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<mods:title id="873624D7691C7EEC295003D28A7E0EE6">Correction of existing generic and species concepts in Platyceroidini (Coleoptera: Lucanidae: Lucaninae) and the description of four new species of Platyceroides Benesh</mods:title>
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<mods:namePart id="40A767D051101FE86F06583CA535EB00">Paulsen, M. J.</mods:namePart>
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<treatment id="0382520FFFC7FFABFF53F6BC8414F8C3" ID-DOI="http://doi.org/10.5281/zenodo.5189919" ID-GBIF-Taxon="130844393" ID-Zenodo-Dep="5189919" LSID="urn:lsid:plazi:treatment:0382520FFFC7FFABFF53F6BC8414F8C3" httpUri="http://treatment.plazi.org/id/0382520FFFC7FFABFF53F6BC8414F8C3" lastPageNumber="362" pageId="16" pageNumber="362">
<subSubSection id="C331B092FFC7FFABFF53F6BC84B9FD74" box="[151,753,655,682]" pageId="16" pageNumber="362" type="nomenclature">
<paragraph id="8B94E319FFC7FFABFF53F6BC84B9FD74" blockId="16.[151,753,655,682]" box="[151,753,655,682]" pageId="16" pageNumber="362">
<heading id="D0DC5475FFC7FFABFF53F6BC84B9FD74" bold="true" box="[151,753,655,682]" fontSize="11" level="1" pageId="16" pageNumber="362" reason="1">
<taxonomicName id="4C2B989AFFC7FFABFF53F6BC84B9FD74" ID-CoL="7ZQBS" authority="Casey" authorityName="Casey" box="[151,753,655,682]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus" subGenus="Platyceroides">
<emphasis id="B95F3F0BFFC7FFABFF53F6BC84B9FD74" bold="true" box="[151,753,655,682]" pageId="16" pageNumber="362">
<emphasis id="B95F3F0BFFC7FFABFF53F6BC8770FD77" bold="true" box="[151,312,655,681]" italics="true" pageId="16" pageNumber="362">Platyceroides</emphasis>
(
<emphasis id="B95F3F0BFFC7FFABFE8DF6BC87A2FD77" bold="true" box="[329,490,655,681]" italics="true" pageId="16" pageNumber="362">Platyceroides</emphasis>
)
<emphasis id="B95F3F0BFFC7FFABFE3FF6BC84C4FD77" bold="true" box="[507,652,655,681]" italics="true" pageId="16" pageNumber="362">californicus</emphasis>
(Casey)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF53F6E585ADFCD5" pageId="16" pageNumber="362" type="reference_group">
<paragraph id="8B94E319FFC7FFABFF53F6E58557FD32" blockId="16.[151,997,726,779]" box="[151,799,726,748]" pageId="16" pageNumber="362">
<treatmentCitationGroup id="AB3BC437FFC7FFABFF53F6E58557FD32" box="[151,799,726,748]" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFF53F6E58472FD32" ID-CoL="7WJBB" authority="Casey, 1885: 331" authorityName="Casey" authorityPageNumber="331" authorityYear="1885" box="[151,570,726,748]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFF53F6E587CAFD32" box="[151,386,726,748]" italics="true" pageId="16" pageNumber="362">Platycerus californicus</emphasis>
<treatmentCitation id="0A8AC508FFC7FFABFE4DF6E58472FD32" author="Casey" box="[393,570,726,748]" page="331" pageId="16" pageNumber="362" year="1885">
<bibRefCitation id="EFBA9EE8FFC7FFABFE4DF6E58440FD32" author="Casey" box="[393,520,726,748]" pageId="16" pageNumber="362" refString="Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1, 283 - 336." type="journal article" year="1885">Casey, 1885</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
, original combination.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B94E319FFC7FFABFF53F6C685ADFCD5" blockId="16.[151,997,726,779]" box="[151,997,757,779]" pageId="16" pageNumber="362">
<treatmentCitationGroup id="AB3BC437FFC7FFABFF53F6C685ADFCD5" box="[151,997,757,779]" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFF53F6C68465FCD5" authority="Casey, 1889: 164" authorityName="Casey" authorityPageNumber="164" authorityYear="1889" box="[151,557,757,779]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFF53F6C6873EFCD5" box="[151,374,757,779]" italics="true" pageId="16" pageNumber="362">Platycerus parvicollis</emphasis>
<treatmentCitation id="0A8AC508FFC7FFABFEB8F6C68465FCD5" author="Casey" box="[380,557,757,779]" page="164" pageId="16" pageNumber="362" year="1889">
<bibRefCitation id="EFBA9EE8FFC7FFABFEB8F6C687B3FCD5" author="Casey" box="[380,507,757,779]" pageId="16" pageNumber="362" refString="Casey, T. L. (1889) Coleopterological notices. I. Annals of the New York Academy of Sciences, 5, 39 - 96. https: // doi. org / 10.1111 / j. 1749 - 6632.1890. tb 57003. x" type="journal article" year="1889">Casey, 1889</bibRefCitation>
: 164
</treatmentCitation>
</taxonomicName>
, original combination.
