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<mods:title id="98C93A5C68F46BCD11D4710E5CCD121C">Systematic studies of the genus Aegialomys Weksler, Percequillo and Voss, 2006 (Rodentia: Cricetidae: Sigmodontinae): Annotated catalogue of the types of the species-group taxa</mods:title>
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<mods:namePart id="297212C985D66148E6380321E6DFF7D4">Prado, Joyce Rodrigues Do</mods:namePart>
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<treatment id="03B487F6FF80FFDAFF56FA4BFF68FA69" ID-DOI="http://doi.org/10.5281/zenodo.5624973" ID-GBIF-Taxon="123514544" ID-Zenodo-Dep="5624973" LSID="urn:lsid:plazi:treatment:03B487F6FF80FFDAFF56FA4BFF68FA69" httpUri="http://treatment.plazi.org/id/03B487F6FF80FFDAFF56FA4BFF68FA69" lastPageId="17" lastPageNumber="494" pageId="15" pageNumber="492">
<subSubSection id="C307656BFF80FFC4FF56FA4BFD33FA1D" box="[151,644,1486,1512]" pageId="15" pageNumber="492" type="nomenclature">
<paragraph id="8BA236E0FF80FFC4FF56FA4BFD33FA1D" blockId="15.[151,644,1486,1512]" box="[151,644,1486,1512]" pageId="15" pageNumber="492">
<heading id="D0EA818CFF80FFC4FF56FA4BFD33FA1D" bold="true" box="[151,644,1486,1512]" fontSize="11" level="1" pageId="15" pageNumber="492" reason="1">
<taxonomicName id="4C1D4D63FF80FFC4FF56FA4BFD33FA1D" authority="Osgood, 1944" authorityName="Osgood" authorityYear="1944" box="[151,644,1486,1512]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="492" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="ica">
<emphasis id="B969EAF2FF80FFC4FF56FA4BFD33FA1D" bold="true" box="[151,644,1486,1512]" pageId="15" pageNumber="492">
<emphasis id="B969EAF2FF80FFC4FF56FA4BFE63FA1D" bold="true" box="[151,468,1486,1512]" italics="true" pageId="15" pageNumber="492">Oryzomys xanthaeolus ica</emphasis>
<bibRefCitation id="EF8C4B11FF80FFC4FE1DFA4BFD33FA1D" author="Osgood" box="[476,644,1486,1512]" pageId="15" pageNumber="492" refString="Osgood, W. H. (1944) Nine new South American rodents. Field Museum of Natural History Publications Zoological Series, 29, 191 - 204." type="journal article" year="1944">Osgood, 1944</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C307656BFF80FFDAFF56F990FF5EFEE9" lastPageId="17" lastPageNumber="494" pageId="15" pageNumber="492" type="materials_examined">
<paragraph id="8BA236E0FF80FFC4FF56F990FED3F9A7" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
<emphasis id="B969EAF2FF80FFC4FF56F990FEBCF9DB" bold="true" box="[151,267,1557,1582]" pageId="15" pageNumber="492">Holotype.</emphasis>
FMNH 53157, a female specimen, deposited at The Field Museum, collected by C. C. Sanborn, in
<date id="FFA31020FF80FFC4FF56F9BFFEB5F9A7" box="[151,258,1594,1618]" pageId="15" pageNumber="492" value="1942-01-17">17.I.1942</date>
(
<figureCitation id="13262A65FF80FFC4FED0F9BCFEE1F9A7" box="[273,342,1593,1618]" captionStart="FIGURE 7" captionStartId="16.[151,250,1950,1972]" captionTargetBox="[209,1377,193,1927]" captionTargetId="figure@16.[209,1377,193,1929]" captionTargetPageId="16" captionText="FIGURE 7. Dorsal, ventral and lateral views of the skull, and lateral view of the mandible of the specimen FMNH 53157, holotype of Oryzomys xanthaeolus ica Osgood, 1944 (ONL: 32.88 mm)." httpUri="https://zenodo.org/record/261826/files/figure.png" pageId="15" pageNumber="492">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BA236E0FF80FFC4FF06F9D8FEA8F96C" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
<emphasis id="B969EAF2FF80FFC4FF06F9D8FECCF983" bold="true" box="[199,379,1629,1654]" pageId="15" pageNumber="492">
<typeStatus id="54A68842FF80FFC4FF06F9D8FEB5F983" box="[199,258,1629,1654]" pageId="15" pageNumber="492">Type</typeStatus>
condition
</emphasis>
. Specimen preserved in skin and skull; both of them are in excellent condition, without any broken part.
