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<document id="271016C88AA9FB8818D1289447B0FC22" ID-CLB-Dataset="7466" ID-DOI="10.11646/zootaxa.4948.4.1" ID-GBIF-Dataset="ae92fbc8-7dd7-4fcd-a469-9f09c96946f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629235" ID-ZooBank="44061E80-81B7-46AF-AD51-9B461C2E2B67" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616487819220" checkinUser="plazi" docAuthor="Diez, Yander L., Monnens, Marlies, Aguirre, Rosa Isabel, Yurduseven, Rana, Jouk, Philippe, Van Steenkiste, Niels W. L., Leander, Brian S., Schockaert, Ernest, Reygel, Patrick, Smeets, Karen &amp; Artois, Tom" docDate="2021" docId="03829F0DFFE8FFBB05DFBBC97B7AFB3F" docLanguage="en" docName="zootaxa.4948.4.1.pdf" docOrigin="Zootaxa 4948 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Simplexcystis asymmetrica Diez, Reygel &amp; Artois 2021, gen. n." docType="treatment" docUuid="BB2226F6-C526-47E4-8B49-F5CF45AB242E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="492" masterDocId="FFBBE775FFCEFF920548BB5E7A00FFF9" masterDocTitle="Taxonomy and phylogeny of Koinocystididae (Platyhelminthes, Kalyptorhynchia) with the description of three new genera and twelve new species" masterLastPageNumber="500" masterPageNumber="451" pageNumber="489" updateTime="1698934503700" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<taxonomicName id="4C2B5598FFE8FFB405DFBBC978F0FF48" authority="Diez, Reygel &amp; Artois" authorityName="Diez, Reygel &amp; Artois" authorityYear="2021" box="[151,752,151,178]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="38" pageNumber="489" phylum="Platyhelminthes" rank="species" species="asymmetrica" status="gen. nov.">
<emphasis id="B95FF209FFE8FFB405DFBBC97BDAFF48" bold="true" box="[151,474,151,178]" italics="true" pageId="38" pageNumber="489">Simplexcystis asymmetrica</emphasis>
Diez, Reygel &amp; Artois
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sp. n.
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(
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<emphasis id="B95FF209FFE8FFB405DFBA7E7BCFFEC3" bold="true" box="[151,463,288,315]" pageId="38" pageNumber="489">Material and distribution.</emphasis>
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Observations on one live specimen, serially sectioned afterwards, designated
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https://id.luomus.fi/
<collectionCode id="ED3AB6DEFFE8FFB4049FBA1A7801FEA6" box="[471,513,324,351]" pageId="38" pageNumber="489">KV</collectionCode>
.655), collected in
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(
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,
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) (Type Locality),
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,
<collectingMunicipality id="6BF0B461FFE8FFB4046DBA377B98FE7A" box="[293,408,361,387]" pageId="38" pageNumber="489">Lanzarote</collectingMunicipality>
,
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(
<date id="FF9508DBFFE8FFB4071EBA36791BFE7A" box="[598,795,360,387]" pageId="38" pageNumber="489" value="2011-10-10">
<collectingDate id="EFD1F133FFE8FFB4071EBA36791BFE7A" box="[598,795,360,387]" pageId="38" pageNumber="489" value="2011-10-10">October 10, 2011</collectingDate>
</date>
), fine sand accumulated at the end of the rocky reef,
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<elevation id="0006C928FFE8FFB4002BBA377F9CFE7A" box="[1379,1436,361,387]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="38" pageNumber="489" unit="m" value="18.0">18 m</elevation>
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deep, salinity 35
</materialsCitation>
‰.
</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFE8FFB4058FBAEF790AFDEB" pageId="38" pageNumber="489" type="etymology">
<paragraph id="8B942E1BFFE8FFB4058FBAEF790AFDEB" blockId="38.[151,1436,288,531]" pageId="38" pageNumber="489">
<emphasis id="B95FF209FFE8FFB4058FBAEF7B4EFE32" bold="true" box="[199,334,433,459]" pageId="38" pageNumber="489">Etymology.</emphasis>
The genus name refers to the fact that the construction of the atrial organs in the new genus is relatively simple compared to that of other koinocystidids. The species name refers to the fact that the copulatory bulb is asymmetric due to the seminal duct entering laterally.
