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<document id="76B661A77A09B90C9AFF2613AC2130D3" ID-DOI="10.5281/zenodo.5169515" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169515" ID-ZooBank="7F2A2366-B4E4-4F37-A5A5-45CB51D4D859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277797618" checkinUser="felipe" docAuthor="Rifkind, Jacques" docDate="2017" docId="03F1E067CC0FFFF310CFFAF4FE51FCA5" docLanguage="en" docName="InsectaMundi.0591.1-18.pdf" docOrigin="Insecta Mundi 2017 (591)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Quadrophenia sumidero , Rifkind 2017, new species" docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="FFC8981FCC06FFF9100FFF9EFF94FFDB" masterDocTitle="New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="8" updateTime="1699039302695" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="968E9AD49E1C0B290867EC9E5577091D">New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea)</mods:title>
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<mods:namePart id="CBFEC52F43DC65D232D55415BCC2C275">Rifkind, Jacques</mods:namePart>
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<mods:title id="F5A681F022378B95F57107E0A8C2DAB6">Insecta Mundi</mods:title>
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<mods:date id="18341BC37BC026F92EB6460479E9E71E">2017</mods:date>
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<mods:number id="09D9E58C2A6FC3FE4506E0F8FEFC8E2E">2017-12-29</mods:number>
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<treatment id="03F1E067CC0FFFF310CFFAF4FE51FCA5" ID-DOI="http://doi.org/10.5281/zenodo.6487887" ID-Zenodo-Dep="6487887" LSID="urn:lsid:plazi:treatment:03F1E067CC0FFFF310CFFAF4FE51FCA5" httpUri="http://treatment.plazi.org/id/03F1E067CC0FFFF310CFFAF4FE51FCA5" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<subSubSection id="C34202FACC0FFFF010CFFAF4FCA1FA59" box="[192,821,1386,1410]" pageId="9" pageNumber="8" type="nomenclature">
<paragraph id="8BE75171CC0FFFF010CFFAF4FCA1FA59" blockId="9.[192,821,1386,1442]" box="[192,821,1386,1410]" pageId="9" pageNumber="8">
<heading id="D0AFE61DCC0FFFF010CFFAF4FCA1FA59" bold="true" box="[192,821,1386,1410]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
<emphasis id="B92C8D63CC0FFFF010CFFAF4FCA1FA59" bold="true" box="[192,821,1386,1410]" pageId="9" pageNumber="8">
<taxonomicName id="4C582AF2CC0FFFF010CFFAF4FD17FA59" authority="Rifkind" authorityName=", Rifkind" authorityYear="2017" box="[192,643,1386,1410]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="sumidero" status="sp. nov.">
<emphasis id="B92C8D63CC0FFFF010CFFAF4FD84FA59" bold="true" box="[192,528,1386,1410]" italics="true" pageId="9" pageNumber="8">Quadrophenia sumidero</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="A21F3018CC0FFFF0129EFAF4FCA1FA59" box="[657,821,1386,1410]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0FFFF010CFFA14FE14FA79" box="[192,384,1418,1442]" pageId="9" pageNumber="8" type="description">
<paragraph id="8BE75171CC0FFFF010CFFA14FE14FA79" blockId="9.[192,821,1386,1442]" box="[192,384,1418,1442]" pageId="9" pageNumber="8">
(
<figureCitation id="13634DF4CC0FFFF010C6FA14FED9FA79" box="[201,333,1418,1442]" captionStart="Figures 1213" captionStartId="17.[192,288,1903,1925]" captionTargetBox="[363,1286,289,1732]" captionTargetId="figure-28@17.[346,1342,267,1759]" captionTargetPageId="17" captionText="Figures 1213. Quadrophenia sumidero new species. 12) habitus; 13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5169523" httpUri="https://zenodo.org/record/5169523/files/figure.png" pageId="9" pageNumber="8">Fig. 12, 13</figureCitation>
,
<figureCitation id="13634DF4CC0FFFF01156FA14FEE3FA79" box="[345,375,1418,1442]" captionStart="Figures 1721" captionStartId="19.[192,288,1875,1897]" captionText="Figures 1721. Quadrophenia new genus, antennae of new species. 17) Q. townshendi, antenna; 18) Q. mooni, capitulum; 19) Q. entwistlei, capitulum; 20) Q. daltreyi, capitulum; 21) Q. sumidero, capitulum." pageId="9" pageNumber="8">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0FFFF010CFFA55FDCBF9D8" pageId="9" pageNumber="8" type="materials_examined">
<paragraph id="8BE75171CC0FFFF010CFFA55FDCBF9D8" blockId="9.[192,1440,1483,1539]" pageId="9" pageNumber="8">
<emphasis id="B92C8D63CC0FFFF010CFFA55FE1FFA38" bold="true" box="[192,395,1483,1507]" pageId="9" pageNumber="8">Type material.</emphasis>
<materialsCitation id="3B305B2CCC0FFFF0119CFA55FF66F9D8" collectingDate="1987-06-15" collectorName="E. Giesbert" country="Mexico" location="Chiapas" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="54E3EFD3CC0FFFF0119CFA55FD91FA38" box="[403,517,1483,1507]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
, male.
<collectingCountry id="F34F11E1CC0FFFF01253FA55FD22FA38" box="[604,694,1483,1507]" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingCountry>
,
<collectingRegion id="499C9F93CC0FFFF012CDFA55FCBEFA38" box="[706,810,1483,1507]" country="Mexico" name="Chiapas" pageId="9" pageNumber="8">Chiapas</collectingRegion>
, Sumidero Cyn., 4000,
<date id="FFE677B1CC0FFFF0145FFA55FA94FA38" box="[1104,1280,1483,1507]" pageId="9" pageNumber="8" value="1987-06-15">
<collectingDate id="EFA28E59CC0FFFF0145FFA55FA94FA38" box="[1104,1280,1483,1507]" pageId="9" pageNumber="8" value="1987-06-15">June 15, 1987</collectingDate>
</date>
,
<collectorName id="26AD34A7CC0FFFF01502FA55FA0FFA38" box="[1293,1435,1483,1507]" pageId="9" pageNumber="8">E. Giesbert</collectorName>
, coll.
</materialsCitation>
<materialsCitation id="3B305B2CCC0FFFF010F6FA75FDCFF9D8" box="[249,603,1515,1539]" collectionCode="CSCA" country="Mexico" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="54E3EFD3CC0FFFF010F6FA75FEFCF9D8" box="[249,360,1515,1539]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="ED49C9B4CC0FFFF01203FA75FDCFF9D8" box="[524,603,1515,1539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="9" pageNumber="8" type="Museum">CSCA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0FFFF010CFF9B5FE6FF9BF" pageId="9" pageNumber="8" type="diagnosis">
<paragraph id="8BE75171CC0FFFF010CFF9B5FE6FF9BF" blockId="9.[192,1440,1579,1636]" pageId="9" pageNumber="8">
<emphasis id="B92C8D63CC0FFFF010CFF9B5FEC4F998" bold="true" box="[192,336,1579,1603]" pageId="9" pageNumber="8">Diagnosis.</emphasis>
This is the only member of its genus with the elytra marked by boldly contrasting black and cream-colored bands.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0FFFF310CFF912FD67FE40" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="description">
<paragraph id="8BE75171CC0FFFF310CFF912FE47FE81" blockId="9.[191,1442,1676,1926]" lastBlockId="10.[192,1440,194,346]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<emphasis id="B92C8D63CC0FFFF010CFF912FEFEF97F" bold="true" box="[192,362,1676,1700]" pageId="9" pageNumber="8">Description.</emphasis>
(
<typeStatus id="54E3EFD3CC0FFFF01177F912FE78F97F" box="[376,492,1676,1700]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
). Length: 8.80 mm. Form: subflattened dorsally, slightly expanded posteriorly. Color: piceous; antennomeres 13 (in part), labrum, and mouthparts (except mandibles), testaceous; elytra with a broad, irregularly bordered cream-colored fascia at middle, fascia narrowed laterally, then extending in a line anteriorly along epipleural margin to approximately anterior 1/5 (
<figureCitation id="13634DF4CC0FFFF01560F973FE84F8FE" captionStart="Figures 1213" captionStartId="17.[192,288,1903,1925]" captionTargetBox="[363,1286,289,1732]" captionTargetId="figure-28@17.[346,1342,267,1759]" captionTargetPageId="17" captionText="Figures 1213. Quadrophenia sumidero new species. 12) habitus; 13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5169523" httpUri="https://zenodo.org/record/5169523/files/figure.png" pageId="9" pageNumber="8">Fig. 1213</figureCitation>
); elytral posterior 1/4 also cream-colored. Antennae: terminal antennomere with internal and external / distal emarginations exaggerated; proximal angle of external emarginate plane produced as a short spine, apex subacuminate (
<figureCitation id="13634DF4CC0FFFF01299F8D0FD7BF8BD" box="[662,751,1870,1894]" captionStart="Figures 1721" captionStartId="19.[192,288,1875,1897]" captionText="Figures 1721. Quadrophenia new genus, antennae of new species. 17) Q. townshendi, antenna; 18) Q. mooni, capitulum; 19) Q. entwistlei, capitulum; 20) Q. daltreyi, capitulum; 21) Q. sumidero, capitulum." pageId="9" pageNumber="8">Fig. 21</figureCitation>
). Pronotum: sides rather convex. Elytra: ratio of length to maximum width 17:10, broadest at posterior 4/11; apices subsinuate; surface coarsely, densely, moderately deeply punctate; asetiferous punctures uniformly dispersed; vestiture sparse, inconspicuous, composed of erect and suberect infuscate setae of various lengths. Abdomen: ventrite 5 with hind angles rounded, posterior margin very feebly, arcuately emarginate; ventrite 6 with hind angles broadly rounded, posterior margin with a shallow U-shaped inflection at middle; tergite 6 with hind margin rounded.
</paragraph>
<paragraph id="8BE75171CC0CFFF310CFFE1DFD67FE40" blockId="10.[192,755,387,411]" box="[192,755,387,411]" pageId="10" pageNumber="9">
<emphasis id="B92C8D63CC0CFFF310CFFE1DFED8FE40" bold="true" box="[192,332,387,411]" pageId="10" pageNumber="9">Variation.</emphasis>
Only the
<typeStatus id="54E3EFD3CC0CFFF311C9FE1DFE6FFE40" box="[454,507,387,411]" pageId="10" pageNumber="9">type</typeStatus>
specimen is known.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0CFFF310CFFE5DFC48FE27" pageId="10" pageNumber="9" type="distribution">
<paragraph id="8BE75171CC0CFFF310CFFE5DFC48FE27" blockId="10.[191,1440,451,508]" pageId="10" pageNumber="9">
<emphasis id="B92C8D63CC0CFFF310CFFE5DFEE0FE00" bold="true" box="[192,372,451,475]" pageId="10" pageNumber="9">Distribution.</emphasis>
<taxonomicName id="4C582AF2CC0CFFF31175FE5DFD30FE00" authorityName=", Rifkind" authorityYear="2017" box="[378,676,451,475]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sumidero">
<emphasis id="B92C8D63CC0CFFF31175FE5DFD30FE00" box="[378,676,451,475]" italics="true" pageId="10" pageNumber="9">Quadrophenia sumidero</emphasis>
</taxonomicName>
is known only from Sumidero Canyon in
<collectingRegion id="499C9F93CC0CFFF31496FE5DFA95FE00" box="[1177,1281,451,475]" country="Mexico" name="Chiapas" pageId="10" pageNumber="9">Chiapas</collectingRegion>
,
<collectingCountry id="F34F11E1CC0CFFF31503FE5DFAF2FE00" box="[1292,1382,451,475]" name="Mexico" pageId="10" pageNumber="9">Mexico</collectingCountry>
. The habitat at the
<typeStatus id="54E3EFD3CC0CFFF3117FFE7AFE31FE27" box="[368,421,484,508]" pageId="10" pageNumber="9">type</typeStatus>
locality is primarily tropical deciduous forest.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0CFFF310CFFDBAFAE0FDE7" box="[192,1396,548,573]" pageId="10" pageNumber="9" type="etymology">
<paragraph id="8BE75171CC0CFFF310CFFDBAFAE0FDE7" blockId="10.[192,1396,548,573]" box="[192,1396,548,573]" pageId="10" pageNumber="9">
<emphasis id="B92C8D63CC0CFFF310CFFDBAFEC8FDE7" bold="true" box="[192,348,548,572]" pageId="10" pageNumber="9">Etymology.</emphasis>
The specific epithet references the
<typeStatus id="54E3EFD3CC0CFFF31300FDBBFCD0FDE6" box="[783,836,549,573]" pageId="10" pageNumber="9">type</typeStatus>
locality. It is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34202FACC0CFFF310CFFDFBFE51FCA5" pageId="10" pageNumber="9" type="discussion">
<paragraph id="8BE75171CC0CFFF310CFFDFBFE51FCA5" blockId="10.[190,1441,613,894]" pageId="10" pageNumber="9">
<emphasis id="B92C8D63CC0CFFF310CFFDFBFED7FDA6" bold="true" box="[192,323,613,637]" pageId="10" pageNumber="9">Remarks.</emphasis>
Based on its bold markings,
<taxonomicName id="4C582AF2CC0CFFF31298FDFBFC2AFDA6" authorityName=", Rifkind" authorityYear="2017" box="[663,958,613,637]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sumidero">
<emphasis id="B92C8D63CC0CFFF31298FDFBFC2AFDA6" box="[663,958,613,637]" italics="true" pageId="10" pageNumber="9">Quadrophenia sumidero</emphasis>
</taxonomicName>
is presumably a mimic, but I am unable as yet to posit a similar, aposematically patterned sympatric model. Another specimen of
<taxonomicName id="4C582AF2CC0CFFF31501FD1BFF7CFD66" authorityName="Rifkind" authorityYear="2017" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B92C8D63CC0CFFF31501FD1BFF7CFD66" italics="true" pageId="10" pageNumber="9">Quadrophenia</emphasis>
</taxonomicName>
from the type locality of
<taxonomicName id="4C582AF2CC0CFFF3122CFD3BFD2DFD66" authorityName=", Rifkind" authorityYear="2017" box="[547,697,677,701]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sumidero">
<emphasis id="B92C8D63CC0CFFF3122CFD3BFD2DFD66" box="[547,697,677,701]" italics="true" pageId="10" pageNumber="9">Q. sumidero</emphasis>
</taxonomicName>
is noted here, but not included in the type series because it shows salient differences in habitus that I cannot with confidence attribute to infraspecific variation. This specimen, a male, measures 8.50 mm in length. It differs from the
<typeStatus id="54E3EFD3CC0CFFF31440FD7BFB2CFD26" box="[1103,1208,741,765]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C582AF2CC0CFFF314EEFD78FAEDFD26" authorityName=", Rifkind" authorityYear="2017" box="[1249,1401,741,765]" class="Insecta" family="Melyridae" genus="Quadrophenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sumidero">
<emphasis id="B92C8D63CC0CFFF314EEFD78FAEDFD26" box="[1249,1401,741,765]" italics="true" pageId="10" pageNumber="9">Q. sumidero</emphasis>
</taxonomicName>
by having the elytra significantly more dorsoventrally flattened, duller, and more coarsely punctate, and by its coloration: the prosternum, the anterior transverse impression dorsolaterally, and the pronotal collar dorsolaterally are cream-colored; the posterior elytral cream-colored band is reduced to a thin line along the apices.
</paragraph>
</subSubSection>
</treatment>
</document>