<emphasis id="B95F3F0BFFC7FFABFCDCF6C685ADFCD5" bold="true" box="[792,997,757,779]" pageId="16" pageNumber="362">Revised synonymy.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF53F7048413FC46" pageId="16" pageNumber="362" type="materials_examined">
<paragraph id="8B94E319FFC7FFABFF53F7048413FC46" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
<emphasis id="B95F3F0BFFC7FFABFF53F704876AFC8E" bold="true" box="[151,290,823,848]" pageId="16" pageNumber="362">
<typeStatus id="54905DBBFFC7FFABFF53F704869AFC8E" box="[151,210,823,848]" pageId="16" pageNumber="362">Type</typeStatus>
series.
</emphasis>
<taxonomicName id="4C2B989AFFC7FFABFEEDF70B8466FC8E" box="[297,558,824,848]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFEEDF70B8466FC8E" box="[297,558,824,848]" italics="true" pageId="16" pageNumber="362">Platycerus californicus</emphasis>
</taxonomicName>
<typeStatus id="54905DBBFFC7FFABFDF0F70484DEFC8E" box="[564,662,823,848]" pageId="16" pageNumber="362" type="holotype">holotype</typeStatus>
female (
<collectionCode id="ED3A7BDCFFC7FFABFD3CF70B8504FC8E" box="[760,844,824,848]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="362">USNM</collectionCode>
) labeled: a) “Cal.”; b) “
<collectionCode id="ED3A7BDCFFC7FFABFB90F70B82FAFC8E" box="[1108,1202,824,848]" pageId="16" pageNumber="362">CASEY</collectionCode>
/ bequest / 1925”; c) on orange paper, “
<typeStatus id="54905DBBFFC7FFABFEA2F76F87E7FCAA" box="[358,431,860,884]" pageId="16" pageNumber="362">
<collectionCode id="ED3A7BDCFFC7FFABFEA2F76F87E7FCAA" box="[358,431,860,884]" pageId="16" pageNumber="362">TYPE</collectionCode>
</typeStatus>
<collectionCode id="ED3A7BDCFFC7FFABFE73F76F8440FCAA" box="[439,520,860,884]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="362">USNM</collectionCode>
/ [36209]”; d) handwritten “
<taxonomicName id="4C2B989AFFC7FFABFC8BF76F859DFCAA" box="[847,981,860,884]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFC8BF76F859DFCAA" box="[847,981,860,884]" italics="true" pageId="16" pageNumber="362">californicus</emphasis>
</taxonomicName>
/ Csy.”; e) “
<taxonomicName id="4C2B989AFFC7FFABFBA1F76F83C4FCAA" box="[1125,1420,860,884]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFBA1F76F83C4FCAA" box="[1125,1420,860,884]" italics="true" pageId="16" pageNumber="362">Platyceroides californicus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFBA9EE8FFC7FFABFF5AF7B38762FC46" author="Casey" box="[158,298,895,920]" pageId="16" pageNumber="362" refString="Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1, 283 - 336." type="journal article" year="1885">Casey, 1885</bibRefCitation>
) / det. M.J. Paulsen 2013”.
