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<document id="D4A3ED5B7B56704D6E070D0616DE2DFB" ID-DOI="10.5281/zenodo.5404983" ID-ISSN="1942-1354" ID-Zenodo-Dep="5404983" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628267599528" checkinUser="felipe" docAuthor="Stebnicka, Z. Teresa &amp; Skelley, Paul E." docDate="2009" docId="03E0CE76FFBA8278F3CDF96DFCC3FF57" docLanguage="en" docName="InsectaMundi.0066.1-18.pdf" docOrigin="Insecta Mundi 2009 (66)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Pseudataenius contortus Cartwright 1974" docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="FFD9B60EFFB48269F30DFFDEFFE1FFAC" masterDocTitle="Taxonomic redefinition of the genera Parataenius Balthasar and Pseudataenius Brown, with descriptions of three new species (Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="13" updateTime="1699038167263" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="BF3A7EA72D6209939256CD32C495FD35" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C109A19AD57E1D5FA445AAA108DC1E7">
<mods:title id="0EDFED5C4862B90CF340725EDA296F24">Taxonomic redefinition of the genera Parataenius Balthasar and Pseudataenius Brown, with descriptions of three new species (Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:name id="02F4128CDE9817893F02012B53A0F796" type="personal">
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<mods:namePart id="5643675E7DE2F76345724FDBDD9D5F10">Stebnicka, Z. Teresa</mods:namePart>
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<mods:namePart id="389542C0ED4DFE0D2900938326C669C6">Skelley, Paul E.</mods:namePart>
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<mods:typeOfResource id="DF2A541DD8F6CEFE211781411CD0352C">text</mods:typeOfResource>
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<mods:titleInfo id="708957944DE46012C68415E5BACC4260">
<mods:title id="77F6E2D098C4BC09938E1E75E1666E77">Insecta Mundi</mods:title>
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<mods:part id="ADC53D2580E3FF03F92D1A2382786B56">
<mods:date id="C7ACE09E7BEE2453B7D20B22B1378A4E">2009</mods:date>
<mods:detail id="9285145F844637D1507E2679FFA92949" type="pubDate">
<mods:number id="53F63F1AC542E83EBC1F60205B5E5C34">2009-01-30</mods:number>
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<mods:detail id="EDA3E512C236325B47D47F43C80E23D7" type="volume">
<mods:number id="41649B7BABE8F961442F9E970B6D7DEA">2009</mods:number>
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<mods:number id="45DB8AEB18EDC9CA1205B3E66E9EED68">66</mods:number>
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<mods:start id="7370107E4D10742E83FC5B2DEC697D8B">1</mods:start>
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<mods:classification id="A9F0C5D5EE78915CF0C537C0702B0443">journal article</mods:classification>
<mods:identifier id="7DE0C50A2D17ED38FAC2A1DE83CEEB46" type="DOI">10.5281/zenodo.5404983</mods:identifier>
<mods:identifier id="7AA0FAF54283EA393A71060D38EB5C12" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="D5CAB52E5D79C37E91059BDD240E2831" type="Zenodo-Dep">5404983</mods:identifier>
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<treatment id="03E0CE76FFBA8278F3CDF96DFCC3FF57" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57" httpUri="http://treatment.plazi.org/id/03E0CE76FFBA8278F3CDF96DFCC3FF57" lastPageId="17" lastPageNumber="16" pageId="14" pageNumber="13">
<subSubSection id="C3532CEBFFBA8267F3CDF96DFD4FF965" box="[192,686,1713,1738]" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="8BF67F60FFBA8267F3CDF96DFD4FF965" blockId="14.[192,686,1713,1769]" box="[192,686,1713,1738]" pageId="14" pageNumber="13">
<heading id="D0BEC80CFFBA8267F3CDF96DFD4FF965" bold="true" box="[192,686,1713,1738]" fontSize="10" level="2" pageId="14" pageNumber="13" reason="3">
<taxonomicName id="4C4904E3FFBA8267F3CDF96DFD4FF965" authority="Cartwright" authorityName="Cartwright" authorityYear="1974" box="[192,686,1713,1738]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="contortus">
<emphasis id="B93DA372FFBA8267F3CDF96DFD4FF965" bold="true" box="[192,686,1713,1738]" pageId="14" pageNumber="13">
<emphasis id="B93DA372FFBA8267F3CDF96DFDEEF966" bold="true" box="[192,527,1715,1738]" italics="true" pageId="14" pageNumber="13">Pseudataenius contortus</emphasis>
Cartwright
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFBA8267F3CDF90FFE65F945" box="[192,388,1745,1769]" pageId="14" pageNumber="13" type="description">