</paragraph>
<paragraph id="8BA236E0FF80FFC4FF06F920FDBDF917" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
<emphasis id="B969EAF2FF80FFC4FF06F920FED9F94B" bold="true" box="[199,366,1701,1726]" pageId="15" pageNumber="492">
<typeStatus id="54A68842FF80FFC4FF06F920FEB5F94B" box="[199,258,1701,1726]" pageId="15" pageNumber="492">Type</typeStatus>
material
</emphasis>
. The
<typeStatus id="54A68842FF80FFC4FE71F923FE57F94B" box="[432,480,1702,1726]" pageId="15" pageNumber="492">type</typeStatus>
material consists of two specimens: the
<typeStatus id="54A68842FF80FFC4FC6AF920FBA7F94B" box="[939,1040,1701,1726]" pageId="15" pageNumber="492" type="holotype">holotype</typeStatus>
, described above, and the
<typeStatus id="54A68842FF80FFC4FAFAF923FA2BF94B" box="[1339,1436,1702,1726]" pageId="15" pageNumber="492" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED0CAE25FF80FFC4FF5FF94FFF46F917" box="[158,241,1738,1762]" pageId="15" pageNumber="492">FMHN</collectionCode>
53158), an adult female.
</paragraph>
<paragraph id="8BA236E0FF80FFC4FF06F968FB07F8DF" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
<materialsCitation id="3B753CBDFF80FFC4FF06F968FB0AF8F0" ID-GBIF-Occurrence="1291011492" box="[199,1213,1773,1798]" country="Peru" location="Hacienda San Jacinto" municipality="Province" pageId="15" pageNumber="492" specimenCount="1" stateProvince="Ica" typeStatus="holotype">
<emphasis id="B969EAF2FF80FFC4FF06F968FED1F8F3" bold="true" box="[199,358,1773,1798]" pageId="15" pageNumber="492">
<typeStatus id="54A68842FF80FFC4FF06F968FEB5F8F3" box="[199,258,1773,1798]" pageId="15" pageNumber="492">Type</typeStatus>
locality
</emphasis>
.
<location id="8EC2603BFF80FFC4FEBCF968FD35F8F3" LSID="urn:lsid:plazi:treatment:03B487F6FF80FFDAFF56FA4BFF68FA69:8EC2603BFF80FFC4FEBCF968FD35F8F3" box="[381,642,1773,1798]" country="Peru" municipality="Province" name="Hacienda San Jacinto" pageId="15" pageNumber="492" stateProvince="Ica">Hacienda San Jacinto</location>
, near
<collectingRegion id="49D9F802FF80FFC4FD17F96BFD4CF8F0" box="[726,763,1774,1797]" country="Peru" name="Ica" pageId="15" pageNumber="492">Ica</collectingRegion>
,
<collectingMunicipality id="6BC6AC9AFF80FFC4FCCEF968FCC4F8F3" box="[783,883,1773,1798]" pageId="15" pageNumber="492">Province</collectingMunicipality>
of
<collectingRegion id="49D9F802FF80FFC4FC6BF96BFC78F8F0" box="[938,975,1774,1797]" country="Peru" name="Ica" pageId="15" pageNumber="492">Ica</collectingRegion>
, southwestern
<collectingCountry id="F30A7670FF80FFC4FB47F96BFB0AF8F0" box="[1158,1213,1774,1797]" name="Peru" pageId="15" pageNumber="492">Peru</collectingCountry>
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. The geographical coordinates of this locality are 14°09S, 75°45W. The type locality is represented in figure 3.
</paragraph>
<paragraph id="8BA236E0FF80FFC4FF06F8B0FADEF887" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
<emphasis id="B969EAF2FF80FFC4FF06F8B0FE09F8BB" bold="true" box="[199,446,1845,1870]" pageId="15" pageNumber="492">Original description</emphasis>
. Osgood described
<taxonomicName id="4C1D4D63FF80FFC4FD6DF8B3FC69F8B8" box="[684,990,1845,1869]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="492" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="ica">
<emphasis id="B969EAF2FF80FFC4FD6DF8B3FC69F8B8" box="[684,990,1845,1869]" italics="true" pageId="15" pageNumber="492">Oryzomys xanthaeolus ica</emphasis>
</taxonomicName>
in the Zoological Series of the Field Museum of Natural History, vol. 29, n.13, 1944:192193, providing the following simplified characterization:
</paragraph>
<paragraph id="8BA236E0FF80FFC4FF06F8F8FE4AF84F" blockId="15.[151,1436,1557,1978]" pageId="15" pageNumber="492">
“Similar in color and general characters to
<taxonomicName id="4C1D4D63FF80FFC4FD7EF8FBFC3DF860" box="[703,906,1917,1941]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="492" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="xanthaeolus">
<emphasis id="B969EAF2FF80FFC4FD7EF8FBFC3DF860" box="[703,906,1917,1941]" italics="true" pageId="15" pageNumber="492">O. x. xanthaeolus</emphasis>
</taxonomicName>
but somewhat larger and having a skull with definitely larger audital bullae.”
</paragraph>
<caption id="DF626668FF9FFFDBFF56F81BFCC0F826" httpUri="https://zenodo.org/record/261826/files/figure.png" pageId="16" pageNumber="493" targetBox="[209,1377,193,1927]" targetPageId="16">
<paragraph id="8BA236E0FF9FFFDBFF56F81BFCC0F826" blockId="16.[151,1436,1950,2003]" pageId="16" pageNumber="493">
<emphasis id="B969EAF2FF9FFFDBFF56F81BFEA2F841" bold="true" box="[151,277,1950,1972]" pageId="16" pageNumber="493">FIGURE 7.</emphasis>
Dorsal, ventral and lateral views of the skull, and lateral view of the mandible of the specimen FMNH 53157, holotype of
<taxonomicName id="4C1D4D63FF9FFFDBFED0F83BFD06F826" authority="Osgood, 1944" authorityName="Osgood" authorityYear="1944" box="[273,689,1981,2003]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="493" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="ica">
<emphasis id="B969EAF2FF9FFFDBFED0F83BFDABF826" box="[273,540,1981,2003]" italics="true" pageId="16" pageNumber="493">Oryzomys xanthaeolus ica</emphasis>
Osgood, 1944
</taxonomicName>
(ONL: 32.88 mm).
</paragraph>
</caption>
<paragraph id="8BA236E0FF9EFFDAFF06FF12FF5EFEE9" blockId="17.[151,1437,151,1436]" pageId="17" pageNumber="494">
<emphasis id="B969EAF2FF9EFFDAFF06FF12FE0DFF45" bold="true" box="[199,442,151,176]" pageId="17" pageNumber="494">Original dimensions</emphasis>
. Total length:
<quantity id="4CE59B05FF9EFFDAFD9FFF12FD08FF5A" box="[606,703,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="17" pageNumber="494" unit="mm" value="325.0">325 mm</quantity>
; tail:
<quantity id="4CE59B05FF9EFFDAFD3EFF12FCE8FF5A" box="[767,863,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.61" pageId="17" pageNumber="494" unit="mm" value="161.0">161 mm</quantity>
; hindfoot:
<quantity id="4CE59B05FF9EFFDAFC18FF1DFB9BFF5A" box="[985,1068,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="17" pageNumber="494" unit="mm" value="33.0">33 mm</quantity>
. Skull-greatest length:
<quantity id="4CE59B05FF9EFFDAFAF1FF1DFA2FFF5A" box="[1328,1432,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.4799999999999995" pageId="17" pageNumber="494" unit="mm" value="44.8">44.8 mm</quantity>
; zygomatic breadth:
<quantity id="4CE59B05FF9EFFDAFEB8FF39FE55FF21" box="[377,482,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="17" pageNumber="494" unit="mm" value="19.3">19.3 mm</quantity>
; breadth of braincase:
<quantity id="4CE59B05FF9EFFDAFD27FF39FCF8FF21" box="[742,847,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="17" pageNumber="494" unit="mm" value="13.5">13.5 mm</quantity>
; interorbital space:
<quantity id="4CE59B05FF9EFFDAFBF1FF39FB3DFF21" box="[1072,1162,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="17" pageNumber="494" unit="mm" value="6.1">6.1 mm</quantity>
; nasals: 14.3 x
<quantity id="4CE59B05FF9EFFDAFAFCFF39FA2FFF21" box="[1341,1432,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="17" pageNumber="494" unit="mm" value="4.5">4.5 mm</quantity>
; diastema,
<quantity id="4CE59B05FF9EFFDAFECBFF5AFED2FF02" box="[266,357,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="17" pageNumber="494" unit="mm" value="8.5">8.5 mm</quantity>
; palatine foramina:
<quantity id="4CE59B05FF9EFFDAFD8BFF65FD12FF02" box="[586,677,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="17" pageNumber="494" unit="mm" value="7.3">7.3 mm</quantity>
; upper toothrow,
<quantity id="4CE59B05FF9EFFDAFCAFFF5AFC7EFF02" box="[878,969,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="494" unit="mm" value="5.5">5.5 mm</quantity>
. Other craniodental measurements are in table 2.
</paragraph>
</subSubSection>
<subSubSection id="C307656BFF9EFFDAFF06FEA2FF68FA69" pageId="17" pageNumber="494" type="description">
<paragraph id="8BA236E0FF9EFFDAFF06FEA2FAABFE00" blockId="17.[151,1437,151,1436]" pageId="17" pageNumber="494">
<emphasis id="B969EAF2FF9EFFDAFF06FEA2FDB3FEB5" bold="true" box="[199,516,295,320]" pageId="17" pageNumber="494">Morphological description</emphasis>
. Dorsal pelage soft, lax, dense and long; yellowish to ochraceous wool and cover hairs, intensely grizzled with brown guard hairs, resulting in dorsal color ochraceous cream grizzled with brown; venter grayish cream; tail slightly shorter than head and body length, densely covered with hairs, weakly bicolor, with large scales with approximately 18 scales/cm and without terminal tuft; pes covered by very long and white hairs; ungual tufts dense and long, white; interdigital and plantar pads developed, thenar and hypothenar fleshy; pinnae with few external hairs and with a creamy and yellowish coloration and inner hairs light yellow.
</paragraph>
<paragraph id="8BA236E0FF9EFFDAFF06FE7AFBFBFC32" blockId="17.[151,1437,151,1436]" pageId="17" pageNumber="494">
Skull very large and robust, with moderately long and broad rostrum (approximately 36% of skull length [LN/ ONL ratio]); zygomatic plate moderately projected anteriorly, not reaching beyond nasolacrimal capsule; rostral fossa very deep; moderately projected plate and very deep fossa, configuring a moderately deep and very wide zygomatic notch; interorbital region strongly diverging posteriorly, with strongly beaded supraorbital margins; zygomatic arches strongly divergent posteriorly, wider near the squamosal root; braincase elongated, with temporal margins squared, without crests; interparietal short and wide; fronto-squamosal suture contiguous with frontoparietal suture; alisphenoid strut absent; anastomotic channel present, configuring pattern 3 of carotid circulation (
<bibRefCitation id="EF8C4B11FF9EFFDAFF5FFD79FEA9FCE1" author="Voss" box="[158,286,764,788]" pageId="17" pageNumber="494" refString="Voss, R. S. (1988) Systematics and ecology of ichthyomyine rodents (Muroidea): patterns of morphological evolution in a small adaptive radiation. Bulletin American Museum of Natural History, 188, 259 - 493." type="journal article" year="1988">Voss 1988</bibRefCitation>
); parietals expanding over surface side of skull; postglenoid foramen apparently small; incisive foramen long, wider medially, with anterior and posterior margins slightly acute, posterior margin reaching anterocone of M1; posterior margin of zygomatic plate situated anterior to the alveolus of M1; palate intermediate, with deep posterolateral palatal pits recessed in deep and large palatine depressions; palate apparently without palatal excrescencies; mesopterygoid fossa perforated by very large sphenopalatine vacuities on the presphenoid and basisphenoid; auditory bulla large, with very short and narrow stapedial process.
</paragraph>
<paragraph id="8BA236E0FF9EFFDAFF06FC50FDB9FB1D" blockId="17.[151,1437,151,1436]" pageId="17" pageNumber="494">Upper incisors opisthodont; upper molars with labial and lingual cusps arranged in opposite pairs; M1 with anteroloph short and narrow, almost entirely fused to anterocone; anterocone apparently with shallow anteromedian flexus; paracone connected medially to protocone; median mure more labially positioned, defining a long and deep hypoflexus; paracone connected labially to mesoloph, forming a small, round and labially positioned mesofosset; mesoloph moderately long, connected with median mure medially; mesoloph separated from metacone by metaflexus; M2 similar to M1, but with median mure more medially positioned; paracone connected medially to protocone; mesoloph long, connected labially to paracone, forming a small, rounded and slightly oblique mesofosset; M3 small, triangular.</paragraph>
<paragraph id="8BA236E0FF9EFFDAFF06FB71FF68FA69" blockId="17.[151,1437,151,1436]" pageId="17" pageNumber="494">
<emphasis id="B969EAF2FF9EFFDAFF06FB71FED1FAF8" bold="true" box="[199,358,1268,1293]" pageId="17" pageNumber="494">Observations</emphasis>
. According to
<bibRefCitation id="EF8C4B11FF9EFFDAFDD0FB71FD0DFAF9" author="Osgood" box="[529,698,1268,1293]" pageId="17" pageNumber="494" refString="Osgood, W. H. (1944) Nine new South American rodents. Field Museum of Natural History Publications Zoological Series, 29, 191 - 204." type="journal article" year="1944">Osgood (1944)</bibRefCitation>
specimens of
<taxonomicName id="4C1D4D63FF9EFFDAFC9FFB71FB83FAF9" box="[862,1076,1268,1292]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="494" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="ica">
<emphasis id="B969EAF2FF9EFFDAFC9FFB71FB83FAF9" box="[862,1076,1268,1292]" italics="true" pageId="17" pageNumber="494">O. xanthaeolus ica</emphasis>
</taxonomicName>
are in overall larger and have a large auditory bulla when compared with the
<taxonomicName id="4C1D4D63FF9EFFDAFD5AFA9DFCF2FAC5" box="[667,837,1304,1328]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="494" phylum="Chordata" rank="species" species="xanthaeolus">
<emphasis id="B969EAF2FF9EFFDAFD5AFA9DFCF2FAC5" box="[667,837,1304,1328]" italics="true" pageId="17" pageNumber="494">O. xanthaeolus</emphasis>
</taxonomicName>
from nearby localities. After its original description,
<taxonomicName id="4C1D4D63FF9EFFDAFF56FAB9FEDBFAA1" box="[151,364,1340,1364]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="494" phylum="Chordata" rank="subSpecies" species="xanthaeolus" subSpecies="ica">
<emphasis id="B969EAF2FF9EFFDAFF56FAB9FEDBFAA1" box="[151,364,1340,1364]" italics="true" pageId="17" pageNumber="494">O. xanthaeolus ica</emphasis>
</taxonomicName>
remained as a subspecies of
<taxonomicName id="4C1D4D63FF9EFFDAFD79FAB9FCD4FAA1" box="[696,867,1340,1364]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="494" phylum="Chordata" rank="species" species="xanthaeolus">
<emphasis id="B969EAF2FF9EFFDAFD79FAB9FCD4FAA1" box="[696,867,1340,1364]" italics="true" pageId="17" pageNumber="494">O. xanthaeolus</emphasis>
</taxonomicName>
until the early 1990s, when
<bibRefCitation id="EF8C4B11FF9EFFDAFB71FAB8FF56FA8D" author="Musser" pageId="17" pageNumber="494" refString="Musser, G. G. &amp; Carleton, M. D. (1993) Family Muridae. In: Wilson, D. E. &amp; Reeder, D. M. (Eds), Mammal species of the world: a taxonomic and geographic reference. 2 nd ed. Smithsonian Institution Press, Washington, DC, pp. 501 - 753." type="book chapter" year="1993">Musser and Carleton (1993)</bibRefCitation>
synonymized it with
<taxonomicName id="4C1D4D63FF9EFFDAFE1DFAE5FD30FA8D" box="[476,647,1376,1400]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="494" phylum="Chordata" rank="species" species="xanthaeolus">
<emphasis id="B969EAF2FF9EFFDAFE1DFAE5FD30FA8D" box="[476,647,1376,1400]" italics="true" pageId="17" pageNumber="494">O. xanthaeolus</emphasis>
</taxonomicName>
in the catalogue “Mammal Species of the World” (Wilson &amp; Reeder 1993).
</paragraph>
</subSubSection>
</treatment>
</document>