF33FBFFB0FFF39ACAFED8FCB0F24B" box="[653,780,282,303]" country="Indonesia" name="Sulawesi Utara" pageId="112" pageNumber="255">SULAWESI</collectingRegion>
: S-,
<collectingMunicipality id="6BC06763FFB0FFF39B71FED8FC7CF24B" box="[822,960,282,304]" pageId="112" pageNumber="255">TanahToraja</collectingMunicipality>
, [Baruppu,]
<location id="8EC4ABC2FFB0FFF398CFFEF5FF4FF229" LSID="urn:lsid:plazi:treatment:03B24C0FFFB0FFF1998AFF7DFE13F2D3:8EC4ABC2FFB0FFF398CFFEF5FF4FF229" box="[136,243,311,333]" country="Indonesia" latitude="-2.8128333" longLatPrecision="1" longitude="119.7936" municipality="TanahToraja" name="Pulu Pulu" pageId="112" pageNumber="255" stateProvince="Sulawesi Utara">Pulu Pulu</location>
[
<geoCoordinate id="EE2F9BDEFFB0FFF398B8FEF5FEC6F228" box="[255,378,311,332]" degrees="02" direction="south" minutes="48.770" orientation="latitude" pageId="112" pageNumber="255" precision="1" value="-2.8128333">02°48.770’S</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFB0FFF399C2FEF4FDB3F228" box="[389,527,310,332]" degrees="119" direction="east" minutes="47.616" orientation="longitude" pageId="112" pageNumber="255" precision="1" value="119.7936">119°47.616’E</geoCoordinate>
],
<quantity id="4CE350FCFFB0FFF39A58FEF5FDD0F228" box="[543,620,311,332]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="112" pageNumber="255" unit="m" value="1700.0">
<elevation id="00361A2AFFB0FFF39A58FEF5FDD0F228" box="[543,620,311,332]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="112" pageNumber="255" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="FFA5DBD9FFB0FFF39A30FEF5FCABF229" box="[631,791,311,333]" pageId="112" pageNumber="255" value="1990-08-13" valueMax="1990-08-16" valueMin="1990-08-13">
<collectingDate id="EFE12231FFB0FFF39A30FEF5FCABF229" box="[631,791,311,333]" pageId="112" pageNumber="255" value="1990-08-13" valueMax="1990-08-16" valueMin="1990-08-13">13-16.VIII.1990</collectingDate>
</date>
, leg.
<collectorName id="26EE98CFFFB0FFF39B0AFEF5FC0EF229" box="[845,946,311,333]" pageId="112" pageNumber="255">A. Riedel</collectorName>
, sifted in for- est (
<collectionCode id="ED0A65DCFFB0FFF398F4FE96FF47F20E" box="[179,251,340,362]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="112" pageNumber="255">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B73F744FFB0FFF3994AFE96FCF0F2B8" accessionNumber="F91724" collectingDate="1990-08-13" collectingDateMax="1991-04-09" collectingDateMin="1990-08-13" collectionCode="SMNS, HNHM, MHNG, NHMW, FMNH" collectorName="A. Riedel &amp; Mamasa &amp; Penanang &amp; WofMamasa &amp; D. Agosti" country="Indonesia" elevation="2" latitude="-2.9105556" longLatPrecision="21" longitude="119.308334" pageId="112" pageNumber="255" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="54A043BBFFB0FFF3994AFE96FEC9F20D" box="[269,373,340,361]" pageId="112" pageNumber="255" type="paratype">Paratypes</typeStatus>
(25): samedataasholotype (
<quantity id="4CE350FCFFB0FFF39AE8FE96FD69F20D" box="[687,725,339,361]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="112" pageNumber="255" unit="m" value="2.0">
<elevation id="00361A2AFFB0FFF39AE8FE96FD69F20D" box="[687,725,339,361]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="112" pageNumber="255" unit="m" value="2.0">2 m</elevation>
</quantity>
, 1 f,
<collectionCode id="ED0A65DCFFB0FFF39B53FE96FCE6F20D" box="[788,858,340,361]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="112" pageNumber="255">SMNS</collectionCode>
,
<quantity id="4CE350FCFFB0FFF39B22FE96FC36F20D" box="[869,906,339,361]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">
<elevation id="00361A2AFFB0FFF39B22FE96FC36F20D" box="[869,906,339,361]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectionCode id="ED0A65DCFFB0FFF39BDFFE96FC51F20D" box="[920,1005,340,361]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="112" pageNumber="255">HNHM</collectionCode>
); Tanah Toraja, [Baruppu,] Ponding - Pulu Pulu [