</paragraph>
</subSubSection>
<caption id="DF547E93FFE8FFB405DFBC627ED8F881" ID-DOI="http://doi.org/10.5281/zenodo.4629305" ID-Zenodo-Dep="4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="38" pageNumber="489" startId="38.[151,250,1852,1876]" targetBox="[159,1428,561,1818]" targetPageId="38">
<paragraph id="8B942E1BFFE8FFB405DFBC627ED8F881" blockId="38.[151,1436,1852,1912]" pageId="38" pageNumber="489">
<emphasis id="B95FF209FFE8FFB405DFBC627B20F8AD" bold="true" box="[151,288,1852,1876]" pageId="38" pageNumber="489">FIGURE 21.</emphasis>
<taxonomicName id="4C2B5598FFE8FFB4046FBC627838F8AD" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[295,568,1852,1876]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="38" pageNumber="489" phylum="Platyhelminthes" rank="species" species="asymmetrica" status="gen. nov.">
<emphasis id="B95FF209FFE8FFB4046FBC627838F8AD" box="[295,568,1852,1876]" italics="true" pageId="38" pageNumber="489">Simplexcystis asymmetrica</emphasis>
</taxonomicName>
<emphasis id="B95FF209FFE8FFB40777BC6278C8F8AD" bold="true" box="[575,712,1852,1876]" pageId="38" pageNumber="489">
<taxonomicNameLabel id="A26C4F72FFE8FFB40777BC627887F8AD" box="[575,647,1852,1876]" pageId="38" pageNumber="489" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side.
</paragraph>
</caption>
<subSubSection id="C3317D90FFE8FFB5058FBCF87BA7FF00" lastPageId="39" lastPageNumber="490" pageId="38" pageNumber="489" type="diagnosis">
<paragraph id="8B942E1BFFE8FFB4058FBCF87F9CF81D" blockId="38.[151,1436,1957,2021]" pageId="38" pageNumber="489">
<emphasis id="B95FF209FFE8FFB4058FBCF8785CF839" bold="true" box="[199,604,1957,1984]" pageId="38" pageNumber="489">
Diagnosis of
<taxonomicName id="4C2B5598FFE8FFB4042DBCFB7801F839" authority="Representative" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[357,513,1957,1984]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="38" pageNumber="489" phylum="Platyhelminthes" rank="genus" status="gen. nov.">
<emphasis id="B95FF209FFE8FFB4042DBCFB7801F839" bold="true" box="[357,513,1957,1984]" italics="true" pageId="38" pageNumber="489">Simplexcystis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C4F72FFE8FFB40742BCF8785CF839" box="[522,604,1958,1984]" pageId="38" pageNumber="489" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
Representative of
<taxonomicName id="4C2B5598FFE8FFB4067FBCF879EFF839" authorityName="Meixner" authorityYear="1924" box="[823,1007,1958,1984]" class="Rhabditophora" family="Koinocystididae" kingdom="Animalia" order="Rhabdocoela" pageId="38" pageNumber="489" phylum="Platyhelminthes" rank="family">Koinocystididae</taxonomicName>
with strong juncture sphincter in between proboscis bulb and cone. Male duct running eccentrically through the copulatory bulb, the latter also enclos-
</paragraph>
<paragraph id="8B942E1BFFE9FFB505DFBBC97BA7FF00" blockId="39.[151,1437,151,1006]" pageId="39" pageNumber="490">ing the prostate vesicle. Copulatory organ devoid of any hard structure. Without bursa. Epithelium of the female, the male, and part of the common atrium lined by a brush border. All these structures surrounded by a sheath of muscles in different orientations.</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFE9FFB5058FBA5D7F59FEE4" box="[199,1369,258,285]" pageId="39" pageNumber="490" type="type_taxon">