<paragraph id="8BF67F60FFBA8267F3CDF90FFE65F945" blockId="14.[192,686,1713,1769]" box="[192,388,1745,1769]" pageId="14" pageNumber="13">
<figureCitation id="137263E5FFBA8267F3CDF90FFEBCF945" box="[192,349,1745,1769]" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="14" pageNumber="13">Figure 24-28</figureCitation>
,
<figureCitation id="137263E5FFBA8267F26BF90FFE65F945" box="[358,388,1745,1769]" captionStart="Figure 34-37" captionStartId="16.[191,275,946,968]" captionTargetBox="[223,1428,257,894]" captionTargetId="figure-371@16.[191,1442,194,965]" captionTargetPageId="16" captionText="Figure 34-37. Pseudataenius spp. 34) P. gracilitarsis, male genitalia. 35) P. socialis, male genitalia. 36) P. contortus, male genitalia. 37) P. socialis, epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5404997" httpUri="https://zenodo.org/record/5404997/files/figure.png" pageId="14" pageNumber="13">36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFBA8267F3CDF8CFFD81F8E5" pageId="14" pageNumber="13" type="reference_group">
<paragraph id="8BF67F60FFBA8267F3CDF8CFFD81F8E5" blockId="14.[192,1440,1809,1865]" pageId="14" pageNumber="13">
<treatmentCitationGroup id="AB59584EFFBA8267F3CDF8CFFD81F8E5" pageId="14" pageNumber="13">
<treatmentCitation id="0AE85971FFBA8267F3CDF8CFFCFFF885" author="Cartwright, O. L." box="[192,798,1809,1833]" page="17" pageId="14" pageNumber="13" year="1974">
<taxonomicName id="4C4904E3FFBA8267F3CDF8CFFCFFF885" authority="Cartwright 1974: 16 - 17" authorityName="Cartwright" authorityPageNumber="16 - 17" authorityYear="1974" box="[192,798,1809,1833]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="contortus">
<emphasis id="B93DA372FFBA8267F3CDF8CFFE11F885" box="[192,496,1809,1833]" italics="true" pageId="14" pageNumber="13">Pseudataenius contortus</emphasis>
<bibRefCitation id="EFD80291FFBA8267F2F5F8CFFCFFF885" author="Cartwright, O. L." box="[504,798,1809,1833]" pageId="14" pageNumber="13" pagination="1 - 106" refId="ref9883" refString="Cartwright, O. L. 1974. Ataenius, Aphotaenius, and Pseudataenius of the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Smithsonian Contributions to Zoology 154: 1 - 106." type="journal article" year="1974">Cartwright 1974: 16-17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1a –
<treatmentCitation id="0AE85971FFBA8267F09AF8CFFB6CF885" author="Woodruff, R. E." box="[919,1165,1809,1833]" page="134" pageId="14" pageNumber="13" year="1973">
<bibRefCitation id="EFD80291FFBA8267F09AF8CFFB6CF885" author="Woodruff, R. E." box="[919,1165,1809,1833]" pageId="14" pageNumber="13" pagination="1 - 220" refId="ref10700" refString="Woodruff, R. E. 1973. The scarab beetles of Florida (Coleoptera: Scarabaeidae). I. The Laparosticti (Subfamilies: Scarabaeinae, Aphodiinae, Hybosorinae, Ochodaeinae, Geotrupinae, Acanthocerinae). Arthropods of Florida and Neighboring Land Areas (Florida Department of Agriculture and Consumer Services, Bureau of Entomology; Gainesville, FL) 8: 1 - 220." type="journal article" year="1973">Woodruff 1973: 134</bibRefCitation>
</treatmentCitation>
, fig. 290 (as unnamed species);
<treatmentCitation id="0AE85971FFBA8267F256F8EFFDBDF8E5" author="Harpootlian, P. J." box="[347,604,1841,1865]" page="36" pageId="14" pageNumber="13" year="2001">
<bibRefCitation id="EFD80291FFBA8267F256F8EFFDBDF8E5" author="Harpootlian, P. J." box="[347,604,1841,1865]" pageId="14" pageNumber="13" refId="ref10028" refString="Harpootlian, P. J. 2001. Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina. Biota of South Carolina, Volume 2. Clemson University Public Service Publishing; Clemson, South Carolina. 157 p." type="book" year="2001">Harpootlian 2001: 36</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF362FE8FFBB8266F3CCFA74FAA3FA70" ID-DOI="http://doi.org/10.5281/zenodo.5404995" ID-Zenodo-Dep="5404995" httpUri="https://zenodo.org/record/5404995/files/figure.png" pageId="15" pageNumber="14" startId="15.[193,277,1450,1472]" targetBox="[197,1443,196,1423]" targetPageId="15">
<paragraph id="8BF67F60FFBB8266F3CCFA74FAA3FA70" blockId="15.[193,1442,1450,1501]" pageId="15" pageNumber="14">
<emphasis id="B93DA372FFBB8266F3CCFA74FE82FA13" bold="true" box="[193,355,1450,1472]" pageId="15" pageNumber="14">Figure 29-33.</emphasis>
<taxonomicName id="4C4904E3FFBB8266F267FA74FD86FA13" baseAuthorityName="Horn" baseAuthorityYear="1871" box="[362,615,1450,1471]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B93DA372FFBB8266F267FA74FD86FA13" box="[362,615,1450,1471]" italics="true" pageId="15" pageNumber="14">Pseudataenius socialis</emphasis>
</taxonomicName>
.