<geoCoordinate id="EE2F9BDEFFB0FFF39A01FEB2FD7DF2E2" box="[582,705,368,390]" degrees="02" direction="south" minutes="49.550" orientation="latitude" pageId="112" pageNumber="255" precision="1" value="-2.8258333">02°49.550’S</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFB0FFF39A89FEB2FCEBF2E2" box="[718,855,368,390]" degrees="119" direction="east" minutes="47.335" orientation="longitude" pageId="112" pageNumber="255" precision="1" value="119.78892">119°47.335’E</geoCoordinate>
],
<quantity id="4CE350FCFFB0FFF39B2EFEB2FC49F2E2" box="[873,1013,368,390]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="112" pageNumber="255" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="00361A2AFFB0FFF39B2EFEB2FC49F2E2" box="[873,1013,368,390]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="112" pageNumber="255" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600–1800 m</elevation>
</quantity>
,
<date id="FFA5DBD9FFB0FFF39C45FEB2FF55F2C6" pageId="112" pageNumber="255" value="1990-08-13" valueMax="1990-08-17" valueMin="1990-08-13">
<collectingDate id="EFE12231FFB0FFF39C45FEB2FF55F2C6" pageId="112" pageNumber="255" value="1990-08-13" valueMax="1990-08-17" valueMin="1990-08-13">13-17. VIII.1990</collectingDate>
</date>
, leg.
<collectorName id="26EE98CFFFB0FFF39962FE4FFE31F2C7" box="[293,397,397,419]" pageId="112" pageNumber="255">A. Riedel</collectorName>
, sifted in forest (
<quantity id="4CE350FCFFB0FFF39A0AFE4FFDCFF2C7" box="[589,627,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">
<elevation id="00361A2AFFB0FFF39A0AFE4FFDCFF2C7" box="[589,627,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">1 m</elevation>
</quantity>
, 2 f,
<collectionCode id="ED0A65DCFFB0FFF39AFEFE4FFD43F2C6" box="[697,767,397,418]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="112" pageNumber="255">SMNS</collectionCode>
,
<quantity id="4CE350FCFFB0FFF39B4AFE4FFC88F2C7" box="[781,820,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">
<elevation id="00361A2AFFB0FFF39B4AFE4FFC88F2C7" box="[781,820,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectionCode id="ED0A65DCFFB0FFF39B03FE4FFC2BF2C6" box="[836,919,397,418]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="112" pageNumber="255" type="Museum">MHNG</collectionCode>
,
<quantity id="4CE350FCFFB0FFF39BE3FE4FFC70F2C7" box="[932,972,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">
<elevation id="00361A2AFFB0FFF39BE3FE4FFC70F2C7" box="[932,972,397,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectionCode id="ED0A65DCFFB0FFF39B9CFE4CFB89F2C7" box="[987,1077,398,419]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="112" pageNumber="255" type="Museum">NHMW</collectionCode>
); W-,
<collectorName id="26EE98CFFFB0FFF398F1FE68FEACF2DB" box="[182,272,426,447]" pageId="112" pageNumber="255">Mamasa</collectorName>
,
<collectorName id="26EE98CFFFB0FFF39959FE68FE3AF2DB" box="[286,390,426,447]" pageId="112" pageNumber="255">Penanang</collectorName>
,
<collectorName id="26EE98CFFFB0FFF399D2FE68FD90F2DB" box="[405,556,426,448]" pageId="112" pageNumber="255">WofMamasa</collectorName>
,
<quantity id="4CE350FCFFB0FFF39A7DFE68FD3BF2DB" box="[570,647,426,448]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="112" pageNumber="255" unit="m" value="1600.0">
<elevation id="00361A2AFFB0FFF39A7DFE68FD3BF2DB" box="[570,647,426,448]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="112" pageNumber="255" unit="m" value="1600.0">1600 m</elevation>
</quantity>
[
<geoCoordinate id="EE2F9BDEFFB0FFF39AD1FE68FCB7F2A4" box="[662,779,426,448]" degrees="02" direction="south" minutes="54" orientation="latitude" pageId="112" pageNumber="255" precision="15" seconds="38" value="-2.9105556">02°54‘38“S</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFB0FFF39B50FE6BFC25F2DB" box="[791,921,425,447]" degrees="119" direction="east" minutes="18" orientation="longitude" pageId="112" pageNumber="255" precision="15" seconds="30" value="119.308334">119°18‘30“E</geoCoordinate>
],
<date id="FFA5DBD9FFB0FFF39BEBFE68FBB2F2DB" box="[940,1038,426,447]" pageId="112" pageNumber="255" value="1991-04-09">
<collectingDate id="EFE12231FFB0FFF39BEBFE68FBB2F2DB" box="[940,1038,426,447]" pageId="112" pageNumber="255" value="1991-04-09">9.IV.1991</collectingDate>
</date>
, leg.
