339 lines
38 KiB
XML
339 lines
38 KiB
XML
<document id="C3AC30DDB58A9D7725C053E80B368A3E" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. & Mathewes, Rolf W." docDate="2021" docId="03B487C20051FFD4FF5BFDC7FCBD15AC" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Okanagrion dorrellae Archibald & Cannings & Erickson & Bybee & Mathewes 2021, new species" docType="treatment" docVersion="7" lastPageNumber="68" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="68" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:titleInfo id="E323C0A5A7DDFC762F117A59113E7CAD">
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<mods:title id="3D1F169E278FB753463C2755B671032B">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="A519CA473F6CB7D6F565C67C4371913D">Archibald, Bruce</mods:namePart>
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<mods:namePart id="FF595D65C97873305C643712E3D1CCD0">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="C284615C285A3D3F4D54E8D468876A5F" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:nameIdentifier id="9E95EE3F81B90E6A6035BB511285078B" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:roleTerm id="1EB3DB23F2D50A610564BABD1BF66AFD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="108389A860753165122B16CE95264320">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="D32ED746A8302D35C56AFCCB0BCC800A" type="ORCID">0000-0003-1162-0355</mods:nameIdentifier>
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<mods:nameIdentifier id="58B79160A548268153B438E04F37BD5D" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="502F72E4D712D438B3E2DCDC6B83D806">Bybee, Seth M.</mods:namePart>
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<mods:nameIdentifier id="CCAC1451519CBDD01476D0507FEEFFAE" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8A49130383A913A2EB4B92DAEC8777AD">Mathewes, Rolf W.</mods:namePart>
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<mods:nameIdentifier id="BBB736FE499E2C2E119C4780C46129CE" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
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<mods:title id="551E48B8B2D34305E5069AE7C0169AEE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3EDCD494D98BDD7C1A6DE14417C838BC">
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<mods:date id="41CFAB77B9C459017D5F6CA4E8C2C8D0">2021</mods:date>
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<mods:number id="52D31749207827498D2BC1C7AD54C0E7">2021-02-24</mods:number>
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<mods:number id="79AFB9BACE0F41AC62567017AB7FFF50">4934</mods:number>
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<mods:classification id="CD1CD27BAB89AE72B93E58B595143065">journal article</mods:classification>
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<mods:identifier id="94DC206C5F067C66D8C346E8B805B8A3" type="CLB-Dataset">7439</mods:identifier>
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<mods:identifier id="A0B9499A7270B4E431A8828657D13597" type="DOI">10.11646/zootaxa.4934.1.1</mods:identifier>
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<mods:identifier id="C87F2E19A8CD46125B59053507500BEA" type="GBIF-Dataset">300bac18-55f5-4e34-90e1-7fccd02d68c1</mods:identifier>
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<mods:identifier id="916E2E211A2CA7B55049E034FEE581CF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="158BFE8909D8AC03DE7C3835AAC5BF7C" type="Zenodo-Dep">4558796</mods:identifier>
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<treatment id="03B487C20051FFD4FF5BFDC7FCBD15AC" ID-DOI="http://doi.org/10.5281/zenodo.4672744" ID-GBIF-Taxon="180881809" ID-Zenodo-Dep="4672744" LSID="urn:lsid:plazi:treatment:03B487C20051FFD4FF5BFDC7FCBD15AC" httpUri="http://treatment.plazi.org/id/03B487C20051FFD4FF5BFDC7FCBD15AC" lastPageNumber="68" pageId="67" pageNumber="68">
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<subSubSection id="C307655F0051FFD4FF5BFDC7FCF11029" box="[151,863,546,573]" pageId="67" pageNumber="68" type="nomenclature">
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<paragraph id="8BA236D40051FFD4FF5BFDC7FCF11029" blockId="67.[151,863,546,609]" box="[151,863,546,573]" pageId="67" pageNumber="68">
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<heading id="D0EA81B80051FFD4FF5BFDC7FCF11029" bold="true" box="[151,863,546,573]" fontSize="11" level="2" pageId="67" pageNumber="68" reason="4">
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<emphasis id="B969EAC60051FFD4FF5BFDC7FCF11029" bold="true" box="[151,863,546,573]" pageId="67" pageNumber="68">
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<taxonomicName id="4C1D4D570051FFD4FF5BFDC7FD6D1029" authority="Archibald & Cannings" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,707,546,573]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="dorrellae" status="sp. nov.">
