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<mods:title id="AB9AE3D60DDE5174BD23553F2DB071D7">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<emphasis id="B907BE5B693DEF13FE55AA6DFD43200D" bold="true" box="[478,651,1363,1390]" italics="true" pageId="27" pageNumber="28">Amphidromus</emphasis>
(
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) 
<emphasis id="B907BE5B693DEF13FCBDAA6DFC7D200E" bold="true" box="[822,949,1363,1389]" italics="true" pageId="27" pageNumber="28">xiengensis</emphasis>
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Morlet, 1891
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<figureCitation id="13487ECC693DEF13FD8EAA49FD8620F2" box="[517,590,1399,1425]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="27" pageNumber="28">Figs 1</figureCitation>
, 
<figureCitation id="13487ECC693DEF13FDD7AA49FDA220F2" box="[604,618,1399,1425]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="27" pageNumber="28">9</figureCitation>
C–D, 12D–F, 13A–H, 14A–B; 
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<bibRefCitation id="EFE21FB8693DEF13FE6DAA87FD7220B0" author="Morlet L." box="[486,698,1465,1492]" pageId="27" pageNumber="28" pagination="25 - 28" refId="ref19021" refString="Morlet L. 1891 a. Diagnoses molluscorum novorum, in Indochina collectorum. Journal de Conchyliologie 39: 25 - 28. Available from http: // biodiversitylibrary. org / page / 15927626 [accessed 9 Sept. 2013]." type="journal article" year="1891">Morlet, 1891a: 27</bibRefCitation>
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. 
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locality: “Xieng-Mai et les forêts des bords du Ménam
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Pinh, 
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occidental” (banks of Ping River, 
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, 
<collectingCountry id="F36422D9693DEF13FC64AAE3FB95209B" box="[1007,1117,1501,1528]" name="Thailand" pageId="27" pageNumber="28">Thailand</collectingCountry>
). 
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<emphasis id="B907BE5B693DEF13FF36A93EFE2A2379" box="[189,482,1536,1562]" italics="true" pageId="27" pageNumber="28">Amphidromus contrarius</emphasis>
var. 
<emphasis id="B907BE5B693DEF13FD97A93EFD7D2379" box="[540,693,1536,1562]" italics="true" pageId="27" pageNumber="28">multifasciata</emphasis>
<bibRefCitation id="EFE21FB8693DEF13FD37A93EFC4A2378" author="Fulton H. C." box="[700,898,1536,1563]" pageId="27" pageNumber="28" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896: 78</bibRefCitation>
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, pl. 7, fig. 5. 
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locality: 
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. 
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<emphasis id="B907BE5B693DEF13FF36A91AFE2C235D" box="[189,484,1572,1598]" italics="true" pageId="27" pageNumber="28">Amphidromus xiengensis</emphasis>
var. 
<emphasis id="B907BE5B693DEF13FDA8A91AFDB3235D" box="[547,635,1572,1598]" italics="true" pageId="27" pageNumber="28">clausus</emphasis>
<bibRefCitation id="EFE21FB8693DEF13FD0FA91AFC54235C" author="Pilsbry H. A." box="[644,924,1572,1599]" pageId="27" pageNumber="28" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900: 195–196</bibRefCitation>
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, pl. 63, figs 79−82. 
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locality: 
<collectingCountry id="F36422D9693DEF13FAB6A91BFABE235C" box="[1341,1398,1573,1599]" name="Laos" pageId="27" pageNumber="28">Laos</collectingCountry>
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Mountain, 
<collectingCountry id="F36422D9693DEF13FEDFA976FE192301" box="[340,465,1608,1634]" name="Cambodia" pageId="27" pageNumber="28">Cambodia</collectingCountry>
.
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<taxonomicName id="4C7319CA693DEF13FF36A9B4FD6D23C7" ID-CoL="D2PK" box="[189,677,1674,1700]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="variety" species="porcellanus" variety="xiengensis">
<emphasis id="B907BE5B693DEF13FF36A9B4FE3923C7" box="[189,497,1674,1700]" italics="true" pageId="27" pageNumber="28">Amphidromus porcellanus</emphasis>
var. 
<emphasis id="B907BE5B693DEF13FDA0A9B5FD6D23C7" box="[555,677,1675,1700]" italics="true" pageId="27" pageNumber="28">xiengensis</emphasis>
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– 
<treatmentCitation id="0AD24458693DEF13FD48A9B4FC4A23C7" author="Fulton H. C." box="[707,898,1674,1700]" page="79" pageId="27" pageNumber="28" year="1896">
<bibRefCitation id="EFE21FB8693DEF13FD48A9B4FC4A23C7" author="Fulton H. C." box="[707,898,1674,1700]" pageId="27" pageNumber="28" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton 1896: 79</bibRefCitation>
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.
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<emphasis id="B907BE5B693DEF13FF36A993FE2B23A4" box="[189,483,1709,1735]" italics="true" pageId="27" pageNumber="28">Amphidromus xiengensis</emphasis>
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– 
<treatmentCitation id="0AD24458693DEF13FD88A993FD2D23A4" author="Morlet L." box="[515,741,1709,1735]" page="232" pageId="27" pageNumber="28" year="1891">
<bibRefCitation id="EFE21FB8693DEF13FD88A993FD2D23A4" author="Morlet L." box="[515,741,1709,1735]" pageId="27" pageNumber="28" pagination="230 - 254" refId="ref19064" refString="Morlet L. 1891 b. Contribution a la faune malacologique de lIndo-Chine. Journal de Conchyliologie 39: 230 - 254. Available from http: // biodiversitylibrary. org / page / 15927831 [accessed 9 Sept. 2013]." type="journal article" year="1891">Morlet 1891b: 232</bibRefCitation>
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, 240–241, pl. 5, fig. 4. — 
<treatmentCitation id="0AD24458693DEF13FBABA990FB3823A4" author="Ancey C. F." box="[1056,1264,1709,1736]" page="134" pageId="27" pageNumber="28" year="1898">
<bibRefCitation id="EFE21FB8693DEF13FBABA990FB3823A4" author="Ancey C. F." box="[1056,1264,1709,1736]" pageId="27" pageNumber="28" pagination="129 - 137" refId="ref17973" refString="Ancey C. F. 1898. Observations sur les mollusques terrestres et fluviatiles recueillis dans lIndo-chine et particulierement au Laos par M. H. Counillon. Bulletin du Museum d'Histoire naturelle de Marseille 1: 129 - 137." type="journal article" year="1898">Ancey 1898: 134</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD24458693DEF13FAA8A993FE43238F" author="Pilsbry H. A." page="194" pageId="27" pageNumber="28" year="1900">
<bibRefCitation id="EFE21FB8693DEF13FAA8A993FE43238F" author="Pilsbry H. A." pageId="27" pageNumber="28" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 194–195</bibRefCitation>
</treatmentCitation>
, pl. 63, figs 75–76. — 
<treatmentCitation id="0AD24458693DEF13FD20A9EFFC1D238F" author="Fischer-Piette E." box="[683,981,1745,1772]" page="159" pageId="27" pageNumber="28" year="1950">
<bibRefCitation id="EFE21FB8693DEF13FD20A9EFFC1D238F" author="Fischer-Piette E." box="[683,981,1745,1772]" pageId="27" pageNumber="28" pagination="149 - 180" refId="ref18386" refString="Fischer-Piette E. 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal. Journal de Conchyliologie 90: 149 - 180." type="journal article" year="1950">Fischer-Piette 1950: 159</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD24458693DEF13FB87A9EFFECC226C" author="Laidlaw F. F. &amp; Solem A." page="564" pageId="27" pageNumber="28" year="1961">
<bibRefCitation id="EFE21FB8693DEF13FB87A9EFFECC226C" author="Laidlaw F. F. &amp; Solem A." pageId="27" pageNumber="28" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 564– 565</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD24458693DEF13FEB8A9CBFDF1226C" author="Solem A." box="[307,569,1780,1807]" page="626" pageId="27" pageNumber="28" year="1965">
<bibRefCitation id="EFE21FB8693DEF13FEB8A9CBFDF1226C" author="Solem A." box="[307,569,1780,1807]" pageId="27" pageNumber="28" pagination="615 - 628" refId="ref19790" refString="Solem A. 1965. Land snails of the genus Amphidromus from Thailand (Mollusca: Pumonata: Camaenidae). Proceedings of the United States National Museum 117: 615 - 628. Available from http: // biodiversitylibrary. org / page / 7759057 [accessed 8 Jun. 2016]." type="journal article" year="1965">Solem 1965: 626–627</bibRefCitation>
</treatmentCitation>
, pl. 2, figs 7-13.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCC6249693DEF13FF36A826FBE42250" blockId="27.[189,1398,1674,1879]" box="[189,1068,1816,1843]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB634567693DEF13FF36A826FBE42250" box="[189,1068,1816,1843]" pageId="27" pageNumber="28">
<taxonomicName id="4C7319CA693DEF13FF36A826FDBC2251" ID-CoL="5M72W" authorityName="Pilsbry" authorityYear="1900" box="[189,628,1816,1843]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="variety" species="xiengensis" variety="clausus">
<emphasis id="B907BE5B693DEF13FF36A826FE2A2251" box="[189,482,1816,1842]" italics="true" pageId="27" pageNumber="28">Amphidromus xiengensis</emphasis>
var. 
<emphasis id="B907BE5B693DEF13FD97A826FDBC2251" box="[540,628,1816,1842]" italics="true" pageId="27" pageNumber="28">clausus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AD24458693DEF13FD1AA826FC952251" author="Solem A." box="[657,861,1816,1843]" page="626" pageId="27" pageNumber="28" year="1965">
<bibRefCitation id="EFE21FB8693DEF13FD1AA826FC952251" author="Solem A." box="[657,861,1816,1843]" pageId="27" pageNumber="28" pagination="615 - 628" refId="ref19790" refString="Solem A. 1965. Land snails of the genus Amphidromus from Thailand (Mollusca: Pumonata: Camaenidae). Proceedings of the United States National Museum 117: 615 - 628. Available from http: // biodiversitylibrary. org / page / 7759057 [accessed 8 Jun. 2016]." type="journal article" year="1965">Solem 1965: 626</bibRefCitation>
</treatmentCitation>
, pl. 2, figs 11–13.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCC6249693DEF13FF36A802FA952235" blockId="27.[189,1398,1674,1879]" box="[189,1373,1852,1879]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB634567693DEF13FF36A802FA952235" box="[189,1373,1852,1879]" pageId="27" pageNumber="28">
<taxonomicName id="4C7319CA693DEF13FF36A802FD092235" box="[189,705,1852,1879]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="variety" species="xiengensis" variety="multifasciatus">
<emphasis id="B907BE5B693DEF13FF36A802FE2A2235" box="[189,482,1852,1878]" italics="true" pageId="27" pageNumber="28">Amphidromus xiengensis</emphasis>
var. 
<emphasis id="B907BE5B693DEF13FD97A802FD092235" box="[540,705,1852,1878]" italics="true" pageId="27" pageNumber="28">multifasciatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AD24458693DEF13FD55A802FC782234" author="Pilsbry H. A." box="[734,944,1852,1879]" page="195" pageId="27" pageNumber="28" year="1900">
<bibRefCitation id="EFE21FB8693DEF13FD55A802FC782234" author="Pilsbry H. A." box="[734,944,1852,1879]" pageId="27" pageNumber="28" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 195</bibRefCitation>
</treatmentCitation>
, pl. 63, fig. 77. — 
<treatmentCitation id="0AD24458693DEF13FB06A802FA912235" author="Solem A." box="[1165,1369,1852,1879]" page="626" pageId="27" pageNumber="28" year="1965">
<bibRefCitation id="EFE21FB8693DEF13FB06A802FA912235" author="Solem A." box="[1165,1369,1852,1879]" pageId="27" pageNumber="28" pagination="615 - 628" refId="ref19790" refString="Solem A. 1965. Land snails of the genus Amphidromus from Thailand (Mollusca: Pumonata: Camaenidae). Proceedings of the United States National Museum 117: 615 - 628. Available from http: // biodiversitylibrary. org / page / 7759057 [accessed 8 Jun. 2016]." type="journal article" year="1965">Solem 1965: 626</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36931C2693DEF16FF36A840FB3421B4" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BCC6249693DEF13FF36A840FE6222FB" blockId="27.[189,426,1918,1944]" box="[189,426,1918,1944]" pageId="27" pageNumber="28">
<heading id="D084D525693DEF13FF36A840FE6222FB" bold="true" box="[189,426,1918,1944]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B907BE5B693DEF13FF36A840FE6222FB" bold="true" box="[189,426,1918,1944]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249693DEF16FF36A893FB8B24F0" blockId="27.[189,1398,1965,2028]" lastBlockId="30.[189,1399,268,403]" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
The species was described based on material from Muséum (
<collectionCode id="ED62FA8C693DEF13FC1FA893FC3922A4" box="[916,1009,1965,1991]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
) ex. A. Pavie Collection and did not include an illustration. 
<bibRefCitation id="EFE21FB8693DEF13FD8BA8EFFD7F228F" author="Morlet L." box="[512,695,2001,2028]" pageId="27" pageNumber="28" pagination="230 - 254" refId="ref19064" refString="Morlet L. 1891 b. Contribution a la faune malacologique de lIndo-Chine. Journal de Conchyliologie 39: 230 - 254. Available from http: // biodiversitylibrary. org / page / 15927831 [accessed 9 Sept. 2013]." type="journal article" year="1891">Morlet (1891b)</bibRefCitation>
subsequently re-published the description and illustrated a single specimen. There is a single specimen of L. Morlet in the 
<collectionCode id="ED62FA8C6938EF16FC29AE33FC342444" box="[930,1020,269,295]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
collection with an original label stating “Type”. 
<bibRefCitation id="EFE21FB86938EF16FEF1AE0EFD7A2428" author="Fischer-Piette E." box="[378,690,304,331]" pageId="30" pageNumber="31" pagination="149 - 180" refId="ref18386" refString="Fischer-Piette E. 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal. Journal de Conchyliologie 90: 149 - 180." type="journal article" year="1950">Fischer-Piette (1950: 159)</bibRefCitation>
wrote the “ 
<typeStatus id="54C8DCEB6938EF16FCB4AE0EFC632429" box="[831,939,304,330]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
, 
<quantity id="4C8BCFAC6938EF16FC32AE0EFBDB2429" box="[953,1043,304,330]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="30" pageNumber="31" unit="mm" value="38.0">38 mm</quantity>
”, which we consider to be an inadvertent 
<typeStatus id="54C8DCEB6938EF16FEC2AE6AFE7D240D" box="[329,437,340,366]" pageId="30" pageNumber="31" type="lectotype">lectotype</typeStatus>
designation (
<collectionCode id="ED62FA8C6938EF16FDD8AE6AFD51240D" box="[595,665,340,366]" pageId="30" pageNumber="31">ICZN</collectionCode>
1999: Art. 74.5). Therefore, 
<typeStatus id="54C8DCEB6938EF16FC65AE6AFB92240D" box="[1006,1114,340,366]" pageId="30" pageNumber="31" type="lectotype">lectotype</typeStatus>
MNHN-IM-130306224 (
<figureCitation id="13487ECC6938EF16FF4DAE46FEEA24F1" box="[198,290,376,402]" captionStart="Fig" captionStartId="17.[189,232,1719,1745]" captionTargetBox="[239,1349,265,1695]" captionTargetId="figure@17.[232,1356,265,1696]" captionTargetPageId="17" captionText="Fig. 8. Genitalia, mating pairs, shell banding formula and radula morphology. A–B. Amphidromus (Syndromus) contrarius (Müller, 1774) from Timor (ZMA). A. Genital system. B. Internal structures of penis and vagina. C. Mating pair of Amphidromus (Syndromus) sp., showing the protruded vaginal simulator pilaster (vsp, red arrows) and its possible function as a stimulating organ. D. Schematic drawing of shell banding, where the numbers 1 to 6 on the last whorl indicate the position of each band. E–G. Radula morphology of Amphidromus (Syndromus) contrarius (Müller, 1774) from Timor (ZMA). E. Central tooth, with the first to fifth lateral teeth. F. Lateral teeth with the tricuspid marginal teeth transition. G. Outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829760" httpUri="https://zenodo.org/record/3829760/files/figure.png" pageId="30" pageNumber="31">Fig. 8A</figureCitation>
) was designated by 
<bibRefCitation id="EFE21FB86938EF16FD9BAE46FC8C24F0" author="Fischer-Piette E." box="[528,836,376,403]" pageId="30" pageNumber="31" pagination="149 - 180" refId="ref18386" refString="Fischer-Piette E. 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal. Journal de Conchyliologie 90: 149 - 180." type="journal article" year="1950">Fischer-Piette (1950: 153)</bibRefCitation>
to stabilise the name.
</paragraph>
<caption id="DF0C32C1693AEF14FF36A973FB4B228F" ID-DOI="http://doi.org/10.5281/zenodo.3829770" ID-Zenodo-Dep="3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1613,1639]" targetBox="[263,1324,265,1582]" targetPageId="28">
<paragraph id="8BCC6249693AEF14FF36A973FB4B228F" blockId="28.[189,1399,1613,2028]" pageId="28" pageNumber="29">
<emphasis id="B907BE5B693AEF14FF36A973FEE92304" bold="true" box="[189,289,1613,1639]" pageId="28" pageNumber="29">Fig. 13.</emphasis>
Shells of 
<taxonomicName id="4C7319CA693AEF14FE25A973FD3D230B" authority="" authorityName="Pilsbry" authorityYear="1900" box="[430,757,1613,1640]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
<emphasis id="B907BE5B693AEF14FE25A973FD992304" box="[430,593,1613,1639]" italics="true" pageId="28" pageNumber="29">Amphidromus</emphasis>
(
<emphasis id="B907BE5B693AEF14FDE1A973FD232304" box="[618,747,1613,1639]" italics="true" pageId="28" pageNumber="29">Syndromus</emphasis>
)
</taxonomicName>
spp. 
<emphasis id="B907BE5B693AEF14FCC9A973FCB52304" bold="true" box="[834,893,1613,1639]" pageId="28" pageNumber="29">A−C</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FC18A973FABB230B" authority="Morlet, 1891" authorityName="Morlet" authorityYear="1891" box="[915,1395,1613,1640]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B693AEF14FC18A973FB772304" box="[915,1215,1613,1639]" italics="true" pageId="28" pageNumber="29">Amphidromus xiengensis</emphasis>
Morlet, 1891
</taxonomicName>
. 
<emphasis id="B907BE5B693AEF14FF36A94EFF1A23E9" bold="true" box="[189,210,1648,1674]" pageId="28" pageNumber="29">A</emphasis>
. Lectotype (MNHN-IM-2000-5249). 
<emphasis id="B907BE5B693AEF14FD19A94FFD6E23E8" bold="true" box="[658,678,1649,1675]" pageId="28" pageNumber="29">B</emphasis>
. Specimen from Thailand (CUMZ 7050). 
<emphasis id="B907BE5B693AEF14FB16A94EFB7A23E9" bold="true" box="[1181,1202,1648,1674]" pageId="28" pageNumber="29">C</emphasis>
. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). 
<emphasis id="B907BE5B693AEF14FCD3A9AAFC5823CD" bold="true" box="[856,912,1684,1710]" pageId="28" pageNumber="29">D−F</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FC2BA9AAFB0F23CD" authorityName="Morlet" authorityYear="1891" box="[928,1223,1684,1710]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B693AEF14FC2BA9AAFB0F23CD" box="[928,1223,1684,1710]" italics="true" pageId="28" pageNumber="29">Amphidromus xiengensis</emphasis>
</taxonomicName>
var. “ 
<emphasis id="B907BE5B693AEF14FA99A9AAFAA223CD" box="[1298,1386,1684,1710]" italics="true" pageId="28" pageNumber="29">clausus</emphasis>
” 
<bibRefCitation id="EFE21FB8693AEF14FF36A989FE9423B2" author="Pilsbry H. A." box="[189,348,1719,1745]" pageId="28" pageNumber="29" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
. 
<emphasis id="B907BE5B693AEF14FEE2A989FEB623B2" bold="true" box="[361,382,1719,1745]" pageId="28" pageNumber="29">D</emphasis>
. Lectotype (ANSP 31496). 
<emphasis id="B907BE5B693AEF14FD47A989FD2823B2" bold="true" box="[716,736,1719,1745]" pageId="28" pageNumber="29">E</emphasis>
. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). 
<emphasis id="B907BE5B693AEF14FEF5A9E5FE582396" bold="true" box="[382,400,1755,1781]" pageId="28" pageNumber="29">F</emphasis>
. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). 
<emphasis id="B907BE5B693AEF14FB0FA9E4FB082397" bold="true" box="[1156,1216,1754,1780]" pageId="28" pageNumber="29">G–H</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FB58A9E4FEFF2274" authorityName="Morlet" authorityYear="1891" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B693AEF14FB58A9E4FEFF2274" italics="true" pageId="28" pageNumber="29">Amphidromus xiengensis</emphasis>
</taxonomicName>
var. “ 
<emphasis id="B907BE5B693AEF14FE0BA9C3FDD12274" box="[384,537,1789,1815]" italics="true" pageId="28" pageNumber="29">multifasciata</emphasis>
” 
<bibRefCitation id="EFE21FB8693AEF14FDA4A9C3FD0E2274" author="Fulton H. C." box="[559,710,1789,1815]" pageId="28" pageNumber="29" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). 
<emphasis id="B907BE5B693AEF14FE9AA81FFE812258" bold="true" box="[273,329,1825,1851]" pageId="28" pageNumber="29">I−M</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FED2A81FFCA52258" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[345,877,1825,1851]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fuscolabris">
<emphasis id="B907BE5B693AEF14FED2A81FFD4F2258" box="[345,647,1825,1851]" italics="true" pageId="28" pageNumber="29">Amphidromus fuscolabris</emphasis>
<bibRefCitation id="EFE21FB8693AEF14FD1BA81FFCA52258" author="Mollendorff O. F. von" box="[656,877,1825,1851]" pageId="28" pageNumber="29" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B907BE5B693AEF14FCF1A81FFC4D2258" bold="true" box="[890,901,1825,1851]" pageId="28" pageNumber="29">I</emphasis>
. Holotype (SMF 7641). 
<emphasis id="B907BE5B693AEF14FB27A81FFB2F2258" bold="true" box="[1196,1255,1825,1851]" pageId="28" pageNumber="29">J−M</emphasis>
. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). 
<emphasis id="B907BE5B693AEF14FCAAA87AFCFE223D" bold="true" box="[801,822,1860,1886]" pageId="28" pageNumber="29">N</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FCCFA87AFB19223D" authority="Ancey, 1897" authorityName="Ancey" authorityYear="1897" box="[836,1233,1860,1886]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="eudeli">
<emphasis id="B907BE5B693AEF14FCCFA87AFBFD223D" box="[836,1077,1860,1886]" italics="true" pageId="28" pageNumber="29">Amphidromus eudeli</emphasis>
Ancey, 1897
</taxonomicName>
, syntype from Binh Dinh, Annam (RBINS 617427). 
<emphasis id="B907BE5B693AEF14FDF0A859FD7A22E2" bold="true" box="[635,690,1895,1921]" pageId="28" pageNumber="29">O–P</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FD4AA859FB3122E2" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[705,1273,1895,1921]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="haematostoma">
<emphasis id="B907BE5B693AEF14FD4AA859FBDF22E2" box="[705,1047,1895,1921]" italics="true" pageId="28" pageNumber="29">Amphidromus haematostoma</emphasis>
<bibRefCitation id="EFE21FB8693AEF14FB95A859FB3122E2" author="Mollendorff O. F. von" box="[1054,1273,1895,1921]" pageId="28" pageNumber="29" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B907BE5B693AEF14FA8FA859FAD322E2" bold="true" box="[1284,1307,1895,1921]" pageId="28" pageNumber="29">O</emphasis>
. Lectotype (SMF 7559). 
<emphasis id="B907BE5B693AEF14FE28A8B5FE7D22C6" bold="true" box="[419,437,1931,1957]" pageId="28" pageNumber="29">P</emphasis>
. Paralectotype (SMF 7560). 
<emphasis id="B907BE5B693AEF14FC91A8B4FC9D22C7" bold="true" box="[794,853,1930,1956]" pageId="28" pageNumber="29">Q–R</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FCE3A8B4FB0A22C7" authorityName="Mollendorff" authorityYear="1898" box="[872,1218,1930,1956]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="haematostoma">
<emphasis id="B907BE5B693AEF14FCE3A8B4FB0A22C7" box="[872,1218,1930,1956]" italics="true" pageId="28" pageNumber="29">Amphidromus haematostoma</emphasis>
</taxonomicName>
var. “ 
<emphasis id="B907BE5B693AEF14FA99A8B5FAA222C7" box="[1298,1386,1931,1956]" italics="true" pageId="28" pageNumber="29">varians</emphasis>
” 
<bibRefCitation id="EFE21FB8693AEF14FF36A893FE5622AB" author="Mollendorff O. F. von" box="[189,414,1965,1992]" pageId="28" pageNumber="29" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
. 
<emphasis id="B907BE5B693AEF14FE3BA893FE0F22A4" bold="true" box="[432,455,1965,1991]" pageId="28" pageNumber="29">Q</emphasis>
. Lectotype (SMF 7561). 
<emphasis id="B907BE5B693AEF14FC81A890FCD722AB" bold="true" box="[778,799,1966,1992]" pageId="28" pageNumber="29">R</emphasis>
. Paralectotype (SMF 7562). 
<emphasis id="B907BE5B693AEF14FB07A893FB7622A4" bold="true" box="[1164,1214,1965,1991]" pageId="28" pageNumber="29">S–T</emphasis>
. 
<taxonomicName id="4C7319CA693AEF14FB58A893FE452288" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="xiengkhaungensis" status="sp. nov.">
<emphasis id="B907BE5B693AEF14FB58A893FE452288" italics="true" pageId="28" pageNumber="29">Amphidromus xiengkhaungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340320693AEF14FE1FA8EFFE272288" box="[404,495,2001,2027]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B907BE5B693AEF14FE7DA8EEFDCE2289" bold="true" box="[502,518,2000,2026]" pageId="28" pageNumber="29">S</emphasis>
. Holotype (CUMZ 7045). 
<emphasis id="B907BE5B693AEF14FCCAA8EFFC9D2288" bold="true" box="[833,853,2001,2027]" pageId="28" pageNumber="29">T</emphasis>
. Paratype (CUMZ 7046).
</paragraph>
</caption>
<caption id="DF0C32C1693BEF15FF36A87CFD76228A" ID-DOI="http://doi.org/10.5281/zenodo.3829772" ID-Zenodo-Dep="3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1858,1884]" targetBox="[189,1399,310,1811]" targetPageId="29">
<paragraph id="8BCC6249693BEF15FF36A87CFD76228A" blockId="29.[189,1399,1858,2025]" pageId="29" pageNumber="30">
<emphasis id="B907BE5B693BEF15FF36A87CFED7223F" bold="true" box="[189,287,1858,1884]" pageId="29" pageNumber="30">Fig. 14.</emphasis>
Genitalia of 
<taxonomicName id="4C7319CA693BEF15FE43A87CFCC5223F" authority="" authorityName="Pilsbry" authorityYear="1900" box="[456,781,1858,1884]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
<emphasis id="B907BE5B693BEF15FE43A87CFDA4223F" box="[456,620,1858,1884]" italics="true" pageId="29" pageNumber="30">Amphidromus</emphasis>
(
<emphasis id="B907BE5B693BEF15FD09A87CFCCB223F" box="[642,771,1858,1884]" italics="true" pageId="29" pageNumber="30">Syndromus</emphasis>
)
</taxonomicName>
spp. 
<emphasis id="B907BE5B693BEF15FCDDA87CFC45223F" bold="true" box="[854,909,1858,1884]" pageId="29" pageNumber="30">A–B</emphasis>
. 
<taxonomicName id="4C7319CA693BEF15FC2AA87CFABF223F" authority="Morlet, 1891" authorityName="Morlet" authorityYear="1891" box="[929,1399,1858,1884]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B693BEF15FC2AA87CFB03223F" box="[929,1227,1858,1884]" italics="true" pageId="29" pageNumber="30">Amphidromus xiengensis</emphasis>
Morlet, 1891
</taxonomicName>
from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). 
<emphasis id="B907BE5B693BEF15FE6EA8B6FDD622C1" bold="true" box="[485,542,1928,1954]" pageId="29" pageNumber="30">C–D</emphasis>
. 
<taxonomicName id="4C7319CA693BEF15FDA5A8B6FBF522C1" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[558,1085,1928,1954]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fuscolabris">
<emphasis id="B907BE5B693BEF15FDA5A8B6FC9322C1" box="[558,859,1928,1954]" italics="true" pageId="29" pageNumber="30">Amphidromus fuscolabris</emphasis>
<bibRefCitation id="EFE21FB8693BEF15FCEFA8B6FBF522C1" author="Mollendorff O. F. von" box="[868,1085,1928,1954]" pageId="29" pageNumber="30" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041).
</paragraph>
</caption>
<paragraph id="8BCC62496938EF16FF61AE85FE5124B7" blockId="30.[189,1399,442,542]" box="[234,409,442,469]" pageId="30" pageNumber="31">
<emphasis id="B907BE5B6938EF16FF61AE85FE5124B7" bold="true" box="[234,409,442,469]" pageId="30" pageNumber="31">
<typeStatus id="54C8DCEB6938EF16FF61AE85FEEF24B6" box="[234,295,443,469]" pageId="30" pageNumber="31">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BCC62496938EF16FF36AEE1FC81277D" blockId="30.[189,1399,442,542]" pageId="30" pageNumber="31">
<collectingCountry id="F36422D96938EF16FF36AEE1FEA9249A" box="[189,353,479,505]" name="Cambodia" pageId="30" pageNumber="31">CAMBODIA</collectingCountry>
: 
<typeStatus id="54C8DCEB6938EF16FEE0AEE1FE1F249A" box="[363,471,479,505]" pageId="30" pageNumber="31" type="lectotype">lectotype</typeStatus>
of var. “ 
<emphasis id="B907BE5B6938EF16FDB1AEE1FD1B249A" box="[570,723,479,505]" italics="true" pageId="30" pageNumber="31">multifasciata</emphasis>
” 
<bibRefCitation id="EFE21FB86938EF16FD6CAEE1FCB6249A" author="Fulton H. C." box="[743,894,479,505]" pageId="30" pageNumber="31" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
, 
<collectionCode id="ED62FA8C6938EF16FC02AEE1FC30249A" box="[905,1016,479,505]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
19601458 (
<bibRefCitation id="EFE21FB86938EF16FB09AEE1FABB249A" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[1154,1395,478,505]" pageId="30" pageNumber="31" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B6938EF16FB7FAEDEFAEE249A" box="[1268,1318,479,505]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 11f); 
<typeStatus id="54C8DCEB6938EF16FEA1AD3DFE1F277E" box="[298,471,515,541]" pageId="30" pageNumber="31" type="paralectotype">paralectotypes</typeStatus>
, 
<collectionCode id="ED62FA8C6938EF16FE69AD3DFD99277E" box="[482,593,515,541]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
19601459 (2 shells).
</paragraph>
<paragraph id="8BCC62496938EF16FF36AD7BFE5827E7" blockId="30.[189,1398,581,644]" pageId="30" pageNumber="31">
<collectingCountry id="F36422D96938EF16FF36AD7BFEA9273C" box="[189,353,581,607]" name="Cambodia" pageId="30" pageNumber="31">CAMBODIA</collectingCountry>
: 
<typeStatus id="54C8DCEB6938EF16FEE0AD7BFE1F273C" box="[363,471,581,607]" pageId="30" pageNumber="31" type="lectotype">lectotype</typeStatus>
of var. “ 
<emphasis id="B907BE5B6938EF16FDB7AD7BFD5C273C" box="[572,660,581,607]" italics="true" pageId="30" pageNumber="31">clausus</emphasis>
” 
<bibRefCitation id="EFE21FB86938EF16FD2CAD7BFC8B273C" author="Pilsbry H. A." box="[679,835,581,607]" pageId="30" pageNumber="31" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
, 
<collectionCode id="ED62FA8C6938EF16FCC6AD7BFC5F273C" box="[845,919,581,607]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
31496 (
<figureCitation id="13487ECC6938EF16FC7CAD7BFBA9273C" box="[1015,1121,581,607]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="30" pageNumber="31">Fig. 11D</figureCitation>
); 
<typeStatus id="54C8DCEB6938EF16FBFCAD7BFAEC273C" box="[1143,1316,581,607]" pageId="30" pageNumber="31" type="paralectotype">paralectotypes</typeStatus>
, 
<collectionCode id="ED62FA8C6938EF16FAA7AD7BFABE273C" box="[1324,1398,581,607]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="30" pageNumber="31">ANSP</collectionCode>
252752 (2 shells).
</paragraph>
<paragraph id="8BCC62496938EF16FF61AD95FE6E27A6" blockId="30.[189,1399,683,999]" box="[234,422,683,709]" pageId="30" pageNumber="31">
<emphasis id="B907BE5B6938EF16FF61AD95FE6E27A6" bold="true" box="[234,422,683,709]" pageId="30" pageNumber="31">Other material</emphasis>
</paragraph>
<paragraph id="8BCC62496938EF16FF36ADEEFACF2685" blockId="30.[189,1399,683,999]" pageId="30" pageNumber="31">
<collectingCountry id="F36422D96938EF16FF36ADEEFEC62789" box="[189,270,720,746]" name="Laos" pageId="30" pageNumber="31">LAOS</collectingCountry>
: 1 shell, Xieng-Moi (W Laos) (
<collectionCode id="ED62FA8C6938EF16FD03ADEEFD312789" box="[648,761,720,746]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
1893.12.8.40, ex. Dautzenberg collection); 32 shells, Ban Phon Pai, Bachieng District, 
<collectingRegion id="49B7ACAB6938EF16FDCEADCAFD06266D" box="[581,718,756,782]" country="Laos" name="Champasak" pageId="30" pageNumber="31">Champasak</collectingRegion>
(
<collectionCode id="ED62FA8C6938EF16FD56ADCAFCFC266D" box="[733,820,756,782]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7030); 9 shells, Thad Phasoam, Boloven Plateau, Paksong District, 
<collectingRegion id="49B7ACAB6938EF16FE10AC26FDEC2651" box="[411,548,792,818]" country="Laos" name="Champasak" pageId="30" pageNumber="31">Champasak</collectingRegion>
(
<collectionCode id="ED62FA8C6938EF16FDB2AC26FD582651" box="[569,656,792,818]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7031); 6 shells, Ban Oudom, Pakbeng District, 
<collectingRegion id="49B7ACAB6938EF16FB7FAC26FABE2651" box="[1268,1398,792,818]" country="Laos" name="Oudomxai" pageId="30" pageNumber="31">Oudomxay</collectingRegion>
(
<collectionCode id="ED62FA8C6938EF16FF4EAC02FED42635" box="[197,284,828,854]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7032); 7 shells, Mouhot’s Tomb, E bank of Kan River, Luang Phrabang District, Luang Phrabang (
<collectionCode id="ED62FA8C6938EF16FF4EAC5EFED42619" box="[197,284,864,890]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7033); 
<specimenCount id="9D75A9C06938EF16FEFEAC5EFDC02619" box="[373,520,864,890]" pageId="30" pageNumber="31" type="generic">2 specimens</specimenCount>
, in ethanol (
<figureCitation id="13487ECC6938EF16FD1DAC5EFD172619" box="[662,735,864,890]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="30" pageNumber="31">Figs 9</figureCitation>
C–D, 14A–B), Thad Kacham Waterfall, Luang Phrabang District, Luang Phrabang (
<collectionCode id="ED62FA8C6938EF16FE75ACBAFD9D26FD" box="[510,597,900,926]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7035); 4 shells (
<figureCitation id="13487ECC6938EF16FCACACBAFC5B26FD" box="[807,915,900,926]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="30" pageNumber="31">Fig. 13E</figureCitation>
, 
<collectionCode id="ED62FA8C6938EF16FC28ACBAFC3026FD" box="[931,1016,900,926]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7036), 
<specimenCount id="9D75A9C06938EF16FBD2ACBAFB2726FD" box="[1113,1263,900,926]" pageId="30" pageNumber="31" type="generic">2 specimens</specimenCount>
, in ethanol (
<figureCitation id="13487ECC6938EF16FF4DAC96FED026A1" box="[198,280,936,962]" captionStart="Fig" captionStartId="24.[189,232,1824,1850]" captionTargetBox="[194,1391,713,1777]" captionTargetId="figure@24.[189,1398,706,1783]" captionTargetPageId="24" captionText="Fig. 12. SEM images of the radula. A–C. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). D–F. Amphidromus xiengensis Morlet, 1891 from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037). G–I. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="30" pageNumber="31">Fig. 12</figureCitation>
D−F, 
<collectionCode id="ED62FA8C6938EF16FED6AC96FE7A26A1" box="[349,434,936,962]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7037), Ban Na Deauy, Luang Phrabang District, Luang Phrabang; 
<specimenCount id="9D75A9C06938EF16FB40AC96FA9C26A1" box="[1227,1364,936,962]" pageId="30" pageNumber="31" type="generic">1 specimen</specimenCount>
, in ethanol, Thad Khaungsy Waterfall, Luang Phrabang District, Luang Phrabang (
<collectionCode id="ED62FA8C6938EF16FBD6ACF2FB7C2685" box="[1117,1204,972,998]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7039).
</paragraph>
<paragraph id="8BCC62496938EF16FF36AB31FE11212E" blockId="30.[189,1398,1038,1101]" pageId="30" pageNumber="31">
<collectingCountry id="F36422D96938EF16FF36AB31FEA9214A" box="[189,353,1039,1065]" name="Cambodia" pageId="30" pageNumber="31">CAMBODIA</collectingCountry>
: 1 shell, Dautzenberg collection (
<collectionCode id="ED62FA8C6938EF16FD7BAB31FC8E214A" box="[752,838,1039,1065]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="30" pageNumber="31" type="Museum">RBINS</collectionCode>
); 3 shells, Lao Moutains (
<collectionCode id="ED62FA8C6938EF16FB0BAB31FB39214A" box="[1152,1265,1039,1065]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="31" type="Museum">NHMUK</collectionCode>
19601539, ex. Cuming Collection).
</paragraph>
<paragraph id="8BCC62496938EF16FF36AB4BFB3421B4" blockId="30.[189,1399,1141,1240]" pageId="30" pageNumber="31">
<collectingCountry id="F36422D96938EF16FF36AB4BFE9121EC" box="[189,345,1141,1167]" name="Thailand" pageId="30" pageNumber="31">THAILAND</collectingCountry>
: 1 shell (
<figureCitation id="13487ECC6938EF16FE36AB4BFDED21EC" box="[445,549,1141,1167]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="30" pageNumber="31">Fig. 13B</figureCitation>
, 
<collectionCode id="ED62FA8C6938EF16FDBBAB4BFD4D21EC" box="[560,645,1141,1167]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7050); 1 shell (
<figureCitation id="13487ECC6938EF16FCBCAB4BFC6821EC" box="[823,928,1141,1167]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="30" pageNumber="31">Fig. 13C</figureCitation>
), Tam Chiang Dao, Chiang Dao District, 
<collectingRegion id="49B7ACAB6938EF16FF36ABA7FE8021D0" box="[189,328,1177,1203]" country="Thailand" name="Chiang Mai" pageId="30" pageNumber="31">Chiang Mai</collectingRegion>
(
<collectionCode id="ED62FA8C6938EF16FED3ABA7FE6721D0" box="[344,431,1177,1203]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7034); 1 shell (
<figureCitation id="13487ECC6938EF16FDECABA7FD0721D0" box="[615,719,1177,1203]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="30" pageNumber="31">Fig. 13F</figureCitation>
), Pha Tang, Pong District, Prayao (
<collectionCode id="ED62FA8C6938EF16FBE0ABA7FB0A21D0" box="[1131,1218,1177,1203]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7052); 3 shells (
<figureCitation id="13487ECC6938EF16FF4DAB83FED021B4" box="[198,280,1213,1239]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="30" pageNumber="31">Fig. 13</figureCitation>
G–H), Doi Phou Nang National Park, Chiangmouan District, 
<collectingRegion id="49B7ACAB6938EF16FC66AB83FB8B21B4" box="[1005,1091,1213,1239]" country="Thailand" name="Phayao" pageId="30" pageNumber="31">Phayao</collectingRegion>
(
<collectionCode id="ED62FA8C6938EF16FBD9AB83FB6121B4" box="[1106,1193,1213,1239]" country="0" name="Cameroon University, Museum of Zoology" pageId="30" pageNumber="31">CUMZ</collectionCode>
7038).
</paragraph>
</subSubSection>
<subSubSection id="C36931C26938EF17FF36ABC1FDBD27B0" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="8BCC62496938EF16FF36ABC1FE86207A" blockId="30.[189,334,1279,1305]" box="[189,334,1279,1305]" pageId="30" pageNumber="31">
<heading id="D084D5256938EF16FF36ABC1FE86207A" bold="true" box="[189,334,1279,1305]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B907BE5B6938EF16FF36ABC1FE86207A" bold="true" box="[189,334,1279,1305]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496938EF16FF36AA11FD622342" blockId="30.[189,1399,1327,1570]" pageId="30" pageNumber="31">Shell sinistral, conical to elongate conical, rather thick and glossy. Spire conical; apex acute with brown or black spot on tip. Whorls 6 to 7 slightly convex; suture depressed; last whorl rounded. Periostracum thin and corneous. Last whorl with reddish to brown Band 1; Bands 2 and 3 as concurrent slanted brownish blotches continuous to expanded lip; Bands 4 and 5 with brownish spiral bands; Band 6 absent or yellowish to reddish. Parietal callus transparent or weakly thickened and white. Aperture subovate; peristome rather thick and expanded; lip whitish. Columella thickened, perpendicular and white. Umbilicus narrowly opened to perforated.</paragraph>
<paragraph id="8BCC62496938EF16FF61A974FE8D2307" blockId="30.[189,1399,1610,1888]" box="[234,325,1610,1636]" pageId="30" pageNumber="31">
<emphasis id="B907BE5B6938EF16FF61A974FE8D2307" bold="true" box="[234,325,1610,1636]" pageId="30" pageNumber="31">Radula</emphasis>
</paragraph>
<paragraph id="8BCC62496938EF16FF36A950FD182203" blockId="30.[189,1399,1610,1888]" pageId="30" pageNumber="31">
Each row contains about 157 (79-(21-17)-1-(17-21)-77) teeth. Central tooth monocuspid, with spatulate shape. Lateral teeth bicuspid; endocone with elongate shape, curved cusp and separated from truncated or curved cusps of ectocone by wide notch, teeth gradually transformed to tricuspid marginal teeth. Marginal teeth tricuspid starting around teeth number 17 to 21 outwards; endocone curved and pointed cusps; mesocone large, with truncate or obtuse cusps; ectocone small, of triangular shape, with pointed cusp. Outermost teeth with pointed cusps on endocone; mesocone large, with curved cusps; extocone sometimes with serrated shape (
<figureCitation id="13487ECC6938EF16FDBFA878FD4F2203" box="[564,647,1862,1888]" captionStart="Fig" captionStartId="24.[189,232,1824,1850]" captionTargetBox="[194,1391,713,1777]" captionTargetId="figure@24.[189,1398,706,1783]" captionTargetPageId="24" captionText="Fig. 12. SEM images of the radula. A–C. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). D–F. Amphidromus xiengensis Morlet, 1891 from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037). G–I. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="30" pageNumber="31">Fig. 12</figureCitation>
D−F).
</paragraph>
<paragraph id="8BCC62496938EF16FF61A8B6FE6C22C0" blockId="30.[189,1399,1928,2028]" box="[234,420,1928,1955]" pageId="30" pageNumber="31">
<emphasis id="B907BE5B6938EF16FF61A8B6FE6C22C0" bold="true" box="[234,420,1928,1955]" pageId="30" pageNumber="31">Genital organs</emphasis>
</paragraph>
<paragraph id="8BCC62496938EF16FF36A893FABF2288" blockId="30.[189,1399,1928,2028]" pageId="30" pageNumber="31">Atrium (at) somewhat short (n = 5). Penis (p) long, cylindrical and enlarged at middle. Epiphallus (e) cylindrical, two times as long as penis; flagellum (fl) short, one third as long as epiphallus; appendix</paragraph>
<paragraph id="8BCC62496939EF17FF36AE35FCC62428" blockId="31.[189,1398,267,331]" pageId="31" pageNumber="32">
absent. Penial retractor muscle (pr) thickened and relatively short. Vas deferens (vd) small tube, connecting epiphallus and free oviduct (
<figureCitation id="13487ECC6939EF17FD1FAE0FFCC92428" box="[660,769,305,331]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="31" pageNumber="32">Fig. 14A</figureCitation>
).
</paragraph>
<paragraph id="8BCC62496939EF17FF36AE4BFC7E24D6" blockId="31.[189,1398,373,437]" pageId="31" pageNumber="32">
Internal penial wall corrugated into weak penial pilasters, which form fringe around penial verge. Penial verge conical, rough surface and orifice open near tip (
<figureCitation id="13487ECC6939EF17FCB6AEA4FC6124D6" box="[829,937,410,437]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="31" pageNumber="32">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="8BCC62496939EF17FF36AEE0FE75270A" blockId="31.[189,1399,478,618]" pageId="31" pageNumber="32">
Vagina cylindrical, long, slender, about 1.5 times as long as penis. Gametolytic duct long, proximal to genital orifice of same diameter as vagina and distally tapering; distal to genital orifice very small tube of about same length as proximal part and connected to gametolytic sac. Oviduct and albumin gland very large (
<figureCitation id="13487ECC6939EF17FEC8AD71FE78270A" box="[323,432,591,617]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="31" pageNumber="32">Fig. 14A</figureCitation>
).
</paragraph>
<paragraph id="8BCC62496939EF17FF36ADADFDBD27B0" blockId="31.[189,1399,659,723]" pageId="31" pageNumber="32">
Internal wall of vagina shows longitudinal vaginal pilasters (vp); pilasters vary narrow and thin throughout vaginal length (
<figureCitation id="13487ECC6939EF17FE77AD87FDAF27B0" box="[508,615,697,723]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="31" pageNumber="32">Fig. 14B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36931C26939EF17FF36ADC3FD35260D" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BCC62496939EF17FF36ADC3FE9E2674" blockId="31.[189,342,765,791]" box="[189,342,765,791]" pageId="31" pageNumber="32">
<heading id="D084D5256939EF17FF36ADC3FE9E2674" bold="true" box="[189,342,765,791]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B907BE5B6939EF17FF36ADC3FE9E2674" bold="true" box="[189,342,765,791]" pageId="31" pageNumber="32">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496939EF17FF36AC10FD35260D" blockId="31.[189,1398,814,878]" pageId="31" pageNumber="32">
The species is widely distributed and can be found in several habitats such as forests, fruit orchards and limestone areas between latitudes 14° and 
<geoCoordinate id="EE47048E6939EF17FD3BAC6DFD31260D" box="[688,761,851,878]" degrees="16" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="55555" value="16.0">16° N</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C36931C26939EF17FF36ACA6FAE323CD" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BCC62496939EF17FF36ACA6FEE726D1" blockId="31.[189,303,920,946]" box="[189,303,920,946]" pageId="31" pageNumber="32">
<heading id="D084D5256939EF17FF36ACA6FEE726D1" bold="true" box="[189,303,920,946]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B907BE5B6939EF17FF36ACA6FEE726D1" bold="true" box="[189,303,920,946]" pageId="31" pageNumber="32">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496939EF17FF36ACF6FC86206C" blockId="31.[189,1399,968,1296]" pageId="31" pageNumber="32">
<taxonomicName id="4C7319CA6939EF17FF36ACF6FE2E2681" authorityName="Morlet" authorityYear="1891" box="[189,486,968,994]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B6939EF17FF36ACF6FE2E2681" box="[189,486,968,994]" italics="true" pageId="31" pageNumber="32">Amphidromus xiengensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7319CA6939EF17FD07ACF7FCD72681" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[652,799,968,995]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="areolatus">
<emphasis id="B907BE5B6939EF17FD07ACF7FCD72681" box="[652,799,968,995]" italics="true" pageId="31" pageNumber="32">A. areolatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7319CA6939EF17FCD4ACF7FC2E2681" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[863,998,968,995]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="zebrinus">
<emphasis id="B907BE5B6939EF17FCD4ACF7FC2E2681" box="[863,998,968,995]" italics="true" pageId="31" pageNumber="32">A. zebrinus</emphasis>
</taxonomicName>
in having a larger shell, reddish Band 1, concurrent slanted and brownish blotched Bands 2 and 3, and brownish spiral Bands 4 and 5. In contrast, the other two species have a smaller shell, Band 1 absent and Bands 2 and 3 merged and becoming brownish slanted blotches in 
<taxonomicName id="4C7319CA6939EF17FD57AB04FCAD2130" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[732,869,1081,1108]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="zebrinus">
<emphasis id="B907BE5B6939EF17FD57AB04FCAD2130" box="[732,869,1081,1108]" italics="true" pageId="31" pageNumber="32">A. zebrinus</emphasis>
</taxonomicName>
or brownish slanted blotches with a flame shape in 
<taxonomicName id="4C7319CA6939EF17FEADAB61FE7D211A" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[294,437,1119,1145]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="areolatus">
<emphasis id="B907BE5B6939EF17FEADAB61FE7D211A" box="[294,437,1119,1145]" italics="true" pageId="31" pageNumber="32">A. areolatus</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="4C7319CA6939EF17FDF8AB61FCBF211A" authority="Ancey, 1897" authorityName="Ancey" authorityYear="1897" box="[627,887,1119,1145]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="eudeli">
<emphasis id="B907BE5B6939EF17FDF8AB61FD14211A" box="[627,732,1119,1145]" italics="true" pageId="31" pageNumber="32">A. eudeli</emphasis>
Ancey, 1897
</taxonomicName>
(
<figureCitation id="13487ECC6939EF17FC0CAB61FC3B211A" box="[903,1011,1119,1145]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="31" pageNumber="32">Fig. 13N</figureCitation>
, 
<typeStatus id="54C8DCEB6939EF17FB8BAB61FB93211A" box="[1024,1115,1119,1145]" pageId="31" pageNumber="32" type="syntype">syntype</typeStatus>
RBINS 617427) and 
<taxonomicName id="4C7319CA6939EF17FAD6AB61FDD521FC" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fuscolabris">
<emphasis id="B907BE5B6939EF17FAD6AB61FEF721FC" italics="true" pageId="31" pageNumber="32">A. fuscolabris</emphasis>
<bibRefCitation id="EFE21FB86939EF17FECEABBBFDD521FC" author="Mollendorff O. F. von" box="[325,541,1157,1183]" pageId="31" pageNumber="32" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13487ECC6939EF17FDA0ABBBFD4221FC" box="[555,650,1157,1183]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="31" pageNumber="32">Fig. 13I</figureCitation>
, 
<typeStatus id="54C8DCEB6939EF17FD1EABBBFD3421FC" box="[661,764,1157,1183]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
SMF 7641) in having a white peristome, reddish Band 1, and Bands 2 and 3 usually separated. In comparison, 
<taxonomicName id="4C7319CA6939EF17FCC3AB95FC2221A7" authorityName="Mollendorff" authorityYear="1898" box="[840,1002,1194,1221]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fuscolabris">
<emphasis id="B907BE5B6939EF17FCC3AB95FC2221A7" box="[840,1002,1194,1221]" italics="true" pageId="31" pageNumber="32">A. fuscolabris</emphasis>
</taxonomicName>
shows a purplish-pink peristome, yellowish Band 1, Bands 2 to 5 merged and roseated Band 6, while 
<taxonomicName id="4C7319CA6939EF17FC67ABEEFB9E2189" authorityName="Ancey" authorityYear="1897" box="[1004,1110,1232,1258]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="eudeli">
<emphasis id="B907BE5B6939EF17FC67ABEEFB9E2189" box="[1004,1110,1232,1258]" italics="true" pageId="31" pageNumber="32">A. eudeli</emphasis>
</taxonomicName>
exhibits a brownish lip, yellow color Band 1 and Bands 2 and 3 usually merged.
</paragraph>
<paragraph id="8BCC62496939EF17FF36AA04FE972019" blockId="31.[189,1398,1337,1402]" pageId="31" pageNumber="32">
Two subspecific entities have been proposed; we, however, recognize them as a single biological species, 
<taxonomicName id="4C7319CA6939EF17FF36AA5EFE90201A" authorityName="Morlet" authorityYear="1891" box="[189,344,1376,1402]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B6939EF17FF36AA5EFE90201A" box="[189,344,1376,1402]" italics="true" pageId="31" pageNumber="32">A. xiengensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCC62496939EF17FF36AAAAFAD920B7" blockId="31.[189,1399,1428,1492]" pageId="31" pageNumber="32">
1. Typical form: Band 1 reddish to brownish; Bands 2 and 3 concurrent slanted brownish blotches continuous to expanded lip; Bands 4 and 5 brownish spiral band; Band 6 absent (
<figureCitation id="13487ECC6939EF17FBFFAA87FB0E20B7" box="[1140,1222,1465,1492]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="31" pageNumber="32">Fig. 13</figureCitation>
A–C).
</paragraph>
<paragraph id="8BCC62496939EF17FF36AAD0FEDA2336" blockId="31.[189,1399,1518,1621]" pageId="31" pageNumber="32">
2. var. “ 
<emphasis id="B907BE5B6939EF17FEABAAD0FE71236B" box="[288,441,1518,1544]" italics="true" pageId="31" pageNumber="32">multifasciata</emphasis>
” 
<bibRefCitation id="EFE21FB86939EF17FE5BAAD0FDA5236B" author="Fulton H. C." box="[464,621,1518,1544]" pageId="31" pageNumber="32" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
: similar to the typical form, but Band 1 yellowish; Bands 2 and 3 usually divided into several smaller bands; Band 6 reddish (
<figureCitation id="13487ECC6939EF17FC23A92AFC34234D" box="[936,1020,1556,1582]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="31" pageNumber="32">Fig. 13</figureCitation>
G–H; 
<bibRefCitation id="EFE21FB86939EF17FBC1A92AFA8B234C" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[1098,1347,1556,1583]" pageId="31" pageNumber="32" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B6939EF17FB36A92BFB3B234D" box="[1213,1267,1556,1582]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 11f–g).
</paragraph>
<paragraph id="8BCC62496939EF17FF36A951FAE323CD" blockId="31.[189,1399,1646,1711]" pageId="31" pageNumber="32">
3. var. “ 
<emphasis id="B907BE5B6939EF17FE97A950FEBC23EB" box="[284,372,1646,1672]" italics="true" pageId="31" pageNumber="32">clausus</emphasis>
” 
<bibRefCitation id="EFE21FB86939EF17FE02A950FDE123EA" author="Pilsbry H. A." box="[393,553,1646,1673]" pageId="31" pageNumber="32" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
: similar to the typical form and var. “ 
<emphasis id="B907BE5B6939EF17FC63A950FB4923EB" box="[1000,1153,1646,1672]" italics="true" pageId="31" pageNumber="32">multifasciata</emphasis>
”, but Bands 2 and 3 have disappeared and turned to a pinkish stain on the second half of the last whorl (
<figureCitation id="13487ECC6939EF17FB04A9AAFB2923CD" box="[1167,1249,1684,1710]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="31" pageNumber="32">Fig. 13</figureCitation>
D−F).
</paragraph>
</subSubSection>
</treatment>
</document>