<collectorName id="26EE98CFFFB0FFF398CFFE05FF52F2B9" box="[136,238,455,477]" pageId="112" pageNumber="255">D. Agosti</collectorName>
(
<accessionNumber id="944860FAFFB0FFF398BAFE05FEF6F2B9" box="[253,330,455,477]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F91724" pageId="112" pageNumber="255" type="EnaNcbi">F91724</accessionNumber>
) (16), montanerainforest, leaflitter (1,
<collectionCode id="ED0A65DCFFB0FFF39AB1FE05FCF9F2B8" box="[758,837,455,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="112" pageNumber="255" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B73F744FFB0FFF39B1EFE05FF5EF29E" accessionNumber="F91722" collectionCode="MHNG" country="Indonesia" pageId="112" pageNumber="255" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
samebut (
<accessionNumber id="944860FAFFB0FFF39B8CFE05FBABF2B9" box="[971,1047,455,477]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F91722" pageId="112" pageNumber="255" type="EnaNcbi">F91722</accessionNumber>
) (2,
<collectionCode id="ED0A65DCFFB0FFF398CFFE26FF60F29E" box="[136,220,484,506]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="112" pageNumber="255" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B73F744FFB0FFF398B4FE26FEDAF108" accessionNumber="F91741" collectingDate="1991-04-11" collectingDateMax="1991-04-12" collectingDateMin="1991-04-11" collectionCode="MHNG, AMNH, ISNB, ZMUC, MNHP" collectorName="D. Agosti" country="Indonesia" elevation="1000" latitude="-2.9191668" longLatPrecision="21" longitude="119.27278" pageId="112" pageNumber="255" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
W-, Mamasa, Ulumambi, WofMamasa,
<quantity id="4CE350FCFFB0FFF39A8DFE26FCA7F29D" box="[714,795,484,505]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1000.0">
<elevation id="00361A2AFFB0FFF39A8DFE26FCA7F29D" box="[714,795,484,505]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="112" pageNumber="255" unit="m" value="1000.0">1000 m</elevation>
</quantity>
[
<geoCoordinate id="EE2F9BDEFFB0FFF39B69FE21FC1EF29D" box="[814,930,483,505]" degrees="02" direction="south" minutes="55" orientation="latitude" pageId="112" pageNumber="255" precision="15" seconds="57" value="-2.9325001">02°55‘57“S</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFB0FFF39BF4FE21FB8AF29D" box="[947,1078,483,505]" degrees="119" direction="east" minutes="15" orientation="longitude" pageId="112" pageNumber="255" precision="15" seconds="05" value="119.25139">119°15‘05“E</geoCoordinate>
],
<date id="FFA5DBD9FFB0FFF398CFFDC2FF4AF171" box="[136,246,512,533]" pageId="112" pageNumber="255" value="1991-04-11">
<collectingDate id="EFE12231FFB0FFF398CFFDC2FF4AF171" box="[136,246,512,533]" pageId="112" pageNumber="255" value="1991-04-11">11.IV.1991</collectingDate>
</date>
, leg.