<emphasis id="B93DA372FFBB8266F178FA74FD7BFA6C" bold="true" box="[629,666,1450,1472]" pageId="15" pageNumber="14">29)</emphasis>
Male, dorsal habitus.
<emphasis id="B93DA372FFBB8266F098FA74FC5BFA6C" bold="true" box="[917,954,1450,1472]" pageId="15" pageNumber="14">30)</emphasis>
Male, protibia.
<emphasis id="B93DA372FFBB8266F760FA74FB73FA6C" bold="true" box="[1133,1170,1450,1472]" pageId="15" pageNumber="14">31)</emphasis>
Female, dorsal habitus.
<emphasis id="B93DA372FFBB8266F3CCFA19FF06FA71" bold="true" box="[193,231,1479,1501]" pageId="15" pageNumber="14">32)</emphasis>
Male, anterolateral view, head and pronotum.
<emphasis id="B93DA372FFBB8266F1FAFA19FCFCFA71" bold="true" box="[759,797,1479,1501]" pageId="15" pageNumber="14">33)</emphasis>
Female, anterolateral view, head and pronotum.
</paragraph>
</caption>
<subSubSection id="C3532CEBFFBB8266F3CDFA2AFE05F9E0" pageId="15" pageNumber="14" type="diagnosis">
<paragraph id="8BF67F60FFBB8266F3CDFA2AFE05F9E0" blockId="15.[192,1437,1524,1612]" pageId="15" pageNumber="14">
<emphasis id="B93DA372FFBB8266F3CDFA2AFEA8F9A0" bold="true" box="[192,329,1524,1548]" pageId="15" pageNumber="14">Diagnosis</emphasis>
. Male head minutely punctate; protibia with one lateral, apical tooth, terminal spur long, twisted, widened apically and bent inward; pronotum with smooth, impunctate midline. Female clypeus deeply, closely punctate.
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFBB8279F3CDF9AAFCFBFADD" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14" type="description">
<paragraph id="8BF67F60FFBB8266F3CDF9AAFA7CF8C0" blockId="15.[192,1437,1652,1900]" pageId="15" pageNumber="14">
<emphasis id="B93DA372FFBB8266F3CDF9AAFE85F920" bold="true" box="[192,356,1652,1676]" pageId="15" pageNumber="14">Description</emphasis>
. Length 4.6-5.1 mm. Male head short and broad (
<figureCitation id="137263E5FFBB8266F0CEF9AAFBFAF920" box="[963,1051,1652,1676]" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="15" pageNumber="14">Fig. 24</figureCitation>
), finely reflexed clypeal margin widely rounded on each side of shallow median emargination, sides arcuate to genae, genae right angled, fimbriate and slightly flaring outward; surface of head minutely alutaceous, minutely to very finely punctate (
<figureCitation id="137263E5FFBB8266F234F90AFE6FF940" box="[313,398,1748,1772]" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="15" pageNumber="14">Fig. 27</figureCitation>
). Pronotum weakly convex, sides strongly arcuate from widely rounded anterior angles to arcuate base; sides fimbriate with long setae; surface with mixed very fine and fine, evenly distributed punctures except for narrow, impunctate longitudinal midline. Elytra subparallel sided, humeri not distinctly dentate; striae fine, strial punctures shallow; intervals slightly convex with median row of very minute punctures, interval 10 flat, alutaceous. Ventral surface finely punctate; abdominal sternites flat-
</paragraph>
<paragraph id="8BF67F60FFA48279F241FCB3FE99FC3C" blockId="16.[332,376,877,912]" box="[332,376,877,912]" pageId="16" pageNumber="15">36</paragraph>
<caption id="DF362FE8FFA48279F3B2FC6CFCEDFC49" ID-DOI="http://doi.org/10.5281/zenodo.5404997" ID-Zenodo-Dep="5404997" httpUri="https://zenodo.org/record/5404997/files/figure.png" pageId="16" pageNumber="15" startId="16.[191,275,946,968]" targetBox="[223,1428,257,894]" targetPageId="16">
<paragraph id="8BF67F60FFA48279F3B2FC6CFCEDFC49" blockId="16.[191,1440,946,997]" pageId="16" pageNumber="15">
<emphasis id="B93DA372FFA48279F3B2FC6CFEBCFC6B" bold="true" box="[191,349,946,968]" pageId="16" pageNumber="15">Figure 34-37</emphasis>
.
<taxonomicName id="4C4904E3FFA48279F263FC6DFDABFC64" box="[366,586,947,968]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93DA372FFA48279F263FC6DFDF0FC64" box="[366,529,947,968]" italics="true" pageId="16" pageNumber="15">Pseudataenius</emphasis>
spp.