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF03F7978716FAFA" pageId="16" pageNumber="362" type="reference_group">
<paragraph id="8B94E319FFC7FFABFF03F797874AFBF6" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFF03F79787F6FC62" box="[199,446,932,956]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFF03F79787F6FC62" box="[199,446,932,956]" italics="true" pageId="16" pageNumber="362">Platycerus parvicollis</emphasis>
</taxonomicName>
<typeStatus id="54905DBBFFC7FFABFE02F7978464FC63" box="[454,556,932,957]" pageId="16" pageNumber="362" type="lectotype">lectotype</typeStatus>
male (USNM),
<emphasis id="B95F3F0BFFC7FFABFD21F79785E8FC63" bold="true" box="[741,928,932,957]" pageId="16" pageNumber="362">here designated</emphasis>
, labeled: a) handwritten “
<taxonomicName id="4C2B989AFFC7FFABFB03F7978377FC62" box="[1223,1343,932,956]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFB03F7978377FC62" box="[1223,1343,932,956]" italics="true" pageId="16" pageNumber="362">parvicollis</emphasis>
</taxonomicName>
/ Cas.”; b) “CASEY / bequest / 1925”; c) on orange paper, “
<typeStatus id="54905DBBFFC7FFABFD3FF7FB850BFC3E" box="[763,835,968,992]" pageId="16" pageNumber="362">TYPE</typeStatus>
USNM / [36208]”; d) “
<taxonomicName id="4C2B989AFFC7FFABFBA5F7FB83C2FC3E" ID-CoL="7ZQBS" box="[1121,1418,968,992]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFBA5F7FB83C2FC3E" box="[1121,1418,968,992]" italics="true" pageId="16" pageNumber="362">Platyceroides californicus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFBA9EE8FFC7FFABFF5AF7DF8766FBDB" author="Casey" box="[158,302,1004,1029]" pageId="16" pageNumber="362" refString="Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1, 283 - 336." type="journal article" year="1885">Casey, 1885</bibRefCitation>
) / det.
<bibRefCitation id="EFBA9EE8FFC7FFABFE4CF7DE842BFBDB" author="Paulsen" box="[392,611,1004,1029]" pageId="16" pageNumber="362" refString="Paulsen, M. J. (2015) A new species of Platyceroides Benesh (Coleoptera: Lucanidae) from Oregon. Insecta Mundi, 430, 1 - 5." type="journal article" year="2015">M.J. Paulsen 2015</bibRefCitation>
”; e) on red paper, “
<taxonomicName id="4C2B989AFFC7FFABFC8BF7DF858FFBDA" ID-CoL="6RQ9" box="[847,967,1004,1028]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="genus">
<emphasis id="B95F3F0BFFC7FFABFC8BF7DF858FFBDA" box="[847,967,1004,1028]" italics="true" pageId="16" pageNumber="362">Platycerus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C2B989AFFC7FFABFC20F7DF82D1FBDA" ID-CoL="7ZQBX" authority="Casey" authorityName="Casey" box="[996,1177,1004,1028]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B95F3F0BFFC7FFABFC20F7DF8201FBDA" box="[996,1097,1004,1028]" italics="true" pageId="16" pageNumber="362">pacificus</emphasis>
Casey
</taxonomicName>
/
<typeStatus id="54905DBBFFC7FFABFB72F7DE836EFBDB" box="[1206,1318,1005,1029]" pageId="16" pageNumber="362" type="lectotype">Lectotype</typeStatus>
/ det. MJ Paulsen”.