<collectorName id="26EE98CFFFB0FFF39969FDC3FE2FF172" box="[302,403,512,534]" pageId="112" pageNumber="255">D. Agosti</collectorName>
(
<accessionNumber id="944860FAFFB0FFF399E6FDC2FE51F172" box="[417,493,512,534]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F91741" pageId="112" pageNumber="255" type="EnaNcbi">F91741</accessionNumber>
) (17), remnantsofrainforest / coffeeplantation, leaflitter (3,
<collectionCode id="ED0A65DCFFB0FFF39888FDDFFE9EF156" box="[207,290,541,562]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="112" pageNumber="255" type="Museum">MHNG</collectionCode>
, 1,
<collectionCode id="ED0A65DCFFB0FFF39900FDDFFE27F157" box="[327,411,541,563]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="112" pageNumber="255">AMNH</collectionCode>
); W-, Mamasa, Ulumambi, WofMamasa,
<quantity id="4CE350FCFFB0FFF39B2BFDDFFC05F157" box="[876,953,541,563]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="112" pageNumber="255" unit="m" value="1300.0">
<elevation id="00361A2AFFB0FFF39B2BFDDFFC05F157" box="[876,953,541,563]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="112" pageNumber="255" unit="m" value="1300.0">1300 m</elevation>
</quantity>
[
<geoCoordinate id="EE2F9BDEFFB0FFF39B80FDDFFB80F157" box="[967,1084,541,563]" degrees="02" direction="south" minutes="55" orientation="latitude" pageId="112" pageNumber="255" precision="15" seconds="09" value="-2.9191668">02°55‘09“S</geoCoordinate>
,
<geoCoordinate id="EE2F9BDEFFB0FFF398CFFDFBFEB6F12B" box="[136,266,569,591]" degrees="119" direction="east" minutes="16" orientation="longitude" pageId="112" pageNumber="255" precision="15" seconds="22" value="119.27278">119°16‘22“E</geoCoordinate>
],
<date id="FFA5DBD9FFB0FFF3995CFDF8FE35F12B" box="[283,393,570,591]" pageId="112" pageNumber="255" value="1991-04-12">
<collectingDate id="EFE12231FFB0FFF3995CFDF8FE35F12B" box="[283,393,570,591]" pageId="112" pageNumber="255" value="1991-04-12">12.IV.1991</collectingDate>
</date>
, leg.
<collectorName id="26EE98CFFFB0FFF399F8FDF8FD9EF134" box="[447,546,570,592]" pageId="112" pageNumber="255">D. Agosti</collectorName>
(18), montane forest, leaf litter (6,
<collectionCode id="ED0A65DCFFB0FFF39BC9FDF8FC5DF12B" box="[910,993,570,591]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="112" pageNumber="255" type="Museum">MHNG</collectionCode>
, 1,
<collectionCode id="ED0A65DCFFB0FFF39C44FDF8FB81F12B" box="[1027,1085,570,591]" pageId="112" pageNumber="255">ISNB</collectionCode>
, 1,
<collectionCode id="ED0A65DCFFB0FFF398E7FD95FF51F108" box="[160,237,599,620]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="112" pageNumber="255">ZMUC</collectionCode>
, 1,
<collectionCode id="ED0A65DCFFB0FFF39949FD95FEE3F108" box="[270,351,599,620]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="112" pageNumber="255">MNHP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA4FD19FFB0FFF39883FD53FDB8F55B" blockId="112.[136,1088,656,1600]" pageId="112" pageNumber="255">
<emphasis id="B96F210BFFB0FFF39883FD53FE8AF1C2" box="[196,310,657,678]" italics="true" pageId="112" pageNumber="255">Description</emphasis>
– Measurements (in mm, n=10): HW = 0.70 (0.64–0.74); TW = 0.69 (0.62– 0.72); PW = 1.02 (0.93–1.08); EW = 1.10 (1.05–1.16); AW = 1.18 (1.08–1.26); HL = 0.55 (0.50– 0.57); EL = 0.14 (0.13–0.15); TL = 0.21 (0.19–0.23); PL = 0.60 (0.54–0.65); SC = 0.65 (0.61–0.69); FB = 1.88 (1.75–1.95); BL = 3.62 (3.35–3.90). Habitus as in
<figureCitation id="1320E19CFFB0FFF39A98FD25FC94F198" box="[735,808,743,765]" captionStart="Figs 96–97" captionStartId="51.[216,261,1553,1575]" captionTargetBox="[174,1124,514,1511]" captionTargetId="figure-152@51.[173,1125,506,1517]" captionTargetPageId="51" captionText="Figs 96–97. Habitus of Anotylus species. 96 = A. rurukan sp. n., 97 = A. tanator sp. n." figureDoi="http://doi.org/10.5281/zenodo.12625481" httpUri="https://zenodo.org/record/12625481/files/figure.png" pageId="112" pageNumber="255">Fig. 97</figureCitation>