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<emphasis id="B969EAC60051FFD4FF5BFDC7FE301028" bold="true" box="[151,414,546,573]" italics="true" pageId="67" pageNumber="68">Okanagrion dorrellae</emphasis>
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Archibald & Cannings
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25A57BD0051FFD4FD03FDC6FCF11029" box="[719,863,546,573]" pageId="67" pageNumber="68" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F0051FFD4FF5BFDA2FF401074" box="[151,238,583,609]" pageId="67" pageNumber="68" type="description">
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<paragraph id="8BA236D40051FFD4FF5BFDA2FF401074" blockId="67.[151,863,546,609]" box="[151,238,583,609]" pageId="67" pageNumber="68">
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<figureCitation id="13262A510051FFD4FF5BFDA2FF441074" box="[151,234,583,609]" captionStart="FIGURE 37" captionStartId="68.[151,250,2004,2028]" captionTargetBox="[288,1302,184,1976]" captionTargetId="figure-19@68.[277,1311,181,1980]" captionTargetPageId="68" captionText="FIGURE 37. Okanagrion dorrellae holotype, SR 93-11-02. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558890" httpUri="https://zenodo.org/record/4558890/files/figure.png" pageId="67" pageNumber="68">Fig. 37</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F0051FFD4FF5BFD6BFBFB16FC" pageId="67" pageNumber="68" type="diagnosis">
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<paragraph id="8BA236D40051FFD4FF5BFD6BFAC11604" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
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<emphasis id="B969EAC60051FFD4FF5BFD6BFEBC10BD" bold="true" box="[151,274,654,680]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
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Wings separated from those of all other species of
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<taxonomicName id="4C1D4D570051FFD4FC91FD6AFC4A10BC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[861,996,655,681]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAC60051FFD4FC91FD6AFC4A10BC" box="[861,996,655,681]" italics="true" pageId="67" pageNumber="68">Okanagrion</emphasis>
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</taxonomicName>
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by MP zigzagged in distal portion [all others: linear (but not known in
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<taxonomicName id="4C1D4D570051FFD4FE36FD56FD2610D9" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[506,648,691,717]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
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||
<emphasis id="B969EAC60051FFD4FE36FD56FDA010D8" box="[506,526,691,717]" italics="true" pageId="67" pageNumber="68">O</emphasis>
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||
.
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||
<emphasis id="B969EAC60051FFD4FDD7FD56FD2610D9" box="[539,648,691,716]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
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</taxonomicName>
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)]; from all but
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<taxonomicName id="4C1D4D570051FFD4FCFCFD56FC1010D9" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[816,958,691,717]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
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<emphasis id="B969EAC60051FFD4FCFCFD56FCEA10D8" box="[816,836,691,717]" italics="true" pageId="67" pageNumber="68">O</emphasis>
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.
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<emphasis id="B969EAC60051FFD4FC9DFD56FC1010D9" box="[849,958,691,716]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
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</taxonomicName>
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by narrower shape, especially in the proximal portion: maximum width 7.0 mm [
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<taxonomicName id="4C1D4D570051FFD4FD81FD32FD5910E4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[589,759,727,753]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="threadgillae">
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<emphasis id="B969EAC60051FFD4FD81FD32FD5910E4" box="[589,759,727,753]" italics="true" pageId="67" pageNumber="68">O. threadgillae</emphasis>
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</taxonomicName>
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: forewing, hind wing: 9.0 mm;
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<taxonomicName id="4C1D4D570051FFD4FB91FD32FB6310E4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1117,1229,727,753]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="hobani">
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<emphasis id="B969EAC60051FFD4FB91FD32FBDF10E4" box="[1117,1137,727,753]" italics="true" pageId="67" pageNumber="68">O</emphasis>
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.