<paragraph id="8B942E1BFFE9FFB5058FBA5D7F59FEE4" blockId="39.[151,1437,151,1006]" box="[199,1369,258,285]" pageId="39" pageNumber="490">
<emphasis id="B95FF209FFE9FFB5058FBA5D7B64FEE5" bold="true" box="[199,356,258,285]" pageId="39" pageNumber="490">
<typeStatus id="549090B9FFE9FFB5058FBA5D7B02FEE4" box="[199,258,259,285]" pageId="39" pageNumber="490">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C2B5598FFE9FFB50424BA5D7817FEE5" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[364,535,259,285]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="39" pageNumber="490" phylum="Platyhelminthes" rank="species" species="asymmetrica" status="sp. nov.">
<emphasis id="B95FF209FFE9FFB50424BA5D7817FEE5" box="[364,535,259,285]" italics="true" pageId="39" pageNumber="490">S. asymmetrica</emphasis>
</taxonomicName>
<emphasis id="B95FF209FFE9FFB50756BA5D785DFEE4" bold="true" box="[542,605,259,285]" pageId="39" pageNumber="490">
<taxonomicNameLabel id="A26C4F72FFE9FFB50756BA5D785DFEE4" box="[542,605,259,285]" pageId="39" pageNumber="490" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(by monotypy). Provisionally with the same diagnosis as the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3317D90FFE9FFBB058FBA787B7AFB3F" lastPageId="41" lastPageNumber="492" pageId="39" pageNumber="490" type="description">
<paragraph id="8B942E1BFFE9FFB5058FBA7879DEFDE3" blockId="39.[151,1437,151,1006]" pageId="39" pageNumber="490">
<emphasis id="B95FF209FFE9FFB5058FBA787B59FEB9" bold="true" box="[199,345,294,320]" pageId="39" pageNumber="490">Description.</emphasis>
Live specimen about
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long, translucent, with pinkish parenchymal glands and a pair of eyes (
<figureCitation id="1310329EFFE9FFB50590BA157B41FE9C" box="[216,321,331,357]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21A</figureCitation>
: e). The brain (
<figureCitation id="1310329EFFE9FFB504A3BA157854FE9C" box="[491,596,331,357]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21A</figureCitation>
: br) is located caudally from the proboscis. The syncytial and fully ciliated epidermis is
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thick, with cilia
<quantity id="4CD383FEFFE9FFB50773BA317892FE70" box="[571,658,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="39" pageNumber="490" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 µm</quantity>
long. The epidermis shows many vacuoles some of which are empty, others filled with a dark, granular secretion. The epidermis shows many rhabdites (
<figureCitation id="1310329EFFE9FFB50167BACD7EB4FE54" box="[1071,1204,403,429]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B–C</figureCitation>
,
<figureCitation id="1310329EFFE9FFB501F7BACD7EDBFE54" box="[1215,1243,403,429]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="39" pageNumber="490">22</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB501B5BACD7F4FFE54" box="[1277,1359,403,429]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23C–E</figureCitation>
: rh) all over the body, except for the part anterior to the proboscis. Rhabdites are also lacking around the mouth and gonopore. The rhabdites situated near the apical surface of the epithelium are globular,
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in diameter, while those more to the basal part of the epithelium are more elongated (
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long).