</taxonomicName>
<emphasis id="B93DA372FFA48279F15EFC6CFD99FC64" bold="true" box="[595,632,946,968]" pageId="16" pageNumber="15">34)</emphasis>
<taxonomicName id="4C4904E3FFA48279F18FFC6DFCCDFC64" authorityName="Stebnicka &amp; Skelley" authorityYear="2009" baseAuthorityName="Stebnicka &amp; Skelley" baseAuthorityYear="1973" box="[642,812,947,968]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gracilitarsis">
<emphasis id="B93DA372FFA48279F18FFC6DFCCDFC64" box="[642,812,947,968]" italics="true" pageId="16" pageNumber="15">P. gracilitarsis</emphasis>
</taxonomicName>
, male genitalia.
<emphasis id="B93DA372FFA48279F0FCFC6CFBF7FC64" bold="true" box="[1009,1046,946,968]" pageId="16" pageNumber="15">35)</emphasis>
<taxonomicName id="4C4904E3FFA48279F72DFC6DFB72FC64" baseAuthorityName="Horn" baseAuthorityYear="1871" box="[1056,1171,947,968]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B93DA372FFA48279F72DFC6DFB72FC64" box="[1056,1171,947,968]" italics="true" pageId="16" pageNumber="15">P. socialis</emphasis>
</taxonomicName>
, male genitalia.
<emphasis id="B93DA372FFA48279F654FC6CFA9EFC64" bold="true" box="[1369,1407,946,968]" pageId="16" pageNumber="15">36)</emphasis>
<taxonomicName id="4C4904E3FFA48279F685FC6DFEC6FC49" authorityName="Cartwright" authorityYear="1974" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="contortus">
<emphasis id="B93DA372FFA48279F685FC6DFEC6FC49" italics="true" pageId="16" pageNumber="15">P. contortus</emphasis>
</taxonomicName>
, male genitalia.
<emphasis id="B93DA372FFA48279F2D2FC11FDE5FC49" bold="true" box="[479,516,975,997]" pageId="16" pageNumber="15">37)</emphasis>
<taxonomicName id="4C4904E3FFA48279F107FC0EFD98FC48" baseAuthorityName="Horn" baseAuthorityYear="1871" box="[522,633,975,997]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B93DA372FFA48279F107FC0EFD98FC48" box="[522,633,975,997]" italics="true" pageId="16" pageNumber="15">P. socialis</emphasis>
</taxonomicName>
, epipharynx.
</paragraph>
</caption>
<paragraph id="8BF67F60FFA48279F3CDFC27FBABFB7D" blockId="16.[192,1440,1017,1393]" pageId="16" pageNumber="15">
tened at middle, finely fluted along sutures, surface minutely alutaceous with scattered punctures more noticeable at sides; apical lip of pygidium convex, shiny, eroded area finely roughened. Legs long; meso- and metafemur without posterior lines; protibia without lateral teeth, terminal tooth elongate, subparallel sided and bluntly rounded, spur long, gradually widened, slightly twisted and bent inward (
<figureCitation id="137263E5FFA48279F634FB87FA73FBDD" box="[1337,1426,1113,1137]" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="16" pageNumber="15">Fig. 25</figureCitation>
); metatibial apex with long setae and accessory spine; tarsi very long and slender, basal tarsomere of metatarsus longer than upper tibial spur and equal in length to following 3 tarsomeres combined, tarsomere 5 as long as 2 preceding tarsomeres combined. Male genitalia as in
<figureCitation id="137263E5FFA48279F0E2FB67FBA7FB7D" box="[1007,1094,1209,1233]" captionStart="Figure 34-37" captionStartId="16.[191,275,946,968]" captionTargetBox="[223,1428,257,894]" captionTargetId="figure-371@16.[191,1442,194,965]" captionTargetPageId="16" captionText="Figure 34-37. Pseudataenius spp. 34) P. gracilitarsis, male genitalia. 35) P. socialis, male genitalia. 36) P. contortus, male genitalia. 37) P. socialis, epipharynx." figureDoi="http://doi.org/10.5281/zenodo.5404997" httpUri="https://zenodo.org/record/5404997/files/figure.png" pageId="16" pageNumber="15">Fig. 36</figureCitation>
.
</paragraph>
<paragraph id="8BF67F60FFA48279F3FDFB07FCFBFADD" blockId="16.[192,1440,1017,1393]" pageId="16" pageNumber="15">
Female color usually darker than in male; head smaller, clypeus more deeply emarginate at middle (
<figureCitation id="137263E5FFA48279F3C4FB27FEFEFABD" box="[201,287,1273,1297]" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="16" pageNumber="15">Fig. 26</figureCitation>
); pronotum more convex, surface of head and pronotum with coarser and closer punctures (
<figureCitation id="137263E5FFA48279F67DFB27FF01FA9D" captionStart="Figure 24-28" captionStartId="14.[194,278,1530,1552]" captionTargetBox="[195,1446,194,1498]" captionTargetId="figure-68@14.[193,1448,192,1500]" captionTargetPageId="14" captionText="Figure 24-28. Pseudataenius contortus. 24) Male, dorsal habitus. 25) Male, protibia. 26) Female, dorsal habitus. 27) Male, anterolateral view, head and pronotum. 28) Female, anterolateral view, head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.5404993" httpUri="https://zenodo.org/record/5404993/files/figure.png" pageId="16" pageNumber="15">Fig. 28</figureCitation>
). Abdominal sternites convex from side to side, pygidium shorter with narrower apical lip. Legs shorter than in male; meso- and metafemora with fine posterior lines; protibia with 3 normal lateral teeth and slender terminal spur; tarsi shorter than in male.
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFA48279F3CDFA47FE23F95D" pageId="16" pageNumber="15" type="materials_examined">
<paragraph id="8BF67F60FFA48279F3CDFA47FDC6FA7D" blockId="16.[192,1440,1433,1489]" pageId="16" pageNumber="15">
<emphasis id="B93DA372FFA48279F3CDFA47FE9CFA1D" bold="true" box="[192,381,1433,1457]" pageId="16" pageNumber="15">Type material</emphasis>
.
<materialsCitation id="3B21753DFFA48279F285FA47FDC6FA7D" collectingDate="1967-06-25" collectingDateMax="1967-08-04" collectingDateMin="1967-06-25" collectionCode="NMNH" collectorName="L. Collins" location="Tall Timbers Res. Sta" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<typeStatus id="54F2C1C2FFA48279F285FA47FE15FA1D" box="[392,500,1433,1457]" pageId="16" pageNumber="15" type="holotype">Holotype</typeStatus>
male “
<collectingRegion id="498DB182FFA48279F149FA47FD7FFA1D" box="[580,670,1433,1457]" country="Uruguay" name="Florida" pageId="16" pageNumber="15">Florida</collectingRegion>
,
<locationDeviation id="9A446642FFA48279F1A5FA47FC57FA1D" box="[680,950,1433,1457]" location="Tallahassee" pageId="16" pageNumber="15">
<quantity id="4CB1D285FFA48279F1A5FA47FD0BFA1D" box="[680,746,1433,1457]" metricMagnitude="4" metricUnit="m" metricValue="2.7358848" pageId="16" pageNumber="15" unit="mi" value="17.0">17 mi</quantity>
N of Tallahassee
</locationDeviation>
,
<location id="8E9629BBFFA48279F0CDFA47FB21FA1D" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F0CDFA47FB21FA1D" box="[960,1216,1433,1457]" name="Tall Timbers Res. Sta" pageId="16" pageNumber="15" stateProvince="Florida">Tall Timbers Res. Sta</location>
<date id="FFF759A0FFA48279F7C8FA47FA7DFA1D" box="[1221,1436,1433,1457]" pageId="16" pageNumber="15" value="1967-06-25" valueMax="1967-08-04" valueMin="1967-06-25">
<collectingDate id="EFB3A048FFA48279F7C8FA47FA7DFA1D" box="[1221,1436,1433,1457]" pageId="16" pageNumber="15" value="1967-06-25" valueMax="1967-08-04" valueMin="1967-06-25">25.VI-4.VIII.1967</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F3CDFA64FED8FA7D" box="[192,313,1465,1489]" pageId="16" pageNumber="15">L. Collins</collectorName>
”, no 71732
<collectionCode id="ED58E7A5FFA48279F2CEFA64FDC3FA7D" box="[451,546,1466,1489]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="15" type="Museum">NMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF67F60FFA48279F3CDFA27FE23F95D" blockId="16.[192,1440,1529,1777]" pageId="16" pageNumber="15">
<emphasis id="B93DA372FFA48279F3CDFA27FE3BF9BD" bold="true" box="[192,474,1529,1553]" pageId="16" pageNumber="15">Specimens examined</emphasis>
.
<materialsCitation id="3B21753DFFA48279F2EAFA27FE23F95D" collectingDate="1948-06-24" collectingDateMax="1987-06-22" collectingDateMin="1948-06-24" collectionCode="FSCA, NMNH, PESC" collectorName="Monticello &amp; Big Bend Hort. Lab. &amp; W. H. Whitcomb &amp; Florida &amp; L. Collins &amp; Tallahassee &amp; Rt. &amp; P. Skelley &amp; Takoma Park &amp; G. H. Nelson &amp; Starkville. &amp; W. H. Cross &amp; Camp Lejeune &amp; J. M. Hirst &amp; Hwy &amp; O. L. Cartwright" country="United States of America" county="Jefferson Co." location="Florida" municipality="Trinity" pageId="16" pageNumber="15" specimenCount="10" stateProvince="Alabama" typeStatus="allotype">
<typeStatus id="54F2C1C2FFA48279F2EAFA27FDAAF9BD" box="[487,587,1529,1553]" pageId="16" pageNumber="15" type="allotype">Allotype</typeStatus>
and
<specimenCount id="9D4FB4E9FFA48279F18FFA27FCFDF9BE" box="[642,796,1529,1554]" pageId="16" pageNumber="15" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
of the same data as holotype and other specimens (88).
<emphasis id="B93DA372FFA48279F3CDF9C7FF1DF99D" bold="true" box="[192,252,1561,1585]" pageId="16" pageNumber="15">
<collectingCountry id="F35E3FF0FFA48279F3CDF9C7FF1DF99D" box="[192,252,1561,1585]" name="United States of America" pageId="16" pageNumber="15">USA</collectingCountry>
</emphasis>
–
<collectingRegion id="498DB182FFA48279F21AF9C7FE6BF99D" box="[279,394,1561,1585]" country="United States of America" name="Alabama" pageId="16" pageNumber="15">Alabama</collectingRegion>
:
<collectingMunicipality id="6B92E51AFFA48279F29EF9C7FE0FF99D" box="[403,494,1561,1585]" pageId="16" pageNumber="15">Trinity</collectingMunicipality>
,
<date id="FFF759A0FFA48279F2F5F9C7FD65F99D" box="[504,644,1561,1585]" pageId="16" pageNumber="15" value="1957-06-17">
<collectingDate id="EFB3A048FFA48279F2F5F9C7FD65F99D" box="[504,644,1561,1585]" pageId="16" pageNumber="15" value="1957-06-17">VI-17-1957</collectingDate>
</date>
, in cow dung (
<collectionCode id="ED58E7A5FFA48279F020F9C7FC9AF99D" box="[813,891,1561,1585]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
);
<location id="8E9629BBFFA48279F080F9C7FC0BF99D" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F080F9C7FC0BF99D" box="[909,1002,1561,1585]" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Florida" pageId="16" pageNumber="15" stateProvince="Alabama">Florida</location>
,
<collectingCounty id="629707ECFFA48279F0F8F9C7FB76F99D" box="[1013,1175,1561,1585]" pageId="16" pageNumber="15">Jefferson Co.</collectingCounty>
,
<collectorName id="26BC1AB6FFA48279F7ACF9C7FAC7F99D" box="[1185,1318,1561,1585]" pageId="16" pageNumber="15">Monticello</collectorName>
,
<collectorName id="26BC1AB6FFA48279F63DF9C7FEA2F9FD" pageId="16" pageNumber="15">Big Bend Hort. Lab.</collectorName>
<date id="FFF759A0FFA48279F241F9E7FE37F9FD" box="[332,470,1593,1617]" pageId="16" pageNumber="15" value="1969-06-12">
<collectingDate id="EFB3A048FFA48279F241F9E7FE37F9FD" box="[332,470,1593,1617]" pageId="16" pageNumber="15" value="1969-06-12">12.VI.1969</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F2E9F9E4FD55F9FD" box="[484,692,1593,1617]" pageId="16" pageNumber="15">W. H. Whitcomb</collectorName>
(
<collectionCode id="ED58E7A5FFA48279F1CBF9E7FCC4F9FD" box="[710,805,1593,1617]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="15" type="Museum">NMNH</collectionCode>
,
<collectionCode id="ED58E7A5FFA48279F03EF9E7FC60F9FD" box="[819,897,1593,1617]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
);
<collectorName id="26BC1AB6FFA48279F09AF9E7FC17F9FD" box="[919,1014,1593,1617]" pageId="16" pageNumber="15">Florida</collectorName>
:
<location id="8E9629BBFFA48279F70FF9E4FB95F9FD" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F70FF9E4FB95F9FD" box="[1026,1140,1593,1617]" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Leon Co." pageId="16" pageNumber="15" stateProvince="Alabama">Leon Co.</location>
,
<location id="8E9629BBFFA48279F78FF9E7FEFDF9DD" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F78FF9E7FEFDF9DD" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Tall Timbers Research Station" pageId="16" pageNumber="15" stateProvince="Alabama">Tall Timbers Research Station</location>
,
<date id="FFF759A0FFA48279F22BF987FE37F9DD" box="[294,470,1625,1649]" pageId="16" pageNumber="15" value="1969-06-21" valueMax="1969-06-27" valueMin="1969-06-21">
<collectingDate id="EFB3A048FFA48279F22BF987FE37F9DD" box="[294,470,1625,1649]" pageId="16" pageNumber="15" value="1969-06-21" valueMax="1969-06-27" valueMin="1969-06-21">21-27-VI-1969</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F2EDF984FDB2F9DD" box="[480,595,1625,1649]" pageId="16" pageNumber="15">L. Collins</collectorName>
(
<collectionCode id="ED58E7A5FFA48279F16DF987FD4CF9DD" box="[608,685,1625,1649]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
);
<collectorName id="26BC1AB6FFA48279F1B2F987FCFDF9DD" box="[703,796,1625,1649]" pageId="16" pageNumber="15">Florida</collectorName>
:
<location id="8E9629BBFFA48279F029F984FC71F9DD" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F029F984FC71F9DD" box="[804,912,1625,1649]" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Leon Co." pageId="16" pageNumber="15" stateProvince="Alabama">Leon Co.</location>
,
<collectorName id="26BC1AB6FFA48279F097F987FBCFF9DD" box="[922,1070,1625,1649]" pageId="16" pageNumber="15">Tallahassee</collectorName>
, jct. I-90 and
<collectorName id="26BC1AB6FFA48279F7C6F984FB11F9DD" box="[1227,1264,1626,1649]" pageId="16" pageNumber="15">Rt.</collectorName>
319,
<date id="FFF759A0FFA48279F620F987FF1EF93D" pageId="16" pageNumber="15" value="1987-06-20" valueMax="1987-06-22" valueMin="1987-06-20">
<collectingDate id="EFB3A048FFA48279F620F987FF1EF93D" pageId="16" pageNumber="15" value="1987-06-20" valueMax="1987-06-22" valueMin="1987-06-20">20-22-VI- 1987</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F204F9A4FE65F93D" box="[265,388,1657,1681]" pageId="16" pageNumber="15">P. Skelley</collectorName>
at light (
<collectionCode id="ED58E7A5FFA48279F2FEF9A7FDA0F93D" box="[499,577,1657,1681]" pageId="16" pageNumber="15">PESC</collectionCode>
);
<collectingRegion id="498DB182FFA48279F15EF9A7FD30F93D" box="[595,721,1657,1681]" country="United States of America" name="Maryland" pageId="16" pageNumber="15">Maryland</collectingRegion>
:
<collectorName id="26BC1AB6FFA48279F1D6F9A7FC64F93D" box="[731,901,1657,1681]" pageId="16" pageNumber="15">Takoma Park</collectorName>
,
<date id="FFF759A0FFA48279F09DF9A7FBC9F93D" box="[912,1064,1657,1681]" pageId="16" pageNumber="15" value="1950-07-13">
<collectingDate id="EFB3A048FFA48279F09DF9A7FBC9F93D" box="[912,1064,1657,1681]" pageId="16" pageNumber="15" value="1950-07-13">13-VII-1950</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F73EF9A7FB33F93D" box="[1075,1234,1657,1681]" pageId="16" pageNumber="15">G. H. Nelson</collectorName>
, at light (
<collectionCode id="ED58E7A5FFA48279F648F9A7FA72F93D" box="[1349,1427,1657,1681]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="16" pageNumber="15" type="Museum">FSCA</collectionCode>
);
<collectingRegion id="498DB182FFA48279F3CDF947FEAAF91D" box="[192,331,1689,1713]" country="United States of America" name="Mississippi" pageId="16" pageNumber="15">Mississippi</collectingRegion>
,
<location id="8E9629BBFFA48279F259F947FDE3F91D" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F259F947FDE3F91D" box="[340,514,1689,1713]" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Oktibbeha Co." pageId="16" pageNumber="15" stateProvince="Alabama">Oktibbeha Co.</location>
,
<collectorName id="26BC1AB6FFA48279F101F947FD6AF91D" box="[524,651,1689,1713]" pageId="16" pageNumber="15">Starkville.</collectorName>
<date id="FFF759A0FFA48279F19DF947FCF7F91D" box="[656,790,1689,1713]" pageId="16" pageNumber="15" value="1982-06-10">
<collectingDate id="EFB3A048FFA48279F19DF947FCF7F91D" box="[656,790,1689,1713]" pageId="16" pageNumber="15" value="1982-06-10">10.VI.1982</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F012F944FC4DF91D" box="[799,940,1689,1713]" pageId="16" pageNumber="15">W. H. Cross</collectorName>
(
<collectionCode id="ED58E7A5FFA48279F0B4F947FBF6F91D" box="[953,1047,1689,1713]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="15" type="Museum">NMNH</collectionCode>
);
<collectingRegion id="498DB182FFA48279F725F947FB02F91D" box="[1064,1251,1689,1713]" country="United States of America" name="North Carolina" pageId="16" pageNumber="15">North Carolina</collectingRegion>
:
<collectorName id="26BC1AB6FFA48279F7E6F947FA7AF91D" box="[1259,1435,1689,1713]" pageId="16" pageNumber="15">Camp Lejeune</collectorName>
,
<date id="FFF759A0FFA48279F3CDF967FEACF97D" box="[192,333,1721,1745]" pageId="16" pageNumber="15" value="1968-06-28">
<collectingDate id="EFB3A048FFA48279F3CDF967FEACF97D" box="[192,333,1721,1745]" pageId="16" pageNumber="15" value="1968-06-28">28-VI-1968</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F255F964FE3EF97D" box="[344,479,1721,1745]" pageId="16" pageNumber="15">J. M. Hirst</collectorName>
,
<collectingMethod id="52080777FFA48279F2E7F967FD45F97D" box="[490,676,1721,1745]" pageId="16" pageNumber="15">blacklight trap</collectingMethod>
;
<collectingRegion id="498DB182FFA48279F1A2F967FC8FF97D" box="[687,878,1721,1745]" country="United States of America" name="South Carolina" pageId="16" pageNumber="15">South Carolina</collectingRegion>
,
<location id="8E9629BBFFA48279F074F967FBF0F97D" LSID="urn:lsid:plazi:treatment:03E0CE76FFBA8278F3CDF96DFCC3FF57:8E9629BBFFA48279F074F967FBF0F97D" box="[889,1041,1721,1745]" country="United States of America" county="Jefferson Co." municipality="Trinity" name="Colleton Co." pageId="16" pageNumber="15" stateProvince="Alabama">Colleton Co.</location>
,
<collectorName id="26BC1AB6FFA48279F711F964FBB7F97E" box="[1052,1110,1722,1746]" pageId="16" pageNumber="15">Hwy</collectorName>
32 &amp; 320,
<date id="FFF759A0FFA48279F7DBF967FABEF97D" box="[1238,1375,1721,1745]" pageId="16" pageNumber="15" value="1948-06-24">
<collectingDate id="EFB3A048FFA48279F7DBF967FABEF97D" box="[1238,1375,1721,1745]" pageId="16" pageNumber="15" value="1948-06-24">24.VI.1948</collectingDate>
</date>
,
<collectorName id="26BC1AB6FFA48279F667F967FEA8F95D" pageId="16" pageNumber="15">O.L. Cartwright</collectorName>
(
<collectionCode id="ED58E7A5FFA48279F25AF907FE56F95D" box="[343,439,1753,1777]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="15" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFA48279F3CDF8C7FBA2F8DD" pageId="16" pageNumber="15" type="discussion">
<paragraph id="8BF67F60FFA48279F3CDF8C7FBA2F8DD" blockId="16.[192,1440,1817,1905]" pageId="16" pageNumber="15">
<emphasis id="B93DA372FFA48279F3CDF8C7FEA4F89D" bold="true" box="[192,325,1817,1841]" pageId="16" pageNumber="15">Remarks.</emphasis>
<taxonomicName id="4C4904E3FFA48279F243F8C7FD61F89D" authorityName="Cartwright" authorityYear="1974" box="[334,640,1817,1841]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="contortus">
<emphasis id="B93DA372FFA48279F243F8C7FD61F89D" box="[334,640,1817,1841]" italics="true" pageId="16" pageNumber="15">Pseudataenius contortus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C4904E3FFA48279F061F8C4FC0DF89D" baseAuthorityName="Horn" baseAuthorityYear="1871" box="[876,1004,1817,1841]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B93DA372FFA48279F061F8C4FC0DF89D" box="[876,1004,1817,1841]" italics="true" pageId="16" pageNumber="15">P. socialis</emphasis>
</taxonomicName>
. The male of this species is easily recognized by its unusual protibial shape, while the female has the clypeal wrinkles inconspicuous or lacking and the pronotal surface less strongly punctate than in
<taxonomicName id="4C4904E3FFA48279F0CFF884FBDCF8DD" baseAuthorityName="Horn" baseAuthorityYear="1871" box="[962,1085,1881,1905]" class="Insecta" family="Aphodiidae" genus="Pseudataenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="socialis">
<emphasis id="B93DA372FFA48279F0CFF884FBDCF8DD" box="[962,1085,1881,1905]" italics="true" pageId="16" pageNumber="15">P. socialis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3532CEBFFA58278F3CDFF1DFCC3FF57" pageId="17" pageNumber="16" type="distribution">
<paragraph id="8BF67F60FFA58278F3CDFF1DFCC3FF57" blockId="17.[192,1440,195,252]" pageId="17" pageNumber="16">
<emphasis id="B93DA372FFA58278F3CDFF1DFE8CFF77" bold="true" box="[192,365,195,219]" pageId="17" pageNumber="16">Distribution</emphasis>
. Eastern and southeastern
<collectingCountry id="F35E3FF0FFA58278F1C7FF1DFCE5FF77" box="[714,772,195,219]" name="United States of America" pageId="17" pageNumber="16">USA</collectingCountry>
–
<collectingRegion id="498DB182FFA58278F02EFF1DFC77FF77" box="[803,918,195,219]" country="United States of America" name="Alabama" pageId="17" pageNumber="16">Alabama</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F0AFFF1DFBFDFF77" box="[930,1052,195,219]" country="United States of America" name="Delaware" pageId="17" pageNumber="16">Delaware</collectingRegion>
, Florida,
<collectingRegion id="498DB182FFA58278F79EFF1DFAF3FF77" box="[1171,1298,195,219]" country="United States of America" name="Louisiana" pageId="17" pageNumber="16">Louisiana</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F613FF1DFA7AFF77" box="[1310,1435,195,219]" country="United States of America" name="Maryland" pageId="17" pageNumber="16">Maryland</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F3CDFF3DFEACFF57" box="[192,333,227,251]" country="United States of America" name="Mississippi" pageId="17" pageNumber="16">Mississippi</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F25AFF3AFE07FF50" box="[343,486,228,252]" country="United States of America" name="New Jersey" pageId="17" pageNumber="16">New Jersey</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F2FDFF3DFD4AFF57" box="[496,683,227,251]" country="United States of America" name="South Carolina" pageId="17" pageNumber="16">South Carolina</collectingRegion>
,
<collectingRegion id="498DB182FFA58278F1B8FF3DFCFFFF57" box="[693,798,227,251]" country="United States of America" name="Virginia" pageId="17" pageNumber="16">Virginia</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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