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<emphasis id="B969EAC60051FFD4FBB3FD32FB6310E4" box="[1151,1229,727,753]" italics="true" pageId="67" pageNumber="68">hobani</emphasis>
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</taxonomicName>
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all wings:
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<quantity id="4CE59B310051FFD4FA84FD33FF681100" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="67" pageNumber="68" unit="mm" value="8.85" valueMax="9.5" valueMin="8.2">8.2–9.5 mm</quantity>
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;
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<taxonomicName id="4C1D4D570051FFD4FF1DFD1EFE941100" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[209,314,763,789]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
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<emphasis id="B969EAC60051FFD4FF1DFD1EFE941100" box="[209,314,763,789]" italics="true" pageId="67" pageNumber="68">O. beardi</emphasis>
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</taxonomicName>
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: 7.7,
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<quantity id="4CE59B310051FFD4FEB4FD1EFE7E1100" box="[376,464,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="67" pageNumber="68" unit="mm" value="8.3">8.3 mm</quantity>
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;
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<taxonomicName id="4C1D4D570051FFD4FE16FD1EFD361100" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[474,664,763,789]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="liquetoalatum">
|
||
<emphasis id="B969EAC60051FFD4FE16FD1EFD361100" box="[474,664,763,789]" italics="true" pageId="67" pageNumber="68">O. liquetoalatum</emphasis>
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||
</taxonomicName>
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: forewing
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||
<quantity id="4CE59B310051FFD4FCDCFD1FFCC61100" box="[784,872,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="67" pageNumber="68" unit="mm" value="8.5">8.5 mm</quantity>
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;
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||
<taxonomicName id="4C1D4D570051FFD4FCBFFD1EFC551100" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[883,1019,763,789]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="worleyae">
|
||
<emphasis id="B969EAC60051FFD4FCBFFD1EFC551100" box="[883,1019,763,789]" italics="true" pageId="67" pageNumber="68">O. worleyae</emphasis>
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||
</taxonomicName>
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: about
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||
<quantity id="4CE59B310051FFD4FB81FD1EFB091100" box="[1101,1191,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="67" pageNumber="68" unit="mm" value="7.8">7.8 mm</quantity>
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], wing length (arculus to apex) / width, 3.7 [
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||
<taxonomicName id="4C1D4D570051FFD4FE43FCFAFDAF112C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[399,513,799,825]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60051FFD4FE43FCFAFE0D112C" box="[399,419,799,825]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FE7FFCFAFDAF112C" box="[435,513,799,825]" italics="true" pageId="67" pageNumber="68">hobani</emphasis>
|
||
</taxonomicName>
|
||
: 3.2;
|
||
<taxonomicName id="4C1D4D570051FFD4FD88FCFAFD41112C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[580,751,799,825]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC60051FFD4FD88FCFAFD41112C" box="[580,751,799,825]" italics="true" pageId="67" pageNumber="68">O. threadgillae</emphasis>
|
||
</taxonomicName>
|
||
: forewing 3.0, hind wing 3.1;
|
||
<taxonomicName id="4C1D4D570051FFD4FB80FCFAFB16112C" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1100,1208,799,825]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
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||