</paragraph>
<paragraph id="8B94E319FFC7FFABFF03F0078716FAFA" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
<materialsCitation id="3B43E944FFC7FFABFF03F00782BBFBAE" ID-GBIF-Occurrence="1503414391" county="Humboldt County" location="Eureka" municipality="Casey" pageId="16" pageNumber="362" specimenCount="1" stateProvince="California" typeStatus="holotype">
The female
<typeStatus id="54905DBBFFC7FFABFE94F00787FAFB93" box="[336,434,1076,1101]" pageId="16" pageNumber="362" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2B989AFFC7FFABFE1FF0068436FB92" ID-CoL="7ZQBS" box="[475,638,1076,1100]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFE1FF0068436FB92" box="[475,638,1076,1100]" italics="true" pageId="16" pageNumber="362">P. californicus</emphasis>
</taxonomicName>
is labeled simply “Cal.”, although
<bibRefCitation id="EFBA9EE8FFC7FFABFBD5F00782E1FB93" author="Casey" box="[1041,1193,1076,1101]" pageId="16" pageNumber="362" refString="Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1, 283 - 336." type="journal article" year="1885">
<collectingMunicipality id="6BF07963FFC7FFABFBD5F007821EFB92" box="[1041,1110,1076,1100]" pageId="16" pageNumber="362">Casey</collectingMunicipality>
(1885)
</bibRefCitation>
fortunately indicated that the specimen was “captured in a dusty wagon road” in
<location id="8EF4B5C2FFC7FFABFC80F06485DEFBAE" LSID="urn:lsid:plazi:treatment:0382520FFFC7FFABFF53F6BC8414F8C3:8EF4B5C2FFC7FFABFC80F06485DEFBAE" box="[836,918,1111,1136]" county="Humboldt County" municipality="Casey" name="Eureka" pageId="16" pageNumber="362" stateProvince="California">Eureka</location>
,
<collectingCounty id="62F59B95FFC7FFABFC67F064823AFBAE" box="[931,1138,1111,1136]" pageId="16" pageNumber="362">Humboldt County</collectingCounty>
,
<collectingRegion id="49EF2DFBFFC7FFABFBBAF06482BBFBAE" box="[1150,1267,1111,1136]" country="United States of America" name="California" pageId="16" pageNumber="362">California</collectingRegion>
</materialsCitation>
.
<materialsCitation id="3B43E944FFC7FFABFAC4F0648716FAFA" ID-GBIF-Occurrence="1503414382" location="For" municipality="Casey" pageId="16" pageNumber="362" specimenCount="2" stateProvince="California" typeStatus="holotype">
Although two species of the nominal subgenus are located at coastal, low elevations near Eureka, the
<typeStatus id="54905DBBFFC7FFABFB70F04F835EFB4B" box="[1204,1302,1148,1173]" pageId="16" pageNumber="362" type="holotype">holotype</typeStatus>
has highly polished elytra and cannot be the female of
<taxonomicName id="4C2B989AFFC7FFABFDBAF0928554FB69" ID-CoL="7ZQBY" box="[638,796,1184,1208]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="pampinatus" status="sp. nov.">
<emphasis id="B95F3F0BFFC7FFABFDBAF0928554FB69" box="[638,796,1184,1208]" italics="true" pageId="16" pageNumber="362">P. pampinatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26C8270FFC7FFABFCEDF09385FBFB66" box="[809,947,1183,1208]" pageId="16" pageNumber="362" rank="species">new species</taxonomicNameLabel>
, described above. Thus the
<typeStatus id="54905DBBFFC7FFABFB23F0AC8301FB66" box="[1255,1353,1183,1208]" pageId="16" pageNumber="362" type="holotype">holotype</typeStatus>
female belongs to the same taxon later described by
<collectingMunicipality id="6BF07963FFC7FFABFD59F0F784AAFB02" box="[669,738,1220,1244]" pageId="16" pageNumber="362">Casey</collectingMunicipality>
from a male specimen as
<emphasis id="B95F3F0BFFC7FFABFBD4F0F682E5FB02" box="[1040,1197,1220,1244]" italics="true" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFBD4F0F682E0FB02" box="[1040,1192,1220,1244]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">P. parvicollis</taxonomicName>
.
</emphasis>
<location id="8EF4B5C2FFC7FFABFB71F0F68294FB02" LSID="urn:lsid:plazi:treatment:0382520FFFC7FFABFF53F6BC8414F8C3:8EF4B5C2FFC7FFABFB71F0F68294FB02" box="[1205,1244,1221,1244]" municipality="Casey" name="For" pageId="16" pageNumber="362" stateProvince="California">For</location>
this reason I am considering
<taxonomicName id="4C2B989AFFC7FFABFEE5F0DA87FDFADE" box="[289,437,1256,1280]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFEE5F0DA87FDFADE" box="[289,437,1256,1280]" italics="true" pageId="16" pageNumber="362">P. parvicollis</emphasis>
</taxonomicName>
, which I had previously considered to constitute a valid species, to be a junior synonym of
<emphasis id="B95F3F0BFFC7FFABFF71F13E8716FAFA" box="[181,350,1292,1316]" italics="true" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFF71F13E8713FAFA" ID-CoL="7ZQBS" box="[181,347,1292,1316]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">P. californicus</taxonomicName>
.