. Forebody weakly shining becauseofsculpture; abdomenwithsuchextremelydensecoriaceousmicrosculpturethat noteventracesofpunctationobservable, opaque. Forebody, legsandabdomenreddish mediumtodarkbrown, exceptexplanatepartofpronotumandclypealarea (bothfor transparency), these, mouthpartsandantennaereddishmediumbrown. Headwithathin rimonanteriormargin, truncateinmiddle, laterallyevenlyarchedtowardseyes, supraan- tennalprominencesasmoderatelyelevatedobliqueridges, discslightlyimpressednear eyes. Epistomalsutureformingsubcircularimpression (almostimpunctateinside), upper surfacewithouttactilesetae, eyesmoderatelysmall, veryslightlybulgingfromsidelineof head; indorsalviewtemples (1.5× lengthofeye) alatiform, laterallyimperfectlyrounded (withoutbeingangled) andposteriorlyconstrictedtoformdistinctneck, notseparated dorsallybyoccipitalfurrow, onlybyneckbeingmicrosculpturedratherthanpunctured. Antennomere 1 club-shaped, finelysculptured, segment 2 elongateoval, segment 3 clubshaped, segment 4 smallandisodiametrical, segments 5–11 withbasaldishes, articles 6–10 transverse (about 1.5× broaderthanlong), segments 9–10 bulkierthanpreceding. Head andpronotumwithmostlylacunosesculpture. Pronotumwithlateralportionexplanate andslightlyup-turned, edgealatiformwithsparseshortsetae; lateralmarginmoreorless evenlyarchedwithanumberofsmall, irregularlydistributeddenticles, posteriormargin broadlyincisedbeforecorners; discmediallywithtwoparallellongitudinalridges, almost confluentposteriorly, impressedalongsidethem, laterallyimpressedaroundmiddleof sides. Elytrawithmembranousposteriormarginextendingfromsuturalcornerto 3/4 of hindmargin. Elytralepipleuralridgepresentwithup-turnedlateralalatiformportions, disclongitudinallyimpressedalongside; shoulderswelldeveloped, surfacefoveolate, in betweenpuncturestorulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesrath- erarcuate, secondsegmentwithparatergitesstronglybroadeningposteriorly (abdomen appearingconstrictedatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad, posterioredgeoftergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasin
<figureCitation id="1320E19CFFB0FFF398CFF9CFFF60F547" box="[136,220,1549,1571]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">Fig. 251</figureCitation>
, sterniteVIII (
<figureCitation id="1320E19CFFB0FFF39933F9CFFE77F547" box="[372,459,1549,1571]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">Fig. 252</figureCitation>
), tergitesIX (
<figureCitation id="1320E19CFFB0FFF39A1FF9CFFD13F547" box="[600,687,1549,1571]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">Fig. 253</figureCitation>
), X (
<figureCitation id="1320E19CFFB0FFF39A98F9CFFC8FF547" box="[735,819,1549,1571]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">Fig. 254</figureCitation>
)
<figureCitation id="1320E19CFFB0FFF39B06F9CCFF62F524" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="112" pageNumber="255">andaedeagusasinFigs 255–257</figureCitation>
,
<figureCitation id="1320E19CFFB0FFF398AFF9E8FDBCF55B" box="[232,512,1578,1600]" captionStart="Figs 265–278" captionStartId="116.[136,181,1475,1497]" captionTargetBox="[138,1088,607,1439]" captionTargetId="figure-539@116.[137,1089,605,1454]" captionTargetPageId="116" captionText="Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali" figureDoi="http://doi.org/10.5281/zenodo.12625539" httpUri="https://zenodo.org/record/12625539/files/figure.png" pageId="112" pageNumber="255">spermathecaasinFig. 277</figureCitation>
.