<emphasis id="B969EAC60051FFD4FB80FCFAFB16112C" box="[1100,1208,799,825]" italics="true" pageId="67" pageNumber="68">O. beardi</emphasis>
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||
</taxonomicName>
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||
: forewings 3.3, 3.4,
|
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<taxonomicName id="4C1D4D570051FFD4FF5BFCA6FEFB1148" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,341,835,861]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="liquetoalatum">
|
||
<emphasis id="B969EAC60051FFD4FF5BFCA6FEFB1148" box="[151,341,835,861]" italics="true" pageId="67" pageNumber="68">O. liquetoalatum</emphasis>
|
||
</taxonomicName>
|
||
: forewing 2.6 (
|
||
<taxonomicName id="4C1D4D570051FFD4FDCDFCA6FD241148" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[513,650,835,861]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lochmum">
|
||
<emphasis id="B969EAC60051FFD4FDCDFCA6FD241148" box="[513,650,835,861]" italics="true" pageId="67" pageNumber="68">O. lochmum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570051FFD4FD5BFCA6FC8E1148" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[663,800,835,861]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="worleyae">
|
||
<emphasis id="B969EAC60051FFD4FD5BFCA6FC8E1148" box="[663,800,835,861]" italics="true" pageId="67" pageNumber="68">O. worleyae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570051FFD4FCE1FCA6FC121149" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[813,956,835,861]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
|
||
<emphasis id="B969EAC60051FFD4FCE1FCA6FCEF1148" box="[813,833,835,861]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FC83FCA6FC121149" box="[847,956,835,860]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
|
||
</taxonomicName>
|
||
not comparable by preservation)]; from
|
||
<taxonomicName id="4C1D4D570051FFD4FA4DFCA6FF4B1194" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60051FFD4FA4DFCA6FF4B1194" italics="true" pageId="67" pageNumber="68">O. hobani</emphasis>
|
||
</taxonomicName>
|
||
by main veins less deeply curved near margin; from
|
||
<taxonomicName id="4C1D4D570051FFD4FCF7FC82FC641195" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[827,970,871,897]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
|
||
<emphasis id="B969EAC60051FFD4FCF7FC82FCE11194" box="[827,847,871,897]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FC91FC82FC641195" box="[861,970,871,896]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570051FFD4FC15FC82FBED1194" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[985,1091,871,897]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC60051FFD4FC15FC82FBED1194" box="[985,1091,871,897]" italics="true" pageId="67" pageNumber="68">O. beardi</emphasis>
|
||
</taxonomicName>
|
||
by IR2 origin between RP1-2, RP3-4 [
|
||
<taxonomicName id="4C1D4D570051FFD4FF3CFC6EFE2E11B1" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[240,384,907,933]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
|
||
<emphasis id="B969EAC60051FFD4FF3CFC6EFEAA11B0" box="[240,260,907,933]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FEDFFC6EFE2E11B1" box="[275,384,907,932]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570051FFD4FE42FC6EFE5711B0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[398,505,907,933]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC60051FFD4FE42FC6EFE5711B0" box="[398,505,907,933]" italics="true" pageId="67" pageNumber="68">O. beardi</emphasis>
|
||
</taxonomicName>
|
||
: origin on RP1-2]; from
|
||
<taxonomicName id="4C1D4D570051FFD4FCD9FC6EFC3011B0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[789,926,907,933]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lochmum">
|
||
<emphasis id="B969EAC60051FFD4FCD9FC6EFC8711B0" box="[789,809,907,933]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FCFBFC6EFC3011B0" box="[823,926,907,933]" italics="true" pageId="67" pageNumber="68">lochmum</emphasis>
|
||
</taxonomicName>
|
||
by cells less dense in apical portion of wing: see RA–RP1 space becomes two cells wide, three cells wide, four–five cells wide as it its diagnosis; from all other