</paragraph>
<paragraph id="8B942E1BFFE9FFB5058FB97D7E72FC17" blockId="39.[151,1437,151,1006]" pageId="39" pageNumber="490">
The proboscis (
<figureCitation id="1310329EFFE9FFB5043DB97D7BDEFDC4" box="[373,478,547,573]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21A</figureCitation>
: pr, 21B, 23A) is of the typical koinocystidid construction (see
<bibRefCitation id="EFBA53EAFFE9FFB501E5B97D7F3BFDC4" author="Brunet, M." box="[1197,1339,547,573]" pageId="39" pageNumber="490" pagination="157 - 174" refId="ref37355" refString="Brunet, M. (1972) Koinocystididae de la region de Marseille. Zoologica Scripta, 1, 157 - 174. https: // doi. org / 10.1111 / j. 1463 - 6409.1972. tb 00673. x" type="journal article" year="1972">Brunet 1972</bibRefCitation>
;
<bibRefCitation id="EFBA53EAFFE9FFB50000B97D7AD1FD98" author="Karling, T. G." pageId="39" pageNumber="490" pagination="241 - 269" refId="ref38008" refString="Karling, T. G. (1980) Revision of Koinocystididae (Turbellaria). Zoologica Scripta, 9, 241 - 269. https: // doi. org / 10.1111 / j. 1463 - 6409.1980. tb 00666. x" type="journal article" year="1980">Karling 1980</bibRefCitation>
), with well-developed cone retractors (
<figureCitation id="1310329EFFE9FFB50730B91978D7FD98" box="[632,727,583,609]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB507B1B919792CFD98" box="[761,812,583,609]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23A</figureCitation>
: cret) and nuclei in the parenchyma of the proboscis bulb. In between the bulb and the cone there is a strong juncture sphincter (
<figureCitation id="1310329EFFE9FFB506C1B9357E0AFD7C" box="[905,1034,619,645]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21A–B</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB50165B9357E60FD7C" box="[1069,1120,619,645]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23A</figureCitation>
: js). In the live specimen, the proboscis is about 15% of the body length. Several
<typeStatus id="549090B9FFE9FFB50790B9D17912FD50" box="[728,786,655,681]" pageId="39" pageNumber="490">types</typeStatus>
of glands enter the proboscis through its caudal wall: dorsally coarse-grained basophilic ones (
<figureCitation id="1310329EFFE9FFB5077BB9ED7898FD34" box="[563,664,691,717]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: prg1), more centrally larger, fine-grained eosinophilic glands (stained pinkish) (
<figureCitation id="1310329EFFE9FFB50449B9897B67FD08" box="[257,359,727,753]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: prg2), and ventrally small fine-grained eosinophilic glands (stained yellowish) (
<figureCitation id="1310329EFFE9FFB501A0B9897F4FFD08" box="[1256,1359,727,753]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: prg3). There are three pairs of proboscis retractors (
<figureCitation id="1310329EFFE9FFB507C8B9A578E4FCEC" box="[640,740,763,789]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: pret). The exact number of fixators (
<figureCitation id="1310329EFFE9FFB50131B9A57ED9FCEC" box="[1145,1241,763,789]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB501B4B9A57F2FFCEC" box="[1276,1327,763,789]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23A</figureCitation>
: pfix) and dilatators (
<figureCitation id="1310329EFFE9FFB50443B8417B6FFCC0" box="[267,367,799,825]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: dil) could not be determined. There is one pair of ventral and one pair of dorsal integument retractors (
<figureCitation id="1310329EFFE9FFB50598B81D7B37FCA4" box="[208,311,835,861]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: iret). The proboscis sheath is lined by a high, nucleated epithelium and is surrounded by an external longitudinal muscle layer. The sheath epithelium is continuous with the epithelium of the proboscis cone, which is very low and devoid of nuclei. The most proximal part of the sheath epithelium contains some oval to circular-shaped eosinophilic gland cells (
<figureCitation id="1310329EFFE9FFB504E2B8F1780BFC30" box="[426,523,943,969]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB50778B8F17863FC30" box="[560,611,943,969]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23A</figureCitation>
: egl). The epithelium surrounding the cone is lined by a brush border (
<figureCitation id="1310329EFFE9FFB50038B8F17ACAFC14" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">Fig. 23A</figureCitation>
: bb). The proboscis pore (
<figureCitation id="1310329EFFE9FFB504AFB88D784BFC14" box="[487,587,979,1005]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: prp) is surrounded by a sphincter (
<figureCitation id="1310329EFFE9FFB50685B88D7E31FC14" box="[973,1073,979,1005]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21B</figureCitation>
: sph).