</emphasis>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF03F11C820CFA23" pageId="16" pageNumber="362" type="description">
<paragraph id="8B94E319FFC7FFABFF03F11C820CFA23" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
The original description of
<taxonomicName id="4C2B989AFFC7FFABFE3EF10284C6FA96" box="[506,654,1328,1352]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFE3EF10284C6FA96" box="[506,654,1328,1352]" italics="true" pageId="16" pageNumber="362">P. parvicollis</emphasis>
</taxonomicName>
lists two specimens, although only one (now the lectotype) is present in the USNM. In the FMNH collection two male specimens are labeled as the type of
<taxonomicName id="4C2B989AFFC7FFABFB95F16682ACFAB2" box="[1105,1252,1364,1388]" class="Insecta" family="Lucanidae" genus="Platycerus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B95F3F0BFFC7FFABFB95F16682ACFAB2" box="[1105,1252,1364,1388]" italics="true" pageId="16" pageNumber="362">P. parvicollis</emphasis>
</taxonomicName>
, and one female was invalidly designated as the allotype by Benesh. Label data indicates that all three specimens are much too recent to have been part of the original type series and are not types; furthermore all three specimens are actually
<emphasis id="B95F3F0BFFC7FFABFA43F1AE86B2FA06" italics="true" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFA43F1AE86BFFA06" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="infernus">P. infernus</taxonomicName>
.
</emphasis>
The second female specimen in the UNSM that was part of the Casey bequest that was identified as
<taxonomicName id="4C2B989AFFC7FFABFA43F1F28755FA22" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFA43F1F28755FA22" italics="true" pageId="16" pageNumber="362">P. californicus</emphasis>
</taxonomicName>
by him is a paratype of the new species
<taxonomicName id="4C2B989AFFC7FFABFD27F1D685C9FA22" box="[739,897,1509,1532]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="pampinatus">
<emphasis id="B95F3F0BFFC7FFABFD27F1D685C9FA22" box="[739,897,1509,1532]" italics="true" pageId="16" pageNumber="362">P. pampinatus</emphasis>
</taxonomicName>
described above.
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF03F2348716F9B6" pageId="16" pageNumber="362" type="distribution">
<paragraph id="8B94E319FFC7FFABFF03F2348716F9B6" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
<emphasis id="B95F3F0BFFC7FFABFF03F2348711F9FE" bold="true" box="[199,345,1543,1568]" pageId="16" pageNumber="362">Distribution</emphasis>
(
<figureCitation id="1310FF9CFFC7FFABFEACF23487F0F9FE" box="[360,440,1543,1568]" captionStart="FIGURES 27 28" captionStartId="29.[151,264,1632,1654]" captionTargetBox="[159,1428,205,1610]" captionTargetId="figure@29.[151,1436,193,1621]" captionTargetPageId="29" captionText="FIGURES 27 28. Distribution of Platyceroides species. 27, Platyceroides californicus species group: P. aeneus (orange triangles); P. umpquus (blue circles); P. californicus (orange squares); P. thoracicus (blue triangles). 28, Platyceroides opacus species group: P. marshalli (blue squares); P. potax (orange squares); P. barrae (blue circles); P. opacus (orange circles)." httpUri="https://zenodo.org/record/582464/files/figure.png" pageId="16" pageNumber="362">Fig. 27</figureCitation>
).
<emphasis id="B95F3F0BFFC7FFABFE0AF23484C4F9FE" bold="true" box="[462,652,1543,1568]" pageId="16" pageNumber="362">
<collectingRegion id="49EF2DFBFFC7FFABFE0AF23484CFF9FE" box="[462,647,1543,1568]" country="United States of America" name="California" pageId="16" pageNumber="362">CALIFORNIA</collectingRegion>
:
</emphasis>
HUMBOLDT: Arcata; Blocksburg; Bull Creek-South Fork Eel River; Dyerville; Eureka; Fieldbrook; Fort Seward; Hydesville; Redcrest; High Rock/Humboldt Redwoods State Park; Van Duzen River.
</paragraph>
</subSubSection>
<subSubSection id="C331B092FFC7FFABFF03F2478414F8C3" pageId="16" pageNumber="362" type="discussion">
<paragraph id="8B94E319FFC7FFABFF03F2478414F8C3" blockId="16.[151,1437,823,1821]" pageId="16" pageNumber="362">
<emphasis id="B95F3F0BFFC7FFABFF03F2478773F953" bold="true" box="[199,315,1652,1677]" pageId="16" pageNumber="362">Remarks.</emphasis>
Compared with
<emphasis id="B95F3F0BFFC7FFABFE33F24684D8F952" box="[503,656,1652,1676]" italics="true" pageId="16" pageNumber="362">
<taxonomicName id="4C2B989AFFC7FFABFE33F24684C5F952" box="[503,653,1652,1676]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="thoracicus">P. thoracicus</taxonomicName>
,
</emphasis>
males of
<taxonomicName id="4C2B989AFFC7FFABFCC4F24685EAF952" box="[768,930,1652,1676]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B95F3F0BFFC7FFABFCC4F24685EAF952" box="[768,930,1652,1676]" italics="true" pageId="16" pageNumber="362">P. californicus</emphasis>
</taxonomicName>
(
<figureCitation id="1310FF9CFFC7FFABFC75F24785BCF952" box="[945,1012,1652,1677]" captionStart="FIGURES 8 11" captionStartId="15.[151,264,1825,1847]" captionTargetBox="[219,1424,201,1801]" captionTargetId="figure@15.[151,1436,193,1804]" captionTargetPageId="15" captionText="FIGURES 8 11. Platyceroides californicus species group, males: A, dorsal habitus; B, flagellum of male genitalia, lateral view; C, apex of flagellum of male genitalia, ventral view. 8, Platyceroides aeneus, arrow indicates large bifurcate process. 9, Platyceroides californicus. 10, Platyceroides thoracicus, arrow indicates small bifurcate process. 11, Platyceroides umpquus new species." httpUri="https://zenodo.org/record/582456/files/figure.png" pageId="16" pageNumber="362">Fig. 9</figureCitation>
A) have shinier, lighter-colored elytra that contrast with their distinctly darker pronotal disc with lighter margins, smaller antennal club size, and the clearly distinct apex of the flagellum of the male genitalia (
<figureCitation id="1310FF9CFFC7FFABFCF4F28F853BF90A" box="[816,883,1724,1749]" captionStart="FIGURES 8 11" captionStartId="15.[151,264,1825,1847]" captionTargetBox="[219,1424,201,1801]" captionTargetId="figure@15.[151,1436,193,1804]" captionTargetPageId="15" captionText="FIGURES 8 11. Platyceroides californicus species group, males: A, dorsal habitus; B, flagellum of male genitalia, lateral view; C, apex of flagellum of male genitalia, ventral view. 8, Platyceroides aeneus, arrow indicates large bifurcate process. 9, Platyceroides californicus. 10, Platyceroides thoracicus, arrow indicates small bifurcate process. 11, Platyceroides umpquus new species." httpUri="https://zenodo.org/record/582456/files/figure.png" pageId="16" pageNumber="362">Fig. 9</figureCitation>
BC). This species lighter coloration is similar to that of
<taxonomicName id="4C2B989AFFC7FFABFF29F2D28713F926" box="[237,347,1761,1784]" class="Insecta" family="Lucanidae" genus="Platyceroides" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="362" phylum="Arthropoda" rank="species" species="aeneus">
<emphasis id="B95F3F0BFFC7FFABFF29F2D28713F926" box="[237,347,1761,1784]" italics="true" pageId="16" pageNumber="362">P. aeneus</emphasis>
</taxonomicName>
from much farther north, but in that species the posterior pronotal angles are obtuse and the genitalia have a bifurcate apex medially.
</paragraph>
</subSubSection>
</treatment>
</document>