|
||
<taxonomicName id="4C1D4D570051FFD4FF5BFC36FEB011F8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,286,979,1005]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC60051FFD4FF5BFC36FEB011F8" box="[151,286,979,1005]" italics="true" pageId="67" pageNumber="68">Okanagrion</emphasis>
|
||
</taxonomicName>
|
||
species but
|
||
<taxonomicName id="4C1D4D570051FFD4FE6AFC36FDBE11F8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[422,528,979,1005]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC60051FFD4FE6AFC36FE1411F8" box="[422,442,979,1005]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FE04FC36FDBE11F8" box="[456,528,979,1005]" italics="true" pageId="67" pageNumber="68">beardi</emphasis>
|
||
</taxonomicName>
|
||
by MP, CuA divergent near margin [others: MP-CuA space remaining narrow;
|
||
<taxonomicName id="4C1D4D570051FFD4FA4DFC36FF711604" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC60051FFD4FA4DFC36FA3B11F8" box="[1409,1429,979,1005]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FF5BFC12FF711604" box="[151,223,1015,1041]" italics="true" pageId="67" pageNumber="68">beardi</emphasis>
|
||
</taxonomicName>
|
||
: distinctly divergent,
|
||
<taxonomicName id="4C1D4D570051FFD4FE18FC12FDF01604" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[468,606,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="dorrellae">
|
||
<emphasis id="B969EAC60051FFD4FE18FC12FE461604" box="[468,488,1015,1041]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FE3BFC12FDF01604" box="[503,606,1015,1041]" italics="true" pageId="67" pageNumber="68">dorrellae</emphasis>
|
||
</taxonomicName>
|
||
: less distinctly as CuA zigzagged distally (
|
||
<taxonomicName id="4C1D4D570051FFD4FB8DFC12FB7E1605" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1089,1232,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
|
||
<emphasis id="B969EAC60051FFD4FB8DFC12FBFB1604" box="[1089,1109,1015,1041]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FBAFFC12FB7E1605" box="[1123,1232,1015,1040]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
|
||
</taxonomicName>
|
||
, not known)].
|
||
</paragraph>
|
||
<paragraph id="8BA236D40051FFD4FF0BFBFEFBFB16FC" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
Further separated by colouration (sex unknown): proximal near half hyaline, dark apical to this except lighter fascia just distal to mid-way between nodus, pterostigma, extending from anterior margin to apparently mid-wing (some damage); size and extent of light fascia differs from:
|
||
<taxonomicName id="4C1D4D570051FFD4FC8FFB86FC1B1668" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[835,949,1123,1149]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60051FFD4FC8FFB86FCF91668" box="[835,855,1123,1149]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FCABFB86FC1B1668" box="[871,949,1123,1149]" italics="true" pageId="67" pageNumber="68">hobani</emphasis>
|
||
</taxonomicName>
|
||
about twice or more as wide, extending to or almost to posterior margin;
|
||
<taxonomicName id="4C1D4D570051FFD4FE20FB62FDD516B5" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[492,635,1159,1185]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="angustum">
|
||
<emphasis id="B969EAC60051FFD4FE20FB62FDAE16B4" box="[492,512,1159,1185]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FDC2FB62FDD516B5" box="[526,635,1159,1184]" italics="true" pageId="67" pageNumber="68">angustum</emphasis>
|
||
</taxonomicName>
|
||
apparently similar to
|
||
<taxonomicName id="4C1D4D570051FFD4FCBCFB62FC4E16B4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[880,992,1159,1185]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="hobani">
|
||
<emphasis id="B969EAC60051FFD4FCBCFB62FC2A16B4" box="[880,900,1159,1185]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FC5EFB62FC4E16B4" box="[914,992,1159,1185]" italics="true" pageId="67" pageNumber="68">hobani</emphasis>
|
||
</taxonomicName>
|
||
as preserved;
|
||
<taxonomicName id="4C1D4D570051FFD4FB4CFB62FA8516B4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1152,1323,1159,1185]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="threadgillae">