</paragraph>
<caption id="DF547E93FFE9FFB505DFBDA87F88F8F6" ID-DOI="http://doi.org/10.5281/zenodo.4629308" ID-Zenodo-Dep="4629308" box="[151,1416,1782,1807]" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="39" pageNumber="490" startId="39.[151,250,1782,1806]" targetBox="[193,1392,1035,1753]" targetPageId="39">
<paragraph id="8B942E1BFFE9FFB505DFBDA87F88F8F6" blockId="39.[151,1416,1782,1807]" box="[151,1416,1782,1807]" pageId="39" pageNumber="490">
<emphasis id="B95FF209FFE9FFB505DFBDA87B20F8F7" bold="true" box="[151,288,1782,1806]" pageId="39" pageNumber="490">FIGURE 22.</emphasis>
<taxonomicName id="4C2B5598FFE9FFB5046EBDA97836F8F7" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[294,566,1783,1807]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="39" pageNumber="490" phylum="Platyhelminthes" rank="species" species="asymmetrica" status="gen. nov.">
<emphasis id="B95FF209FFE9FFB5046EBDA97836F8F7" box="[294,566,1783,1807]" italics="true" pageId="39" pageNumber="490">Simplexcystis asymmetrica</emphasis>
</taxonomicName>
<emphasis id="B95FF209FFE9FFB50775BDA978C3F8F6" bold="true" box="[573,707,1783,1807]" pageId="39" pageNumber="490">
<taxonomicNameLabel id="A26C4F72FFE9FFB50775BDA97884F8F6" box="[573,644,1783,1807]" pageId="39" pageNumber="490" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
Sagittal reconstruction of the genital system from the right hand side.
</paragraph>
</caption>
<paragraph id="8B942E1BFFE9FFBA058FBC637ECFFE0C" blockId="39.[151,1437,1853,2024]" lastBlockId="40.[151,1437,151,501]" lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490">
The pharynx (
<figureCitation id="1310329EFFE9FFB5042DBC637BCEF8AE" box="[357,462,1853,1879]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21A</figureCitation>
: ph, 21C, 23B) is located at 40%. In the live specimen, it has a diameter of 10% of the body length. The prepharyngeal cavity (
<figureCitation id="1310329EFFE9FFB50711BC3F78C1F882" box="[601,705,1889,1915]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21C</figureCitation>
: ppc) is lined by a nucleated epithelium, and is surrounded by an internal circular and an outer longitudinal muscle layer. The mouth (
<figureCitation id="1310329EFFE9FFB506D8BCDB79F7F866" box="[912,1015,1925,1951]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21C</figureCitation>
: m) can be closed by a sphincter (
<figureCitation id="1310329EFFE9FFB50027BCDB7ACAF83A" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21C</figureCitation>
: sph). Three
<typeStatus id="549090B9FFE9FFB50416BCF77B99F83A" box="[350,409,1961,1987]" pageId="39" pageNumber="490">types</typeStatus>
of glands open into the pharynx lumen: coarse-grained eosinophilic ones (stained reddish) opening most distally (
<figureCitation id="1310329EFFE9FFB504E8BC937806F81E" box="[416,518,1997,2023]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="39" pageNumber="490">Fig. 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE9FFB50778BC937862F81E" box="[560,610,1997,2023]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="39" pageNumber="490">23B</figureCitation>
: phg1), and, opening more proximally coarse-grained eosinophilic ones (stained dark pinkish) (
<figureCitation id="1310329EFFE6FFBA04D5BBC97801FF48" box="[413,513,151,177]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE6FFBA076FBBC9785AFF48" box="[551,602,151,177]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">23B</figureCitation>
: phg2) and coarse-grained basophilic ones (stained greenish) (
<figureCitation id="1310329EFFE6FFBA0053BBC97F7EFF48" box="[1307,1406,151,177]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE6FFBA05DFBBE57ACAFF2C" box="[151,202,187,213]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">23B</figureCitation>
: phg3). The oesophagus (
<figureCitation id="1310329EFFE6FFBA04ADBBE57884FF2C" box="[485,644,187,213]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21A, 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE6FFBA07EFBBE578DAFF2C" box="[679,730,187,213]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">23B</figureCitation>