</paragraph>
<caption id="DF64AD91FFB1FFF298EAFA10FBDAF546" ID-DOI="http://doi.org/10.5281/zenodo.12625533" ID-Zenodo-Dep="12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="113" pageNumber="256" startId="113.[173,218,1490,1512]" targetBox="[187,1114,190,1463]" targetPageId="113">
<paragraph id="8BA4FD19FFB1FFF298EAFA10FBDAF546" blockId="113.[173,1126,1490,1599]" pageId="113" pageNumber="256">
<emphasis id="B96F210BFFB1FFF298EAFA10FE87F68C" bold="true" box="[173,315,1490,1512]" pageId="113" pageNumber="256">Figs 251–257.</emphasis>
<taxonomicName id="4C1B869AFFB1FFF29905FA11FE55F68C" authorityName="Makranczy" authorityYear="2017" box="[322,489,1491,1512]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="256" phylum="Arthropoda" rank="species" species="tanator" status="sp. nov.">
<emphasis id="B96F210BFFB1FFF29905FA11FE55F68C" box="[322,489,1491,1512]" italics="true" pageId="113" pageNumber="256">Anotylus tanator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C9C70FFB1FFF299A8FA11FD9BF68C" box="[495,551,1491,1512]" pageId="113" pageNumber="256" rank="species">sp. n.</taxonomicNameLabel>
, male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257,
</paragraph>
</caption>
<paragraph id="8BA4FD19FFB1FFF299D4F9EBFC3CF55B" blockId="113.[173,1126,1490,1599]" box="[403,896,1577,1599]" pageId="113" pageNumber="256">
<quantity id="4CE350FCFFB1FFF299D4F9EBFE51F55B" box="[403,493,1577,1599]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="113" pageNumber="256" unit="mm" value="0.11">0.11 mm</quantity>
for
<figureCitation id="1320E19CFFB1FFF29A5FF9EBFD1CF55B" box="[536,672,1577,1599]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="113" pageNumber="256">Figs 253–254</figureCitation>
,
<quantity id="4CE350FCFFB1FFF29AEDF9EBFCB8F55B" box="[682,772,1577,1599]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="113" pageNumber="256" unit="mm" value="0.13">0.13 mm</quantity>
for
<figureCitation id="1320E19CFFB1FFF29B68F9EBFC3CF55B" box="[815,896,1577,1599]" captionStart="Figs 251–257" captionStartId="113.[173,218,1490,1512]" captionTargetBox="[187,1114,190,1463]" captionTargetId="figure-13@113.[185,1114,189,1464]" captionTargetPageId="113" captionText="Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257," figureDoi="http://doi.org/10.5281/zenodo.12625533" httpUri="https://zenodo.org/record/12625533/files/figure.png" pageId="113" pageNumber="256">Fig. 252</figureCitation>
</paragraph>
<paragraph id="8BA4FD19FFB2FFF19883FF7CFD5CF395" blockId="114.[136,1088,190,241]" pageId="114" pageNumber="257">
<emphasis id="B96F210BFFB2FFF19883FF7CFE91F3B7" box="[196,301,190,211]" italics="true" pageId="114" pageNumber="257">Etymology</emphasis>
– The specific epithet is composed of the beginnings of the two words of thetypelocalityarea, TanahToraja (nouninapposition).
</paragraph>
<paragraph id="8BA4FD19FFB2FFF19883FED4FDB3F210" blockId="114.[136,1088,278,439]" pageId="114" pageNumber="257">
<emphasis id="B96F210BFFB2FFF19883FED4FEEEF254" box="[196,338,278,304]" italics="true" pageId="114" pageNumber="257">Distribution</emphasis>
– Accordingtoadministrativeareasthespeciesisknown fromanareapartlybelongingtoWSulawesi (Mamasaregency), partlyS
<collectingRegion id="49DF33FBFFB2FFF198CFFE9BFF49F217" box="[136,245,345,372]" country="Indonesia" name="Sulawesi Utara" pageId="114" pageNumber="257">Sulawesi</collectingRegion>
(TanaTorajaregency).
</paragraph>
<paragraph id="8BA4FD19FFB2FFF19883FEB9FE13F2D3" blockId="114.[136,1088,278,439]" pageId="114" pageNumber="257">
<emphasis id="B96F210BFFB2FFF19883FEB9FE9AF2F1" box="[196,294,379,405]" italics="true" pageId="114" pageNumber="257">Remarks</emphasis>
– Itisapparentlyrelatedto
<taxonomicName id="4C1B869AFFB2FFF19ACEFEB9FCB6F2F1" authorityName=", Gunung Mahawu" authorityYear="1200" box="[649,778,379,405]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="257" phylum="Arthropoda" rank="species" species="rurukan">
<emphasis id="B96F210BFFB2FFF19ACEFEB9FCB6F2F1" box="[649,778,379,405]" italics="true" pageId="114" pageNumber="257">A. rurukan</emphasis>
</taxonomicName>
(alsofromSulawesi, but fromthenorthernpart).
</paragraph>
</subSubSection>
</treatment>
</document>
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