|
||
<emphasis id="B969EAC60051FFD4FB4CFB62FB3A16B4" box="[1152,1172,1159,1185]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FB6FFB62FA8516B4" box="[1187,1323,1159,1185]" italics="true" pageId="67" pageNumber="68">threadgillae</emphasis>
|
||
</taxonomicName>
|
||
narrower, extends to mid-wing;
|
||
<taxonomicName id="4C1D4D570051FFD4FE46FB4EFE5D16D0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[394,499,1195,1221]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="beardi">
|
||
<emphasis id="B969EAC60051FFD4FE46FB4EFE3016D0" box="[394,414,1195,1221]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FE67FB4EFE5D16D0" box="[427,499,1195,1221]" italics="true" pageId="67" pageNumber="68">beardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D570051FFD4FDCCFB4EFD2716D0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[512,649,1195,1221]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lochmum">
|
||
<emphasis id="B969EAC60051FFD4FDCCFB4EFDBA16D0" box="[512,532,1195,1221]" italics="true" pageId="67" pageNumber="68">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60051FFD4FDEEFB4EFD2716D0" box="[546,649,1195,1221]" italics="true" pageId="67" pageNumber="68">lochmum</emphasis>
|
||
</taxonomicName>
|
||
, similar width, apparently extends closer to posterior margin;
|
||
<taxonomicName id="4C1D4D570051FFD4FAF6FB4EFF5516FC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="liquetoalatum">
|
||
<emphasis id="B969EAC60051FFD4FAF6FB4EFF5516FC" italics="true" pageId="67" pageNumber="68">O. liquetoalatum</emphasis>
|
||
</taxonomicName>
|
||
: hyaline throughout;
|
||
<taxonomicName id="4C1D4D570051FFD4FE21FB2AFDD816FC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[493,630,1231,1257]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="worleyae">
|
||
<emphasis id="B969EAC60051FFD4FE21FB2AFDD816FC" box="[493,630,1231,1257]" italics="true" pageId="67" pageNumber="68">O. worleyae</emphasis>
|
||
</taxonomicName>
|
||
: lightly infuscate throughout known wing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0051FFD4FF0BFB16FB401724" pageId="67" pageNumber="68" type="materials_examined">
|
||
<paragraph id="8BA236D40051FFD4FF0BFB16FB401724" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
<emphasis id="B969EAC60051FFD4FF0BFB16FED91719" bold="true" box="[199,375,1266,1293]" pageId="67" pageNumber="68">Type material.</emphasis>
|
||
<emphasis id="B969EAC60051FFD4FE4CFB16FE491718" box="[384,487,1267,1293]" italics="true" pageId="67" pageNumber="68">
|
||
<typeStatus id="54A688760051FFD4FE4CFB16FE491718" box="[384,487,1267,1293]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: SR 93-11-02 (
|
||
<figureCitation id="13262A510051FFD4FD56FB16FD5E1718" box="[666,752,1267,1293]" captionStart="FIGURE 37" captionStartId="68.[151,250,2004,2028]" captionTargetBox="[288,1302,184,1976]" captionTargetId="figure-19@68.[277,1311,181,1980]" captionTargetPageId="68" captionText="FIGURE 37. Okanagrion dorrellae holotype, SR 93-11-02. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558890" httpUri="https://zenodo.org/record/4558890/files/figure.png" pageId="67" pageNumber="68">Fig. 37</figureCitation>
|
||
), collected Republic B9201 by Laurie Dorrell,
|
||
<date id="FFA310140051FFD4FADCFB16FA2C1718" box="[1296,1410,1267,1293]" pageId="67" pageNumber="68" value="1992-10-31">31.x.1992</date>
|
||
, a complete wing except for diagonal break across mid-section, with colour patterning; SR collection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0051FFD4FF0BFADFFB8814A8" pageId="67" pageNumber="68" type="description">
|
||
<paragraph id="8BA236D40051FFD4FF0BFADFFB8814A8" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
<emphasis id="B969EAC60051FFD4FF0BFADFFEF71741" bold="true" box="[199,345,1338,1364]" pageId="67" pageNumber="68">Description.</emphasis>
|
||
<typeStatus id="54A688760051FFD4FEAEFADEFE631740" box="[354,461,1339,1365]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
|
||
, SR 93-11-02, wing. Shape: oval distal to nodus, symmetrical to lateral mid-line, but apex slightly posterior to this. Length: arculus to apex:
|
||
<quantity id="4CE59B310051FFD4FCC7FABBFCDC176C" box="[779,882,1374,1401]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="67" pageNumber="68" unit="mm" value="25.6">25.6 mm</quantity>
|
||
, nodus to apex:
|
||
<quantity id="4CE59B310051FFD4FBE0FABAFB3D176C" box="[1068,1171,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="67" pageNumber="68" unit="mm" value="21.1">21.1 mm</quantity>
|
||
, origin of RP2 to apex:
|
||
<quantity id="4CE59B310051FFD4FF5BFA67FF511788" box="[151,255,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="67" pageNumber="68" unit="mm" value="16.5">16.5 mm</quantity>
|
||
, arculus to base of pterostigma:
|
||
<quantity id="4CE59B310051FFD4FDBBFA66FD711788" box="[631,735,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="67" pageNumber="68" unit="mm" value="19.7">19.7 mm</quantity>
|
||
, nodus to base of pterostigma:
|
||
<quantity id="4CE59B310051FFD4FB84FA67FB1F1788" box="[1096,1201,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="67" pageNumber="68" unit="mm" value="15.3">15.3 mm</quantity>
|
||
, nodus at 26% wing length; width 7.0 mm. Colouration as in diagnosis. Pterostigma incompletely preserved, region of possible oblique brace vein not preserved. 35 crossveins in postnodal space (total), those in postsubnodal space not aligned. At least three cells wide maximum in costal space distal to pterostigma, but most of space not preserved. RA meets margin anterior to apex. RA–RP1 space narrows near margin. IR1: origin nine cells distal to origin of RP2, IR1 two cells wide probably (poorly preserved) eight cells distal to origin. RP2: origin seven cells distal to origin of IR2 (one very small). IR2: origin at subnodus. RP3-4 origin in distal part of middle third between arculus, nodus. Arculus just proximal to Ax2. No crossvein O. All main veins linear except base of IR1, MP distal to mid-wing between nodus, pterostigma. CuA–A space apparently (by preservation) five cells wide at widest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0051FFD4FF0BF922FED31558" pageId="67" pageNumber="68" type="etymology">
|
||
<paragraph id="8BA236D40051FFD4FF0BF922FDDB1510" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
<emphasis id="B969EAC60051FFD4FF0BF922FEE614F4" bold="true" box="[199,328,1735,1761]" pageId="67" pageNumber="68">Etymology</emphasis>
|
||
. The specific epithet is a patronymic formed from the surname of Laurie Dorrell, the collector of the
|
||
<typeStatus id="54A688760051FFD4FF0CF90EFE8B1510" box="[192,293,1771,1797]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
, recognising her contribution.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40051FFD4FF0BF8EAFED31558" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
<emphasis id="B969EAC60051FFD4FF0BF8EAFED6153C" bold="true" box="[199,376,1807,1833]" pageId="67" pageNumber="68">Range and age</emphasis>
|
||
. Republic locality B9201 of the Tom Thumb Tuff Member of the Klondike Mountain Formation; latest Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0051FFD4FF0BF8B3FCBD15AC" pageId="67" pageNumber="68" type="discussion">
|
||
<paragraph id="8BA236D40051FFD4FF0BF8B3FCBD15AC" blockId="67.[151,1437,654,1977]" pageId="67" pageNumber="68">
|
||
<emphasis id="B969EAC60051FFD4FF0BF8B3FEE21565" bold="true" box="[199,332,1878,1904]" pageId="67" pageNumber="68">Discussion.</emphasis>
|
||
Although the
|
||
<typeStatus id="54A688760051FFD4FE22F8B2FDFE1564" box="[494,592,1879,1905]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
|
||
wing and sole specimen of
|
||
<taxonomicName id="4C1D4D570051FFD4FC42F8B2FBB61564" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[910,1048,1879,1905]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="dorrellae">
|
||
<emphasis id="B969EAC60051FFD4FC42F8B2FBB61564" box="[910,1048,1879,1905]" italics="true" pageId="67" pageNumber="68">O. dorrellae</emphasis>
|
||
</taxonomicName>
|
||
has a diagonal break across it, we are confident of the shape as the veins are not shifted across this break. The wing has not been compressed, as internal elements such as the shape of cells are not deformed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |