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<document id="DD84B5E6BE75D48B49D4D77C5B3F5613" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. &amp; Mathewes, Rolf W." docDate="2021" docId="03B487C2005CFFC3FF5BFF72FC3014A8" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Okanopteryx jeppesenorum Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes 2021, new species" docType="treatment" docVersion="7" lastPageNumber="85" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="79" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:title id="B5ED464A264A5EC71CEAC3B49E29E054">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="1569B3F5891EEAE562D596252816FA14">Archibald, Bruce</mods:namePart>
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<mods:namePart id="87913D0E62FE528AF484E4254B030ECA">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="E055D5A7909E35642A3852D56EF29882" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="5428A0E464FBC34E8665B9499F47E923">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="FFE049A88098D173D640C93F4BDDD986" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A94CA43FE05E9905D3E3D8D58F2BCEBC">Bybee, Seth M.</mods:namePart>
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<mods:nameIdentifier id="AB85107A1FD59F17E0B7ECC6B908B229" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5E96B93D6B9813544C10BF57425E4542">Mathewes, Rolf W.</mods:namePart>
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<mods:nameIdentifier id="45CF3734CEE57A877B82ABD80704F27C" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
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<mods:title id="009F633CF259D378D00A59540A87B1E0">Zootaxa</mods:title>
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<mods:date id="586AEF1875337DC9B99EE3468DE24835">2021</mods:date>
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<mods:number id="1AC2B5C2B320F0F14D377B71B45B486E">4934</mods:number>
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<treatment id="03B487C2005CFFC3FF5BFF72FC3014A8" ID-DOI="http://doi.org/10.5281/zenodo.4672752" ID-GBIF-Taxon="180881797" ID-Zenodo-Dep="4672752" LSID="urn:lsid:plazi:treatment:03B487C2005CFFC3FF5BFF72FC3014A8" httpUri="http://treatment.plazi.org/id/03B487C2005CFFC3FF5BFF72FC3014A8" lastPageId="84" lastPageNumber="85" pageId="78" pageNumber="79">
<subSubSection id="C307655F005CFFD9FF5BFF72FC0B12A4" box="[151,933,151,178]" pageId="78" pageNumber="79" type="nomenclature">
<paragraph id="8BA236D4005CFFD9FF5BFF72FC0B12A4" blockId="78.[151,933,151,213]" box="[151,933,151,178]" pageId="78" pageNumber="79">
<heading id="D0EA81B8005CFFD9FF5BFF72FC0B12A4" bold="true" box="[151,933,151,178]" fontSize="11" level="2" pageId="78" pageNumber="79" reason="4">
<emphasis id="B969EAC6005CFFD9FF5BFF72FC0B12A4" bold="true" box="[151,933,151,178]" pageId="78" pageNumber="79">
<taxonomicName id="4C1D4D57005CFFD9FF5BFF72FCA712A4" authority="Archibald &amp; Cannings" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[151,777,151,178]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="jeppesenorum" status="sp. nov.">
<emphasis id="B969EAC6005CFFD9FF5BFF72FE4A12A4" bold="true" box="[151,484,151,178]" italics="true" pageId="78" pageNumber="79">Okanopteryx jeppesenorum</emphasis>
Archibald &amp; Cannings
</taxonomicName>
,
<taxonomicNameLabel id="A25A57BD005CFFD9FCDAFF72FC0B12A4" box="[790,933,151,177]" pageId="78" pageNumber="79" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C307655F005CFFD9FF5BFF5EFE8D12C0" box="[151,291,187,213]" pageId="78" pageNumber="79" type="description">
<paragraph id="8BA236D4005CFFD9FF5BFF5EFE8D12C0" blockId="78.[151,933,151,213]" box="[151,291,187,213]" pageId="78" pageNumber="79">
<figureCitation id="13262A51005CFFD9FF5BFF5EFEB112C0" box="[151,287,187,213]" captionStart-0="FIGURE 44" captionStart-1="FIGURE 45" captionStart-2="FIGURE 46" captionStart-3="FIGURE 47" captionStart-4="FIGURE 48" captionStartId-0="79.[151,250,1955,1979]" captionStartId-1="80.[151,250,1985,2009]" captionStartId-2="81.[151,250,1999,2023]" captionStartId-3="82.[151,250,1943,1967]" captionStartId-4="83.[151,250,1976,2000]" captionTargetBox-0="[159,1427,185,1927]" captionTargetBox-1="[332,1252,185,1959]" captionTargetBox-2="[270,1315,181,1971]" captionTargetBox-3="[169,1416,181,1906]" captionTargetBox-4="[255,1326,182,1939]" captionTargetId-0="figure-17@79.[159,1429,181,1931]" captionTargetId-1="figure-19@80.[317,1270,181,1961]" captionTargetId-2="figure-17@81.[259,1328,181,1975]" captionTargetId-3="figure-19@82.[169,1419,181,1919]" captionTargetId-4="figure-17@83.[247,1341,181,1952]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionTargetPageId-2="81" captionTargetPageId-3="82" captionTargetPageId-4="83" captionText-0="FIGURE 44. Okanopteryx jeppesenorum holotype, SR 13-005-013. A, photograph of part SR 13-005-013A; B, photograph of counterpart SR 13-005-013B; C, drawing from part and counterpart.All to scale, 5 mm" captionText-1="FIGURE 45. Okanopteryx jeppesenorum paratype 1, SR 11-43-09. A, photograph; B, drawing. Both to scale, 5 mm." captionText-2="FIGURE 46. Okanopteryx jeppesenorum paratype 2, SR 18-002-001. A, photograph; B, drawing. Both to scale, 5 mm." captionText-3="FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm." captionText-4="FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558908" figureDoi-1="http://doi.org/10.5281/zenodo.4558910" figureDoi-2="http://doi.org/10.5281/zenodo.4558912" figureDoi-3="http://doi.org/10.5281/zenodo.4558914" figureDoi-4="http://doi.org/10.5281/zenodo.4558918" httpUri-0="https://zenodo.org/record/4558908/files/figure.png" httpUri-1="https://zenodo.org/record/4558910/files/figure.png" httpUri-2="https://zenodo.org/record/4558912/files/figure.png" httpUri-3="https://zenodo.org/record/4558914/files/figure.png" httpUri-4="https://zenodo.org/record/4558918/files/figure.png" pageId="78" pageNumber="79">Figs. 4448</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F005CFFD9FF5BFEE6FBBC13BB" pageId="78" pageNumber="79" type="diagnosis">
<paragraph id="8BA236D4005CFFD9FF5BFEE6FBBC13BB" blockId="78.[151,1437,259,2014]" pageId="78" pageNumber="79">
<emphasis id="B969EAC6005CFFD9FF5BFEE6FEBC1308" bold="true" box="[151,274,259,285]" pageId="78" pageNumber="79">Diagnosis.</emphasis>
Wings distinct from those of
<taxonomicName id="4C1D4D57005CFFD9FD9AFEE6FCA81308" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[598,774,259,285]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6005CFFD9FD9AFEE6FCA81308" box="[598,774,259,285]" italics="true" pageId="78" pageNumber="79">O. macabeensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D57005CFFD9FCDFFEE6FCD11308" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[787,895,259,285]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC6005CFFD9FCDFFEE6FCD11308" box="[787,895,259,285]" italics="true" pageId="78" pageNumber="79">O. fraseri</emphasis>
</taxonomicName>
by CuAA space wider, four cells wide at widest (difficult to count number of columns by preservation), between the levels of origins of IR1, RP2 [
<taxonomicName id="4C1D4D57005CFFD9FB2FFEC2FA3A1354" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1251,1428,295,321]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6005CFFD9FB2FFEC2FA3A1354" box="[1251,1428,295,321]" italics="true" pageId="78" pageNumber="79">O. macabeensis</emphasis>
</taxonomicName>
: 510 columns of three cells wide;
<taxonomicName id="4C1D4D57005CFFD9FDEDFEAEFD211370" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[545,655,331,357]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC6005CFFD9FDEDFEAEFD211370" box="[545,655,331,357]" italics="true" pageId="78" pageNumber="79">O. fraseri</emphasis>
</taxonomicName>
: all two cells wide, but sometimes a single column of three]; further from
<taxonomicName id="4C1D4D57005CFFD9FF1AFE8AFE26139C" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[214,392,367,393]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6005CFFD9FF1AFE8AFE26139C" box="[214,392,367,393]" italics="true" pageId="78" pageNumber="79">O. macabeensis</emphasis>
</taxonomicName>
, by longer wing,
<typeStatus id="54A68876005CFFD9FD9DFE8AFD18139C" box="[593,694,367,393]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
,
<quantity id="4CE59B31005CFFD9FD0EFE8AFC89139F" box="[706,807,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="78" pageNumber="79" unit="mm" value="22.4">22.4 mm</quantity>
nodus to apex;
<typeStatus id="54A68876005CFFD9FC17FE95FB95139F" box="[987,1083,368,394]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
1 (SR 11-43-09):
<quantity id="4CE59B31005CFFD9FAC7FE8AFADE139F" box="[1291,1392,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="78" pageNumber="79" unit="mm" value="23.5">23.5 mm</quantity>
[
<taxonomicName id="4C1D4D57005CFFD9FA4DFE8AFE8813B8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6005CFFD9FA4DFE8AFE8813B8" italics="true" pageId="78" pageNumber="79">O. macabeensis</emphasis>
</taxonomicName>
<typeStatus id="54A68876005CFFD9FEE1FE76FE3D13B8" box="[301,403,403,429]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
: estimated
<quantity id="4CE59B31005CFFD9FDDCFE76FDD913B8" box="[528,631,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="78" pageNumber="79" unit="mm" value="18.4">18.4 mm</quantity>
] (all
<taxonomicName id="4C1D4D57005CFFD9FD7DFE76FCB013B8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[689,798,403,429]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC6005CFFD9FD7DFE76FCB013B8" box="[689,798,403,429]" italics="true" pageId="78" pageNumber="79">O. fraseri</emphasis>
</taxonomicName>
wings missing apex).
</paragraph>
</subSubSection>
<subSubSection id="C307655F005CFFD9FF0BFE52FEC41100" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="8BA236D4005CFFD9FF0BFE52FEC41100" blockId="78.[151,1437,259,2014]" pageId="78" pageNumber="79">
<emphasis id="B969EAC6005CFFD9FF0BFE52FE9813C4" bold="true" box="[199,310,439,465]" pageId="78" pageNumber="79">Material.</emphasis>
<materialsCitation id="3B753C89005CFFD9FE8CFE52FEC41100" ID-GBIF-Occurrence="3069296335" collectingDate="2002-06-06" collectingDateMax="2019-05-01" collectingDateMin="2002-06-06" collectorName="Republic B &amp; Gregg Wilson" location="Stonerose Interpretive" pageId="78" pageNumber="79" specimenCount="7" stateProvince="Centre" typeStatus="holotype">
<emphasis id="B969EAC6005CFFD9FE8CFE52FE0913C4" box="[320,423,439,465]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FE8CFE52FE0913C4" box="[320,423,439,465]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
</emphasis>
: SR 13-005-013 (
<figureCitation id="13262A51005CFFD9FDBBFE52FD6313C4" box="[631,717,439,465]" captionStart="FIGURE 44" captionStartId="79.[151,250,1955,1979]" captionTargetBox="[159,1427,185,1927]" captionTargetId="figure-17@79.[159,1429,181,1931]" captionTargetPageId="79" captionText="FIGURE 44. Okanopteryx jeppesenorum holotype, SR 13-005-013. A, photograph of part SR 13-005-013A; B, photograph of counterpart SR 13-005-013B; C, drawing from part and counterpart.All to scale, 5 mm" figureDoi="http://doi.org/10.5281/zenodo.4558908" httpUri="https://zenodo.org/record/4558908/files/figure.png" pageId="78" pageNumber="79">Fig. 44</figureCitation>
), collected at B4131 by Jackie and Tim Jeppesen,
<date id="FFA31014005CFFD9FADAFE52FA3613C7" box="[1302,1432,439,466]" pageId="78" pageNumber="79" value="2013-07-15">
<collectingDate id="EFE7E9FC005CFFD9FADAFE52FA3613C7" box="[1302,1432,439,466]" pageId="78" pageNumber="79" value="2013-07-15">15.vii.2013</collectingDate>
</date>
, housed in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FDF4FE3EFD2D13E0" box="[568,643,475,501]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FCDFFE39FC2313E0" box="[787,909,475,501]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FCDFFE39FCD913E0" box="[787,887,476,501]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
1
</emphasis>
: SR 11-43-09, (
<figureCitation id="13262A51005CFFD9FB88FE3EFB3613E3" box="[1092,1176,475,502]" captionStart="FIGURE 45" captionStartId="80.[151,250,1985,2009]" captionTargetBox="[332,1252,185,1959]" captionTargetId="figure-19@80.[317,1270,181,1961]" captionTargetPageId="80" captionText="FIGURE 45. Okanopteryx jeppesenorum paratype 1, SR 11-43-09. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558910" httpUri="https://zenodo.org/record/4558910/files/figure.png" pageId="78" pageNumber="79">Fig. 45</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FAFBFE3EFF06100C" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by Christin Gustafson,
<date id="FFA31014005CFFD9FE21FE1AFDC8100F" box="[493,614,511,538]" pageId="78" pageNumber="79" value="2011-06-15">
<collectingDate id="EFE7E9FC005CFFD9FE21FE1AFDC8100F" box="[493,614,511,538]" pageId="78" pageNumber="79" value="2011-06-15">15.vi.2011</collectingDate>
</date>