: oe) is lined by a nucleated epithelium. A bundle of glands (
<figureCitation id="1310329EFFE6FFBA0027BBE57ACAFF00" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">Fig. 23B</figureCitation>
: oeg) opens into the oesophagus. It consists of coarse-grained basophilic glands (
<figureCitation id="1310329EFFE6FFBA011CBB817EBBFF00" box="[1108,1211,223,249]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: oeg1), fine-grained basophilic ones (
<figureCitation id="1310329EFFE6FFBA041EBA5D7BBEFEE4" box="[342,446,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: oeg2), and fine-grained eosinophilic ones (
<figureCitation id="1310329EFFE6FFBA06E7BA5D7E17FEE4" box="[943,1047,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: oeg3). The pharynx lumen is surrounded by a low, nucleated epithelium. The musculature of the pharynx consists of a layer of longitudinal muscles outside of the septum (
<figureCitation id="1310329EFFE6FFBA04DCBA157BFAFE9C" box="[404,506,331,357]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: lm), which is continuous with the longitudinal muscles surrounding the prepharyngeal cavity, and a circular one just inside of the septum (
<figureCitation id="1310329EFFE6FFBA0647BA317976FE70" box="[783,886,367,393]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: cm1). The distal opening of the pharynx is lined by a thick layer of longitudinal muscles, which in sagittal section, gives the impression of forming a lip-like structure (
<figureCitation id="1310329EFFE6FFBA059BBAE97B37FE28" box="[211,311,439,465]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE6FFBA0415BAE97B90FE28" box="[349,400,439,465]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">23B</figureCitation>
: slm). The pharynx lumen is surrounded by a circular muscle layer (
<figureCitation id="1310329EFFE6FFBA01DFBAE97EFFFE28" box="[1175,1279,439,465]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
: cm2). Radial muscles run (
<figureCitation id="1310329EFFE6FFBA0465BA857B90FE0C" box="[301,400,475,501]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="40" pageNumber="491">Fig. 21C</figureCitation>
&amp;
<figureCitation id="1310329EFFE6FFBA04FCBA857BE7FE0C" box="[436,487,475,501]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491">23B</figureCitation>
: rm) between the wall of the pharynx lumen and the outer septum.
</paragraph>
<caption id="DF547E93FFE6FFBA05DFBCFA78D4F818" ID-DOI="http://doi.org/10.5281/zenodo.4629310" ID-Zenodo-Dep="4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="40" pageNumber="491" startId="40.[151,250,1956,1980]" targetBox="[170,1416,525,1933]" targetPageId="40">
<paragraph id="8B942E1BFFE6FFBA05DFBCFA78D4F818" blockId="40.[151,1436,1956,2017]" pageId="40" pageNumber="491">
<emphasis id="B95FF209FFE6FFBA05DFBCFA7B20F845" bold="true" box="[151,288,1956,1980]" pageId="40" pageNumber="491">FIGURE 23.</emphasis>
<taxonomicName id="4C2B5598FFE6FFBA046EBCFA7836F845" authority="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois, 2021" authorityName="Diez &amp; Monnens &amp; Aguirre &amp; Yurduseven &amp; Jouk &amp; Van Steenkiste &amp; Leander &amp; Schockaert &amp; Reygel &amp; Smeets &amp; Artois" authorityYear="2021" box="[294,566,1956,1980]" class="Rhabditophora" family="Koinocystididae" genus="Simplexcystis" kingdom="Animalia" order="Rhabdocoela" pageId="40" pageNumber="491" phylum="Platyhelminthes" rank="species" species="asymmetrica" status="gen. nov.">
<emphasis id="B95FF209FFE6FFBA046EBCFA7836F845" box="[294,566,1956,1980]" italics="true" pageId="40" pageNumber="491">Simplexcystis asymmetrica</emphasis>
</taxonomicName>
<emphasis id="B95FF209FFE6FFBA0774BCFB78C3F844" bold="true" box="[572,707,1957,1981]" pageId="40" pageNumber="491">
<taxonomicNameLabel id="A26C4F72FFE6FFBA0774BCFB7883F844" box="[572,643,1957,1981]" pageId="40" pageNumber="491" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs.