, in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FC74FE1AFBAD100C" box="[952,1027,511,537]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FB5EFDE5FAA5100C" box="[1170,1291,511,537]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FB5EFDE5FB58100C" box="[1170,1270,512,537]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
2
</emphasis>
: SR 18-002- 001 (
<figureCitation id="13262A51005CFFD9FF18FDC6FE85102B" box="[212,299,547,574]" captionStart="FIGURE 46" captionStartId="81.[151,250,1999,2023]" captionTargetBox="[270,1315,181,1971]" captionTargetId="figure-17@81.[259,1328,181,1975]" captionTargetPageId="81" captionText="FIGURE 46. Okanopteryx jeppesenorum paratype 2, SR 18-002-001. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558912" httpUri="https://zenodo.org/record/4558912/files/figure.png" pageId="78" pageNumber="79">Fig. 46</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FE1EFDC6FDFC1028" box="[466,594,547,573]" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by Valerie Paquin,
<date id="FFA31014005CFFD9FCBAFDC6FC411028" box="[886,1007,547,573]" pageId="78" pageNumber="79" value="2018-04-28">
<collectingDate id="EFE7E9FC005CFFD9FCBAFDC6FC411028" box="[886,1007,547,573]" pageId="78" pageNumber="79" value="2018-04-28">28.iv.2018</collectingDate>
</date>
, in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FA9EFDC6FA331028" box="[1362,1437,547,573]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FEEDFDADFE341074" box="[289,410,583,609]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FEEDFDADFE2B1074" box="[289,389,584,609]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
3
</emphasis>
: SR 15-003-001 (
<figureCitation id="13262A51005CFFD9FDA4FDA2FD131074" box="[616,701,583,609]" captionStart="FIGURE 47" captionStartId="82.[151,250,1943,1967]" captionTargetBox="[169,1416,181,1906]" captionTargetId="figure-19@82.[169,1419,181,1919]" captionTargetPageId="82" captionText="FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558914" httpUri="https://zenodo.org/record/4558914/files/figure.png" pageId="78" pageNumber="79">Fig. 47</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FC90FDA2FC741074" box="[860,986,583,609]" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by Ethan Fisher,
<date id="FFA31014005CFFD9FB12FDA2FAE11077" box="[1246,1359,583,610]" pageId="78" pageNumber="79" value="2015-05-30">
<collectingDate id="EFE7E9FC005CFFD9FB12FDA2FAE11077" box="[1246,1359,583,610]" pageId="78" pageNumber="79" value="2015-05-30">30.v.2015</collectingDate>
</date>
, in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FE59FD8EFE4E1090" box="[405,480,619,645]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FDA2FD89FD481090" box="[622,742,619,645]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FDA2FD89FD7C1090" box="[622,722,620,645]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
4
</emphasis>
: SR 02-32-23 (
<figureCitation id="13262A51005CFFD9FC5FFD8EFC481090" box="[915,998,619,645]" captionStart="FIGURE 48" captionStartId="83.[151,250,1976,2000]" captionTargetBox="[255,1326,182,1939]" captionTargetId="figure-17@83.[247,1341,181,1952]" captionTargetPageId="83" captionText="FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558918" httpUri="https://zenodo.org/record/4558918/files/figure.png" pageId="78" pageNumber="79">Fig. 48</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FB4CFD8EFB531090" box="[1152,1277,619,645]" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by Chris Engle,
<date id="FFA31014005CFFD9FF2BFD6AFEFD10BC" box="[231,339,655,681]" pageId="78" pageNumber="79" value="2002-06-06">
<collectingDate id="EFE7E9FC005CFFD9FF2BFD6AFEFD10BC" box="[231,339,655,681]" pageId="78" pageNumber="79" value="2002-06-06">6.vi.2002</collectingDate>
</date>
, in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FD69FD6AFD5E10BC" box="[677,752,655,681]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FC4CFD75FC5710BC" box="[896,1017,655,681]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FC4CFD75FC4A10BC" box="[896,996,656,681]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
5
</emphasis>
: SR 07-22-05AB (
<figureCitation id="13262A51005CFFD9FB03FD6AFA8C10BC" box="[1231,1314,655,681]" captionStart="FIGURE 47" captionStartId="82.[151,250,1943,1967]" captionTargetBox="[169,1416,181,1906]" captionTargetId="figure-19@82.[169,1419,181,1919]" captionTargetPageId="82" captionText="FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558914" httpUri="https://zenodo.org/record/4558914/files/figure.png" pageId="78" pageNumber="79">Fig. 47</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FF7FFD56FE9E10D8" box="[179,304,691,717]" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by Kathleen Malloch
<date id="FFA31014005CFFD9FDA9FD56FD4110D8" box="[613,751,691,717]" pageId="78" pageNumber="79" value="2006-08-13">
<collectingDate id="EFE7E9FC005CFFD9FDA9FD56FD4110D8" box="[613,751,691,717]" pageId="78" pageNumber="79" value="2006-08-13">13.viii.2006</collectingDate>
</date>
, in the Stonerose Interpretive
<collectingRegion id="49D9F836005CFFD9FB8DFD56FB2210D8" box="[1089,1164,691,717]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections;
<emphasis id="B969EAC6005CFFD9FAD0FD51FA3A10D8" box="[1308,1428,691,717]" italics="true" pageId="78" pageNumber="79">
<typeStatus id="54A68876005CFFD9FAD0FD51FA2E10D8" box="[1308,1408,692,717]" pageId="78" pageNumber="79" type="paratype">paratype</typeStatus>
6
</emphasis>
: SR 19-005-002AB (
<figureCitation id="13262A51005CFFD9FEB5FD32FE6210E4" box="[377,460,727,753]" captionStart="FIGURE 48" captionStartId="83.[151,250,1976,2000]" captionTargetBox="[255,1326,182,1939]" captionTargetId="figure-17@83.[247,1341,181,1952]" captionTargetPageId="83" captionText="FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558918" httpUri="https://zenodo.org/record/4558918/files/figure.png" pageId="78" pageNumber="79">Fig. 48</figureCitation>
), collected at
<collectorName id="26E85302005CFFD9FDABFD32FD4A10E4" box="[615,740,727,753]" pageId="78" pageNumber="79">Republic B</collectorName>
4131 by
<collectorName id="26E85302005CFFD9FC8BFD32FC4810E4" box="[839,998,727,753]" pageId="78" pageNumber="79">Gregg Wilson</collectorName>
,
<date id="FFA31014005CFFD9FC3DFD32FBFA10E4" box="[1009,1108,727,753]" pageId="78" pageNumber="79" value="2019-05-01">
<collectingDate id="EFE7E9FC005CFFD9FC3DFD32FBFA10E4" box="[1009,1108,727,753]" pageId="78" pageNumber="79" value="2019-05-01">1.v.2019</collectingDate>
</date>
, in the
<location id="8EC2600F005CFFD9FB68FD32FA3310E4" LSID="urn:lsid:plazi:treatment:03B487C2005CFFC3FF5BFF72FC3014A8:8EC2600F005CFFD9FB68FD32FA3310E4" box="[1188,1437,727,753]" name="Stonerose Interpretive" pageId="78" pageNumber="79" stateProvince="Centre">Stonerose Interpretive</location>
<collectingRegion id="49D9F836005CFFD9FF5BFD1EFF4C1100" box="[151,226,763,789]" country="France" name="Centre" pageId="78" pageNumber="79">Center</collectingRegion>
collections.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307655F005CFFC3FF0BFCFAFEBE141C" lastPageId="84" lastPageNumber="85" pageId="78" pageNumber="79" type="description">
<paragraph id="8BA236D4005CFFD9FF0BFCFAFA251727" blockId="78.[151,1437,259,2014]" pageId="78" pageNumber="79">
<emphasis id="B969EAC6005CFFD9FF0BFCFAFEF7112C" bold="true" box="[199,345,799,825]" pageId="78" pageNumber="79">Description.</emphasis>
<typeStatus id="54A68876005CFFD9FEADFCFAFE62112C" box="[353,460,799,825]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
, SR 13-005-013, wing (
<figureCitation id="13262A51005CFFD9FD13FCFAFC9B112C" box="[735,821,799,825]" captionStart="FIGURE 44" captionStartId="79.[151,250,1955,1979]" captionTargetBox="[159,1427,185,1927]" captionTargetId="figure-17@79.[159,1429,181,1931]" captionTargetPageId="79" captionText="FIGURE 44. Okanopteryx jeppesenorum holotype, SR 13-005-013. A, photograph of part SR 13-005-013A; B, photograph of counterpart SR 13-005-013B; C, drawing from part and counterpart.All to scale, 5 mm" figureDoi="http://doi.org/10.5281/zenodo.4558908" httpUri="https://zenodo.org/record/4558908/files/figure.png" pageId="78" pageNumber="79">Fig. 44</figureCitation>
). Mostly complete, with portions of mid-posterior not preserved. Petiolate. Evenly oval distal to nodus, nodus to apex
<quantity id="4CE59B31005CFFD9FC93FCA6FC6B114B" box="[863,965,835,862]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="78" pageNumber="79" unit="mm" value="22.4">22.4 mm</quantity>
, maximum width mid-wing
<quantity id="4CE59B31005CFFD9FACEFCA1FAF4114B" box="[1282,1370,836,862]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="78" pageNumber="79" unit="mm" value="7.7">7.7 mm</quantity>
. Hyaline except for broad dark fascia across wing between level of nodus, just over half way to pterostigma. Pterostigma broad, about 2.5 times longer than wide, proximal side nearly vertical; subtends 2.5 cells. Oblique brace vein joins RA at proximal corner of pterostigma. Most crossveins roughly aligned in proximal half of postnodal, postsubnodal spaces. Costal space distal to pterostigma two cells wide. RA meets margin posterior to apex, RA slightly sigmoidal near margin. RARP1, RP1IR1 spaces narrow near margin. RARP1 space mostly one cell wide, briefly two near apex. RP1IR1 space two cells wide four cells distal to origin of IR1. Origin of IR1 presumed four cells distal to origin of RP2, mid-way between arculus, pterostigma. IR1RP2, RP2IR2 spaces two cells wide shortly proximal to level of pterostigma. Origin of RP2 about 56 cells distal to subnodus. RP2 mostly rather straight, curved slightly posteriorly in distal quarter. IR2: origin at subnodus; mostly rather straight, linear, slightly zigzagged in distal quarter, strongly so near margin. Origin of RP3-4 about 2/3 between arculus, nodus; mostly rather straight. MA linear, straight at least to half way from quadrangle (partially not preserved), strongly zigzagged at least distal third. MP linear, rather straight from quadrangle to terminus. CuA linear until shortly after level of nodus, then increasingly zigzagged. CuAA space with 4 cells at widest (from just proximal to, distal to level of RP2 origin as preserved).
</paragraph>
<paragraph id="8BA236D4005CFFD9FF0BFAD9FD4A153F" blockId="78.[151,1437,259,2014]" pageId="78" pageNumber="79">
<materialsCitation id="3B753C89005CFFD9FF0BFAD9FD4A153F" ID-GBIF-Occurrence="3069296302" collectionCode="RA, MA, MP, CuA" pageId="78" pageNumber="79" specimenCode="RP1, RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A68876005CFFD9FF0BFAD9FE871743" box="[199,297,1340,1366]" pageId="78" pageNumber="79" type="paratype">Paratype</typeStatus>
1, SR 11-43-09, wing (
<figureCitation id="13262A51005CFFD9FDFCFADEFD2A1743" box="[560,644,1339,1366]" captionStart="FIGURE 45" captionStartId="80.[151,250,1985,2009]" captionTargetBox="[332,1252,185,1959]" captionTargetId="figure-19@80.[317,1270,181,1961]" captionTargetPageId="80" captionText="FIGURE 45. Okanopteryx jeppesenorum paratype 1, SR 11-43-09. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558910" httpUri="https://zenodo.org/record/4558910/files/figure.png" pageId="78" pageNumber="79">Fig. 45</figureCitation>
). Petiolate. Evenly oval distal to nodus; nodus to apex
<quantity id="4CE59B31005CFFD9FB25FADEFAFF1743" box="[1257,1361,1339,1366]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="78" pageNumber="79" unit="mm" value="23.5">23.5 mm</quantity>
: maximum
<quantity id="4CE59B31005CFFD9FF14FABAFE80176F" box="[216,302,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="78" pageNumber="79" unit="mm" value="7.2">7.2 mm</quantity>
wide. Hyaline except for broad dark fascia across wing between level of nodus, just over half way to pterostigma. Pterostigma broad, about 2.5 times longer than wide; proximal side with shallow angle to costa. Region of oblique brace vein not preserved. Crossveins in postnodal, postsubnodal spaces mostly rather aligned. Two cells wide distal to pterostigma in costal space. RARP1 space one cell wide.
<collectionCode id="ED0CAE11005CFFD9FBC7FA2EFB9E17F0" box="[1035,1072,1483,1509]" pageId="78" pageNumber="79">RA</collectionCode>
meets margin posterior to apex.
<specimenCode id="DBBB9EAF005CFFD9FF5BFA0AFF69141C" box="[151,199,1519,1545]" collectionCode="RP" pageId="78" pageNumber="79">RP1</specimenCode>
IR1 space goes to two cells wide five cells distal to origin of IR1; narrows near margin. Origin of IR1 four cells distal to origin of
<specimenCode id="DBBB9EAF005CFFD9FEAAF9F6FE371438" box="[358,409,1555,1581]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
, mid-way between arculus, pterostigma. IR1
<specimenCode id="DBBB9EAF005CFFD9FC68F9F6FC791438" box="[932,983,1555,1581]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
,
<specimenCode id="DBBB9EAF005CFFD9FC28F9F6FBBA1438" box="[996,1044,1555,1581]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
IR2 spaces mostly not preserved.
<specimenCode id="DBBB9EAF005CFFD9FF5BF9D2FF651444" box="[151,203,1591,1617]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
: origin about 3 cells distal to subnodus; proximal half linear, rather straight; distal half not preserved. IR2: origin at subnodus; mostly rather straight, liner, slightly zigzagged in distal quarter, missing by preservation near margin.
<specimenCode id="DBBB9EAF005CFFD9FF38F99AFE91148C" box="[244,319,1663,1689]" collectionCode="RP" pageId="78" pageNumber="79">RP3-4</specimenCode>
: origin &lt;2/3 between arculus, nodus; mostly rather straight (not preserved near margin).
<collectionCode id="ED0CAE11005CFFD9FAE8F99AFAFE148C" box="[1316,1360,1663,1689]" pageId="78" pageNumber="79">MA</collectionCode>
linear, straight at least to half way from quadrangle; slightly zigzagged in small portion preserved distal to this.
<collectionCode id="ED0CAE11005CFFD9FAE4F941FAE114AB" box="[1320,1359,1700,1726]" country="0" name="Mohonk Preserve, Inc." pageId="78" pageNumber="79">MP</collectionCode>
linear, rather straight from quadrangle to terminus in regions preserved.
<collectionCode id="ED0CAE11005CFFD9FCBAF922FC0714F4" box="[886,937,1735,1761]" pageId="78" pageNumber="79">CuA</collectionCode>
slightly zigzagged from level of quadrangle to level of origin of
<specimenCode id="DBBB9EAF005CFFD9FEB2F90EFE1F1510" box="[382,433,1771,1797]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
, mostly not preserved apical to this. CuAA space with 4 cells wide shortly proximal to level of origin of
<specimenCode id="DBBB9EAF005CFFD9FE97F8EAFE25153C" box="[347,395,1807,1833]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
(portion distal to this missing)
</materialsCitation>
</paragraph>
<paragraph id="8BA236D4005CFFC3FF0BF8D1FF4C1370" blockId="78.[151,1437,259,2014]" lastBlockId="84.[151,1437,151,1725]" lastPageId="84" lastPageNumber="85" pageId="78" pageNumber="79">
<materialsCitation id="3B753C89005CFFC3FF0BF8D1FF701370" ID-GBIF-Occurrence="3069296305" collectionCode="MA, MP, CuA" lastPageId="84" lastPageNumber="85" pageId="78" pageNumber="79" specimenCode="RP1, RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A68876005CFFD9FF0BF8D1FE87155B" box="[199,297,1844,1870]" pageId="78" pageNumber="79" type="paratype">Paratype</typeStatus>
2, SR 18-002-001, wing (
<figureCitation id="13262A51005CFFD9FD8BF8D6FD34155B" box="[583,666,1843,1870]" captionStart="FIGURE 46" captionStartId="81.[151,250,1999,2023]" captionTargetBox="[270,1315,181,1971]" captionTargetId="figure-17@81.[259,1328,181,1975]" captionTargetPageId="81" captionText="FIGURE 46. Okanopteryx jeppesenorum paratype 2, SR 18-002-001. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558912" httpUri="https://zenodo.org/record/4558912/files/figure.png" pageId="78" pageNumber="79">Fig. 46</figureCitation>
). Petiolate. Evenly oval distal to nodus (missing apical region), nodus to base of pterostigma
<quantity id="4CE59B31005CFFD9FE50F8B2FDAD1567" box="[412,515,1879,1906]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="78" pageNumber="79" unit="mm" value="16.5">16.5 mm</quantity>
, maximum width mid-wing
<quantity id="4CE59B31005CFFD9FC84F8B2FC0C1567" box="[840,930,1879,1906]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="78" pageNumber="79" unit="mm" value="7.2">7.2 mm</quantity>
. Hyaline except for broad dark fascia across wing between level of nodus, to about presumed 2/3 distance to pterostigma. Pterostigma not preserved. Most crossveins roughly aligned in proximal half of postnodal, postsubnodal spaces, not in distal half.
<specimenCode id="DBBB9EAF005CFFD9FB21F87AFAB315AC" box="[1261,1309,1951,1977]" collectionCode="RP" pageId="78" pageNumber="79">RP1</specimenCode>
IR1 space two cells wide eight cells distal to origin of IR1. IR1 origin two cells distal to origin of
<specimenCode id="DBBB9EAF005CFFD9FBACF826FB3A15C8" box="[1120,1172,1987,2013]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF005CFFD9FB52F826FB7C15C8" box="[1182,1234,1987,2013]" collectionCode="RP" pageId="78" pageNumber="79">RP2</specimenCode>
: origin apparently five cells distal to subnodus; mostly rather straight in region preserved. IR2: origin at subnodus; mostly rather straight, linear, apical quarter missing.
<specimenCode id="DBBB9EAF0046FFC3FD9AFF5EFD0F12C0" box="[598,673,187,213]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
: origin missing; mostly rather straight, missing apical quarter.
<collectionCode id="ED0CAE110046FFC3FABDFF5EFA3312C0" box="[1393,1437,187,213]" pageId="84" pageNumber="85">MA</collectionCode>
linear, straight at least to half way between nodus, presumed level of pterostigma.
<collectionCode id="ED0CAE110046FFC3FB8EFF3AFBC712EC" box="[1090,1129,223,249]" country="0" name="Mohonk Preserve, Inc." pageId="84" pageNumber="85">MP</collectionCode>
linear, rather straight from quadrangle to terminus.
<collectionCode id="ED0CAE110046FFC3FE64FEE6FE751308" box="[424,475,259,285]" pageId="84" pageNumber="85">CuA</collectionCode>
linear until shortly after level of nodus, then slightly zigzagged to distal quarter, then increasingly so. CuAA space four cells at widest from just proximal to level of origin of
<specimenCode id="DBBB9EAF0046FFC3FB55FEC2FB641354" box="[1177,1226,295,321]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
to about level IP2 origin
</materialsCitation>
.
</paragraph>
<caption id="DF62665C005DFFD8FF5BF846FBA415CA" ID-DOI="http://doi.org/10.5281/zenodo.4558908" ID-Zenodo-Dep="4558908" httpUri="https://zenodo.org/record/4558908/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,250,1955,1979]" targetBox="[159,1427,185,1927]" targetPageId="79">
<paragraph id="8BA236D4005DFFD8FF5BF846FBA415CA" blockId="79.[151,1436,1955,2015]" pageId="79" pageNumber="80">
<emphasis id="B969EAC6005DFFD8FF5BF846FE8E15AE" bold="true" box="[151,288,1955,1979]" pageId="79" pageNumber="80">FIGURE 44.</emphasis>
<taxonomicName id="4C1D4D57005DFFD8FEEAF846FD9015AE" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,574,1955,1979]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC6005DFFD8FEEAF846FD9015AE" box="[294,574,1955,1979]" italics="true" pageId="79" pageNumber="80">Okanopteryx jeppesenorum</emphasis>
</taxonomicName>
holotype, SR 13-005-013. A, photograph of part SR 13-005-013A; B, photograph of counterpart SR 13-005-013B; C, drawing from part and counterpart. All to scale, 5 mm
</paragraph>
</caption>
<caption id="DF62665C0042FFC7FF5BF824FA9915CC" ID-DOI="http://doi.org/10.5281/zenodo.4558910" ID-Zenodo-Dep="4558910" box="[151,1335,1985,2010]" httpUri="https://zenodo.org/record/4558910/files/figure.png" pageId="80" pageNumber="81" startId="80.[151,250,1985,2009]" targetBox="[332,1252,185,1959]" targetPageId="80">
<paragraph id="8BA236D40042FFC7FF5BF824FA9915CC" blockId="80.[151,1335,1985,2010]" box="[151,1335,1985,2010]" pageId="80" pageNumber="81">
<emphasis id="B969EAC60042FFC7FF5BF824FE8E15CC" bold="true" box="[151,288,1985,2009]" pageId="80" pageNumber="81">FIGURE 45.</emphasis>
<taxonomicName id="4C1D4D570042FFC7FEEAF824FD9315CC" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,573,1985,2009]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC60042FFC7FEEAF824FD9315CC" box="[294,573,1985,2009]" italics="true" pageId="80" pageNumber="81">Okanopteryx jeppesenorum</emphasis>
</taxonomicName>
paratype 1, SR 11-43-09. A, photograph; B, drawing. Both to scale, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C0043FFC6FF5BF82AFAFF15F2" ID-DOI="http://doi.org/10.5281/zenodo.4558912" ID-Zenodo-Dep="4558912" box="[151,1361,1999,2024]" httpUri="https://zenodo.org/record/4558912/files/figure.png" pageId="81" pageNumber="82" startId="81.[151,250,1999,2023]" targetBox="[270,1315,181,1971]" targetPageId="81">
<paragraph id="8BA236D40043FFC6FF5BF82AFAFF15F2" blockId="81.[151,1361,1999,2024]" box="[151,1361,1999,2024]" pageId="81" pageNumber="82">
<emphasis id="B969EAC60043FFC6FF5BF82AFE8E15F2" bold="true" box="[151,288,1999,2023]" pageId="81" pageNumber="82">FIGURE 46.</emphasis>
<taxonomicName id="4C1D4D570043FFC6FEEAF82AFD9315F2" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,573,1999,2023]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC60043FFC6FEEAF82AFD9315F2" box="[294,573,1999,2023]" italics="true" pageId="81" pageNumber="82">Okanopteryx jeppesenorum</emphasis>
</taxonomicName>
paratype 2, SR 18-002-001. A, photograph; B, drawing. Both to scale, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C0040FFC5FF5BF872FDF915C6" ID-DOI="http://doi.org/10.5281/zenodo.4558914" ID-Zenodo-Dep="4558914" httpUri="https://zenodo.org/record/4558914/files/figure.png" pageId="82" pageNumber="83" startId="82.[151,250,1943,1967]" targetBox="[169,1416,181,1906]" targetPageId="82">
<paragraph id="8BA236D40040FFC5FF5BF872FDF915C6" blockId="82.[151,1436,1943,2004]" pageId="82" pageNumber="83">
<emphasis id="B969EAC60040FFC5FF5BF872FE8E15BA" bold="true" box="[151,288,1943,1967]" pageId="82" pageNumber="83">FIGURE 47.</emphasis>
<taxonomicName id="4C1D4D570040FFC5FEEAF87DFD9315BA" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,573,1944,1968]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC60040FFC5FEEAF87DFD9315BA" box="[294,573,1944,1968]" italics="true" pageId="82" pageNumber="83">Okanopteryx jeppesenorum</emphasis>
</taxonomicName>
paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C0041FFC4FF5BF85DFDF915E6" ID-DOI="http://doi.org/10.5281/zenodo.4558918" ID-Zenodo-Dep="4558918" httpUri="https://zenodo.org/record/4558918/files/figure.png" pageId="83" pageNumber="84" startId="83.[151,250,1976,2000]" targetBox="[255,1326,182,1939]" targetPageId="83">
<paragraph id="8BA236D40041FFC4FF5BF85DFDF915E6" blockId="83.[151,1436,1976,2036]" pageId="83" pageNumber="84">
<emphasis id="B969EAC60041FFC4FF5BF85DFE8E15C5" bold="true" box="[151,288,1976,2000]" pageId="83" pageNumber="84">FIGURE 48.</emphasis>
<taxonomicName id="4C1D4D570041FFC4FEEAF85DFD9315C5" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,573,1976,2000]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC60041FFC4FEEAF85DFD9315C5" box="[294,573,1976,2000]" italics="true" pageId="83" pageNumber="84">Okanopteryx jeppesenorum</emphasis>
</taxonomicName>
paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm.
</paragraph>
</caption>
<paragraph id="8BA236D40046FFC3FF0BFE8AFC4C10BC" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<materialsCitation id="3B753C890046FFC3FF0BFE8AFDFF13C4" ID-GBIF-Occurrence="3069296321" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP1" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A688760046FFC3FF0BFE8AFE87139C" box="[199,297,367,393]" pageId="84" pageNumber="85" type="paratype">Paratype</typeStatus>
3, SR 15-003-001, wing (
<figureCitation id="13262A510046FFC3FD9DFE8AFD08139C" box="[593,678,367,393]" captionStart="FIGURE 47" captionStartId="82.[151,250,1943,1967]" captionTargetBox="[169,1416,181,1906]" captionTargetId="figure-19@82.[169,1419,181,1919]" captionTargetPageId="82" captionText="FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558914" httpUri="https://zenodo.org/record/4558914/files/figure.png" pageId="84" pageNumber="85">Fig. 47</figureCitation>
). Proximal two-thirds. Petiolate. Maximum width
<quantity id="4CE59B310046FFC3FB13FE8AFA96139C" box="[1247,1336,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="84" pageNumber="85" unit="mm" value="7.7">7.7 mm</quantity>
. Hyaline proximal to level of nodus, darkly infuscate distal to it. Most crossveins roughly aligned in proximal half preserved of postnodal, postsubnodal spaces.