</paragraph>
</caption>
<paragraph id="8B942E1BFFE7FFBB058FBBC9793FFCEC" blockId="41.[151,1437,151,1222]" pageId="41" pageNumber="492">
The two elongated testes (
<figureCitation id="1310329EFFE7FFBB04BDBBC97860FF48" box="[501,608,151,177]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
: t) are located rostrally from the pharynx. They seem to be divided into three follicles that are closely packed together. However, this could also be an optic effect caused by the testes being heavily folded. The vasa deferentia run towards the caudal body end, and distally form the elongated seminal vesicles (
<figureCitation id="1310329EFFE7FFBB05B7BA5D7B63FEE4" box="[255,355,259,285]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB04CFBA5D7BA7FEE4" box="[391,423,259,285]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
: sv). The seminal vesicles are lined by a low, nucleated epithelium, and are surrounded by longitudinal muscles. The copulatory bulb is oviform, and lies in the caudal body third (
<figureCitation id="1310329EFFE7FFBB0130BA797EDCFEB8" box="[1144,1244,295,321]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB0049BA797F33FEB8" box="[1281,1331,295,321]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C</figureCitation>
: cb). It is surrounded by a longitudinal muscle layer. Distally from the seminal vesicles, the vasa deferentia enter the ejaculatory duct separately (
<figureCitation id="1310329EFFE7FFBB04D4BA317805FE70" box="[412,517,367,393]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB075ABA31782EFE70" box="[530,558,367,393]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB071CBA3178A6FE70" box="[596,678,367,393]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C–E</figureCitation>
: ed). The ejaculatory duct is elongated, with its distal half situated eccentrically within the copulatory bulb (
<figureCitation id="1310329EFFE7FFBB0729BACD78C3FE54" box="[609,707,403,429]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB07ADBACD7918FE54" box="[741,792,403,429]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C</figureCitation>
: cb). Distally the seminal duct opens into the male atrium, surrounded by the prostate vesicle (
<figureCitation id="1310329EFFE7FFBB0761BAE9787BFE28" box="[553,635,439,465]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB07E8BAE978F2FE28" box="[672,754,439,465]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C–E</figureCitation>
: pv), which also opens in the male atrium. The seminal duct is lined by a low nucleated epithelium, and is surrounded by longitudinal muscles. The prostate glands (
<figureCitation id="1310329EFFE7FFBB000ABA857F99FE0C" box="[1346,1433,475,501]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
: pg) are extracapsular, without clearly separated gland necks in the prostate vesicle. The latter contains a coarsegrained eosinophilic secretion in its proximal half and a fine-grained one in its distal half. The male atrium (
<figureCitation id="1310329EFFE7FFBB0004B97D7F9CFDC4" box="[1356,1436,547,573]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB05FDB9197B07FD98" box="[181,263,583,609]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23D–F</figureCitation>
: ma) is divided into a short, broad proximal part, and a tubiform distal part, which ends in the common genital atrium (
<figureCitation id="1310329EFFE7FFBB040DB9357B97FD7C" box="[325,407,619,645]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB04F5B9357BEFFD7C" box="[445,495,619,645]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C</figureCitation>
: ca). The male atrium, the female atrium, and the common genital atrium are lined by a nucleated epithelium and are surrounded by longitudinal muscles. Outside of this longitudinal muscle layer, a thick layer of muscles with different orientations (probably circular and oblique ones) and connective tissue occurs (
<figureCitation id="1310329EFFE7FFBB05E8B9897AF1FD08" box="[160,241,727,753]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB045EB9897B66FD08" box="[278,358,727,753]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C–F</figureCitation>
: tl). At its apical side the epithelium is lined by a brush border (
<figureCitation id="1310329EFFE7FFBB017EB9897E87FD08" box="[1078,1159,727,753]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB01E4B9897EFEFD08" box="[1196,1278,727,753]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23D–F</figureCitation>
: bb), which is lacking in the most distal part of the common genital atrium.
</paragraph>
<paragraph id="8B942E1BFFE7FFBB058FB8417B7AFB3F" blockId="41.[151,1437,151,1222]" pageId="41" pageNumber="492">
In the live specimen, we only observed a single, poorly differentiated ovary (
<figureCitation id="1310329EFFE7FFBB0176B8417EA4FCC0" box="[1086,1188,799,825]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB0184B8417EECFCC0" box="[1228,1260,799,825]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
: ov), located at the level of the seminal vesicles. In the serially-sectioned specimen, we observed two weakly developed oviducts, suggesting that the female system in full maturity has two ovaries. The very long oviducts (
<figureCitation id="1310329EFFE7FFBB01E2B8397F00FC78" box="[1194,1280,871,897]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
: od) are lined by a nucleated epithelium; muscles were not observed. They open proximally into the female duct (
<figureCitation id="1310329EFFE7FFBB01B9B8D57F58FC5C" box="[1265,1368,907,933]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB002BB8D57F7FFC5C" box="[1379,1407,907,933]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB05DFB8F17AE9FC30" box="[151,233,943,969]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23D–F</figureCitation>
: fd). The female duct is lined by a nucleated epithelium and contains sperm. It opens into the female atrium (
<figureCitation id="1310329EFFE7FFBB05E8B88D7AF4FC14" box="[160,244,979,1005]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB0448B88D7B85FC14" box="[256,389,979,1005]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23D &amp; 23F</figureCitation>
: fa) through a strong sphincter (
<figureCitation id="1310329EFFE7FFBB07A7B88D7953FC14" box="[751,851,979,1005]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB0631B88D79C8FC14" box="[889,968,979,1005]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23E–F</figureCitation>
: sph, 22: sph1). The female atrium enters the common genital atrium dorsally. There is no bursa. The uterus (
<figureCitation id="1310329EFFE7FFBB06DEB8A979FFFBE8" box="[918,1023,1015,1041]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB0143B8A97E2AFBE8" box="[1035,1066,1015,1041]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB017FB8A97E69FBE8" box="[1079,1129,1015,1041]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C</figureCitation>
: ut) is lined by a nucleated epithelium and surrounded by longitudinal muscles. Coarse-grained eosinophilic uterine glands (
<figureCitation id="1310329EFFE7FFBB0187BF457F23FBCC" box="[1231,1315,1051,1077]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
: ueg) open into the uterus somewhat proximally from the entry point into the common genital atrium. Just distally from the entry point of the uterus, some fine-grained eosinophilic atrial glands (
<figureCitation id="1310329EFFE7FFBB06ECBF3D79F8FB84" box="[932,1016,1123,1149]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
: ag) enter the common genital atrium. The common gonopore lies ventrally, at about 95% (
<figureCitation id="1310329EFFE7FFBB0796BFD97945FB58" box="[734,837,1159,1185]" captionStart="FIGURE 21" captionStartId="38.[151,250,1852,1876]" captionTargetBox="[159,1428,561,1818]" captionTargetId="figure-135@38.[151,1436,554,1827]" captionTargetPageId="38" captionText="FIGURE 21. Simplexcystis asymmetrica gen. n. sp. n. A, general organisation (from a live animal). B, sagittal reconstruction of the proboscis from the right hand side. C, sagittal reconstruction of the pharynx from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629305" httpUri="https://zenodo.org/record/4629305/files/figure.png" pageId="41" pageNumber="492">Fig. 21A</figureCitation>
,
<figureCitation id="1310329EFFE7FFBB0607BFD9796BFB58" box="[847,875,1159,1185]" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">22</figureCitation>
&amp;
<figureCitation id="1310329EFFE7FFBB06C5BFD979BFFB58" box="[909,959,1159,1185]" captionStart="FIGURE 23" captionStartId="40.[151,250,1956,1980]" captionTargetBox="[170,1416,525,1933]" captionTargetId="figure-211@40.[170,1416,525,1933]" captionTargetPageId="40" captionText="FIGURE 23. Simplexcystis asymmetrica gen. n. sp. n. A, sagittal section through the proboscis. B, sagittal section through the pharynx. C–F, sagittal sections through the atrial organs." figureDoi="http://doi.org/10.5281/zenodo.4629310" httpUri="https://zenodo.org/record/4629310/files/figure.png" pageId="41" pageNumber="492">23C</figureCitation>
: cg), and can be closed by a sphincter (
<figureCitation id="1310329EFFE7FFBB0027BFD97AB7FB3C" captionStart="FIGURE 22" captionStartId="39.[151,250,1782,1806]" captionTargetBox="[193,1392,1035,1753]" captionTargetId="figure-449@39.[189,1398,1029,1759]" captionTargetPageId="39" captionText="FIGURE 22. Simplexcystis asymmetrica gen. n. sp. n. Sagittal reconstruction of the genital system from the right hand side." figureDoi="http://doi.org/10.5281/zenodo.4629308" httpUri="https://zenodo.org/record/4629308/files/figure.png" pageId="41" pageNumber="492">Fig. 22</figureCitation>
: sph2, 23C: sph).
</paragraph>
</subSubSection>
</treatment>
</document>
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