<specimenCode id="DBBB9EAF0046FFC3FDEEFE52FDFF13C4" box="[546,593,439,465]" collectionCode="RP" pageId="84" pageNumber="85">RP1</specimenCode>
</materialsCitation>
<materialsCitation id="3B753C890046FFC3FDAEFE52FDB913E0" ID-GBIF-Occurrence="3069296333" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
IR1 space two cells wide five cells distal to origin of IR1. IR1: origin four cells distal to origin of
<specimenCode id="DBBB9EAF0046FFC3FE6FFE3EFE7913E0" box="[419,471,475,501]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0046FFC3FE2FFE3EFDB913E0" box="[483,535,475,501]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
</materialsCitation>
:
<materialsCitation id="3B753C890046FFC3FDE8FE3EFBEC100C" ID-GBIF-Occurrence="3069296375" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP3-4" specimenCount="1" typeStatus="paratype">
origin five cells distal to subnodus; mostly rather straight in region preserved. IR2: origin at subnodus; mostly rather straight, linear, apical half missing.
<specimenCode id="DBBB9EAF0046FFC3FC3BFE1AFBEC100C" box="[1015,1090,511,537]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
</materialsCitation>
: origin missing; mostly rather straight, missing apical third. MA
<materialsCitation id="3B753C890046FFC3FDD1FDC6FB5E1028" ID-GBIF-Occurrence="3069296338" box="[541,1264,547,573]" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
mostly linear, slightly zigzagged around levels of origin of
<specimenCode id="DBBB9EAF0046FFC3FB0CFDC6FB5E1028" box="[1216,1264,547,573]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
</materialsCitation>
to IR1, mostly not preserved distal to that. MP mostly linear, slightly zigzagged in portions, gently curved from quadrangle, near terminus not preserved. CuA
<materialsCitation id="3B753C890046FFC3FE12FD8EFCD31090" ID-GBIF-Occurrence="3069296387" box="[478,893,619,645]" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
rather linear to level of origin of
<specimenCode id="DBBB9EAF0046FFC3FC86FD8EFCD31090" box="[842,893,619,645]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B753C890046FFC3FC44FD8EFC7010BC" ID-GBIF-Occurrence="3069296388" collectionCode="RP" pageId="84" pageNumber="85" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
then increasingly zigzagged. CuAA space four cells at widest from just proximal to, to just distal to level of origin of
<specimenCode id="DBBB9EAF0046FFC3FC66FD6AFC7010BC" box="[938,990,655,681]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA236D40046FFC3FF0BFD56FAF5164C" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<materialsCitation id="3B753C890046FFC3FF0BFD56FAF9164C" ID-GBIF-Occurrence="3069296339" collectionCode="RA, MA, MP, CuA" pageId="84" pageNumber="85" specimenCode="RP1, RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A688760046FFC3FF0BFD56FE8710D8" box="[199,297,691,717]" pageId="84" pageNumber="85" type="paratype">Paratype</typeStatus>
4, SR 02-32-23, wing (
<figureCitation id="13262A510046FFC3FDF8FD56FD2710D8" box="[564,649,691,717]" captionStart="FIGURE 48" captionStartId="83.[151,250,1976,2000]" captionTargetBox="[255,1326,182,1939]" captionTargetId="figure-17@83.[247,1341,181,1952]" captionTargetPageId="83" captionText="FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558918" httpUri="https://zenodo.org/record/4558918/files/figure.png" pageId="84" pageNumber="85">Fig. 48</figureCitation>
). Apical half, missing much of anterior region. Apical portion hyaline from about half way between origin of IR1, pterostigma; darkly infuscate proximal to this. Pterostigma broad, proximal portion not preserved. Postnodal, most of postsubnodal spaces not preserved. Costal space distal to pterostigma two cells wide.
<collectionCode id="ED0CAE110046FFC3FE8AFCFAFEC5112C" box="[326,363,799,825]" pageId="84" pageNumber="85">RA</collectionCode>
meets margin posterior to apex. RARP1,
<specimenCode id="DBBB9EAF0046FFC3FC87FCFAFCD4112C" box="[843,890,799,825]" collectionCode="RP" pageId="84" pageNumber="85">RP1</specimenCode>
IR1 spaces narrow near margin. RARP1 space mostly one cell wide, briefly two near apex.
<specimenCode id="DBBB9EAF0046FFC3FD4EFCA6FD1C1148" box="[642,690,835,861]" collectionCode="RP" pageId="84" pageNumber="85">RP1</specimenCode>
IR1 space constantly two cells wide twelve cells distal to origin of IR1. Origin of IR1 presumed three cells distal to origin of
<specimenCode id="DBBB9EAF0046FFC3FCECFC82FCFA1194" box="[800,852,871,897]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
. IR1
<specimenCode id="DBBB9EAF0046FFC3FC58FC82FC691194" box="[916,967,871,897]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
space two cells wide at presumed level of proximal side of pterostigma; IR2
<specimenCode id="DBBB9EAF0046FFC3FDD2FC6EFDCA11B0" box="[542,612,907,933]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
space two cells wide from a few cells proximal to this.
<specimenCode id="DBBB9EAF0046FFC3FB0BFC6EFB5511B0" box="[1223,1275,907,933]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
: linear, mostly rather straight, slightly curved posteriorly in apical quarter. IR2 straight, linear to level
<specimenCode id="DBBB9EAF0046FFC3FB94FC4AFB2611DC" box="[1112,1160,943,969]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
IR2 space becomes two cells wide, then slightly zigzagged, curves posteriorly near terminus.
<specimenCode id="DBBB9EAF0046FFC3FC6FFC36FC4011F8" box="[931,1006,979,1005]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
: mostly rather straight, wing damaged near margin.
<collectionCode id="ED0CAE110046FFC3FEE5FC12FEFB1604" box="[297,341,1015,1041]" pageId="84" pageNumber="85">MA</collectionCode>
preserved portion linear to about level of origin of IR1, then increasingly zigzagged.
<collectionCode id="ED0CAE110046FFC3FACEFC12FA871604" box="[1282,1321,1015,1041]" country="0" name="Mohonk Preserve, Inc." pageId="84" pageNumber="85">MP</collectionCode>
preserved portion linear, slightly curved to terminus.
<collectionCode id="ED0CAE110046FFC3FD4FFBFEFD181620" box="[643,694,1051,1077]" pageId="84" pageNumber="85">CuA</collectionCode>
increasingly zigzagged in region preserved, strongly so apically. CuAA space four cells wide from just proximal to level of
<specimenCode id="DBBB9EAF0046FFC3FCF4FBDAFCC6164C" box="[824,872,1087,1113]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
origin to just distal to level of origin of IR1
</materialsCitation>
.
</paragraph>
<paragraph id="8BA236D40046FFC3FF0BFB86FE561724" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<materialsCitation id="3B753C890046FFC3FF0BFB86FE5A1724" ID-GBIF-Occurrence="3069296347" collectionCode="A, MA, MP" pageId="84" pageNumber="85" specimenCode="RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A688760046FFC3FF0BFB86FE871668" box="[199,297,1123,1149]" pageId="84" pageNumber="85" type="paratype">Paratype</typeStatus>
5, SR 07-22-05
<collectionCode id="ED0CAE110046FFC3FE2EFB87FE561668" box="[482,504,1122,1149]" pageId="84" pageNumber="85">A</collectionCode>
, B, wing (
<figureCitation id="13262A510046FFC3FDBDFB86FD691668" box="[625,711,1123,1149]" captionStart="FIGURE 47" captionStartId="82.[151,250,1943,1967]" captionTargetBox="[169,1416,181,1906]" captionTargetId="figure-19@82.[169,1419,181,1919]" captionTargetPageId="82" captionText="FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558914" httpUri="https://zenodo.org/record/4558914/files/figure.png" pageId="84" pageNumber="85">Fig. 47</figureCitation>
). Parts of proximal half. Petiolate. Hyaline proximal to level of nodus, darkly infuscate distal to it. Most crossveins in postnodal, postsubnodal spaces roughly aligned in region preserved. Origin of IR1 three cells distal to origin of
<specimenCode id="DBBB9EAF0046FFC3FD1DFB4EFCAB16D0" box="[721,773,1195,1221]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0046FFC3FCDCFB4EFCEA16D0" box="[784,836,1195,1221]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
: origin 2.5 cells distal to subnodus; portion preserved rather straight, linear. IR2: origin at subnodus; straight, linear.
<specimenCode id="DBBB9EAF0046FFC3FCAAFB2AFC1E16FC" box="[870,944,1231,1257]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
,
<collectionCode id="ED0CAE110046FFC3FC72FB2AFC4016FC" box="[958,1006,1231,1257]" pageId="84" pageNumber="85">MA</collectionCode>
,
<collectionCode id="ED0CAE110046FFC3FC37FB2AFB8616FC" box="[1019,1064,1231,1257]" country="0" name="Mohonk Preserve, Inc." pageId="84" pageNumber="85">MP</collectionCode>
: origins, distal portions missing; preserved portion mostly rather straight, rather linear. CuAA space 4 cells wide just proximal to level of
<specimenCode id="DBBB9EAF0046FFC3FAF0FB16FAC21718" box="[1340,1388,1267,1293]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
origin, not preserved distal to this
</materialsCitation>
.
</paragraph>
<paragraph id="8BA236D40046FFC3FF0BFADEFEBE141C" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<materialsCitation id="3B753C890046FFC3FF0BFADEFEA2141C" ID-GBIF-Occurrence="3069296327" collectionCode="A, MA, MP" pageId="84" pageNumber="85" specimenCode="RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A688760046FFC3FF0BFADEFE871740" box="[199,297,1339,1365]" pageId="84" pageNumber="85" type="paratype">Paratype</typeStatus>
6, SR 19-005-002
<collectionCode id="ED0CAE110046FFC3FE35FADFFDA11740" box="[505,527,1338,1365]" pageId="84" pageNumber="85">A</collectionCode>
, B, wing (
<figureCitation id="13262A510046FFC3FD4DFADEFD7B1740" box="[641,725,1339,1365]" captionStart="FIGURE 48" captionStartId="83.[151,250,1976,2000]" captionTargetBox="[255,1326,182,1939]" captionTargetId="figure-17@83.[247,1341,181,1952]" captionTargetPageId="83" captionText="FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558918" httpUri="https://zenodo.org/record/4558918/files/figure.png" pageId="84" pageNumber="85">Fig. 48</figureCitation>
). Broken up middle portion from closely distal to nodus to base of pterostigma, posterior region missing or damaged. Proximal portion darkly infuscate, distal portion hyaline. Many crossveins roughly aligned in preserved portion of postnodal, postsubnodal spaces. IR1 origin four cells distal to origin of
<specimenCode id="DBBB9EAF0046FFC3FED0FA42FEFE17D4" box="[284,336,1447,1473]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0046FFC3FE96FA42FE2317D4" box="[346,397,1447,1473]" collectionCode="RP" pageId="84" pageNumber="85">RP2</specimenCode>
, IR2,
<specimenCode id="DBBB9EAF0046FFC3FE1CFA42FDB517D4" box="[464,539,1447,1473]" collectionCode="RP" pageId="84" pageNumber="85">RP3-4</specimenCode>
,
<collectionCode id="ED0CAE110046FFC3FDEAFA42FDF917D4" box="[550,599,1447,1473]" pageId="84" pageNumber="85">MA</collectionCode>
: preserved portions mostly linear, rather straight, except
<collectionCode id="ED0CAE110046FFC3FB1EFA42FB5017D4" box="[1234,1278,1447,1473]" pageId="84" pageNumber="85">MA</collectionCode>
zigzagging in distal portion preserved. Small portions of
<collectionCode id="ED0CAE110046FFC3FDB2FA2EFD0B17F0" box="[638,677,1483,1509]" country="0" name="Mohonk Preserve, Inc." pageId="84" pageNumber="85">MP</collectionCode>
reserved. Disarticulated fragment of CuAA space preserved, four cells wide
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F0046FFC3FF0BF9F6FC3014A8" pageId="84" pageNumber="85" type="etymology">
<paragraph id="8BA236D40046FFC3FF0BF9F6FED11460" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<emphasis id="B969EAC60046FFC3FF0BF9F6FEE61438" bold="true" box="[199,328,1555,1581]" pageId="84" pageNumber="85">Etymology</emphasis>
. The specific epithet is a patronymic formed from the surname of Jackie and Tim Jeppesen, the collectors and donors of the
<typeStatus id="54A688760046FFC3FE1AF9D2FD951444" box="[470,571,1591,1617]" pageId="84" pageNumber="85" type="holotype">holotype</typeStatus>
, recognising their contribution. The Latin suffix
<emphasis id="B969EAC60046FFC3FBA0F9D2FB091445" box="[1132,1191,1591,1616]" italics="true" pageId="84" pageNumber="85">orum</emphasis>
indicates the genitive case of plural nouns.
</paragraph>
<paragraph id="8BA236D40046FFC3FF0BF99AFC3014A8" blockId="84.[151,1437,151,1725]" pageId="84" pageNumber="85">
<emphasis id="B969EAC60046FFC3FF0BF99AFED5148C" bold="true" box="[199,379,1663,1689]" pageId="84" pageNumber="85">Range and age</emphasis>
. Tom Thumb Tuff Member of the Klondike Mountain Formation, locality B4131 [and other maybe?] at Republic,
<collectingRegion id="49D9F8360046FFC3FE40F946FDBA14A8" box="[396,532,1699,1725]" country="United States of America" name="Washington" pageId="84" pageNumber="85">Washington</collectingRegion>
,
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; second half of the Ypresian.
</paragraph>
</subSubSection>
</treatment>
</document>