treatments-xml/data/03/80/87/038087D7FFFFFF8EFC4DF99CFEB434EB.xml
2024-06-21 12:22:17 +02:00

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<document id="DAA1FFCC5BF3BF3FAACF40E6F5132475" ID-DOI="10.5252/g2015n2a5" ID-ISSN="1638-9395" ID-Zenodo-Dep="4535125" ID-ZooBank="urn:lsid:zoobank.org:pub:0117CBA4-4CE0-4431-B5F6-721F998C72C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613058030568" checkinUser="felipe" docAuthor="Fourvel, Jean-Baptiste, Fosse, Philippe, Fernandez, Philippe &amp; Antoine, Pierre-Olivier" docDate="2015" docId="038087D7FFFFFF8EFC4DF99CFEB434EB" docLanguage="en" docName="Geodiversitas.37.2.237-266.pdf" docOrigin="Geodiversitas 37 (2)" docSource="http://dx.doi.org/10.5252/g2015n2a5" docStyle="DocumentStyle:4DA100580BA89F561A6D37CADC50A8FD.3:Geodiversitas.2015-2017.journal_article.type2" docStyleId="4DA100580BA89F561A6D37CADC50A8FD" docStyleName="Geodiversitas.2015-2017.journal_article.type2" docStyleVersion="3" docTitle="Rangifer tarandus" docType="treatment" docVersion="4" lastPageNumber="261" masterDocId="FFB9FFAFFFE8FF96FFEEFFAEFFA53307" masterDocTitle="Large mammals of Fouvent-Saint-Andoche (Haute-Saône, France): a glimpse into a Late Pleistocene hyena den" masterLastPageNumber="266" masterPageNumber="237" pageNumber="260" updateTime="1698870744442" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="BB99C2749BC583FD6BC87C2298BE5139">Large mammals of Fouvent-Saint-Andoche (Haute-Saône, France): a glimpse into a Late Pleistocene hyena den</mods:title>
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<mods:namePart id="2047EC1E068793AD896613579F36DD5A">Fourvel, Jean-Baptiste</mods:namePart>
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<mods:namePart id="467E9DDCC68434358D47D7E71096059E">Fosse, Philippe</mods:namePart>
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<mods:namePart id="925CFB1568BF676371CDC08CE16BD16B">Fernandez, Philippe</mods:namePart>
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<mods:namePart id="FE62469BA063B991C4278529919EC1DE">Antoine, Pierre-Olivier</mods:namePart>
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<treatment id="038087D7FFFFFF8EFC4DF99CFEB434EB" ID-DOI="http://doi.org/10.5281/zenodo.4535457" ID-Zenodo-Dep="4535457" LSID="urn:lsid:plazi:treatment:038087D7FFFFFF8EFC4DF99CFEB434EB" httpUri="http://treatment.plazi.org/id/038087D7FFFFFF8EFC4DF99CFEB434EB" lastPageId="24" lastPageNumber="261" pageId="23" pageNumber="260">
<subSubSection id="C333654AFFFFFF81FC4DF99CFA9D354A" box="[931,1336,1586,1613]" pageId="23" pageNumber="260" type="nomenclature">
<paragraph id="8B9636C1FFFFFF81FC4DF99CFA9D354A" blockId="23.[931,1336,1586,1613]" box="[931,1336,1586,1613]" pageId="23" pageNumber="260">
<heading id="D0DE81ADFFFFFF81FC4DF99CFA9D354A" box="[931,1336,1586,1613]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="260" reason="2">
<taxonomicName id="4C294D42FFFFFF81FC4DF99CFA9D354A" ID-CoL="4RF6J" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[931,1336,1586,1613]" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageId="23" pageNumber="260" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="B95DEAD3FFFFFF81FC4DF99CFBCA354A" bold="true" box="[931,1135,1586,1613]" italics="true" pageId="23" pageNumber="260">Rangifer tarandus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C333654AFFFFFF81FCC3F9D4FAD835AB" pageId="23" pageNumber="260" type="materials_examined">
<paragraph id="8B9636C1FFFFFF81FCC3F9D4FAA83595" blockId="23.[811,1457,1658,2026]" box="[813,1293,1658,1682]" pageId="23" pageNumber="260">
MATERIAL EXAMINED. —
<collectionCode id="ED38AE04FFFFFF81FBC1F9D4FBCE3595" box="[1071,1131,1658,1682]" pageId="23" pageNumber="260">NISP</collectionCode>
=113;
<collectionCode id="ED38AE04FFFFFF81FB47F9D4FB443595" box="[1193,1249,1658,1682]" pageId="23" pageNumber="260">MNI</collectionCode>
=16.
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<paragraph id="8B9636C1FFFFFF81FCC3F93AFAD835AB" blockId="23.[811,1457,1658,2026]" box="[813,1405,1684,1708]" pageId="23" pageNumber="260">
<emphasis id="B95DEAD3FFFFFF81FCC3F93AFC1235AB" bold="true" box="[813,951,1684,1708]" pageId="23" pageNumber="260">1842 sample:</emphasis>
1 left antler; 3 antler fragments; 1 left talus.
</paragraph>
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<subSubSection id="C333654AFFFFFF8EFCC3F901FEB434EB" lastPageId="24" lastPageNumber="261" pageId="23" pageNumber="260" type="description">
<paragraph id="8B9636C1FFFFFF81FCC3F901FB9534ED" blockId="23.[811,1457,1658,2026]" pageId="23" pageNumber="260">
<emphasis id="B95DEAD3FFFFFF81FCC3F901FC5235C0" bold="true" box="[813,1015,1711,1735]" pageId="23" pageNumber="260">1989-1992 sample:</emphasis>
2 right antlers; 1 left antler; 3 antler fragments; 3 petrosal bones; 1 left P2; 1 left P2-3; 2 right P3-4; 1 right M1-2; 3 left M1-2; 1 right M2; 1 right M3; 1 right i1; 1 left i1; 4 left p4; 1 left m1; 2 left m2; 1 right m3; 4 left m3; 2 right m1-2; 1 left m1-2; 4 tooth fragments; 5 right humeri; 2 left humeri; 4 humeri; 2 left radii; 4 radii; 1 ulna; 2 left magnum; 1 right lunate; 1 left scaphoid; 1 left metacarpal III-IV; 1 left femur; 4 femorae; 1 patella; 4 right tibiae; 1 left tibia; 4 tibiae; 2 naviculo-cuboids; 3 right metatarsals III-IV; 1 left metatarsal III-IV; 2 metatarsals; 1 sesamoid; 2 vestigial metapodials; 1 long bone fragment; 4 metapodials; 7 first phalanges; 5 second phalanges; 4 third phalanges.
</paragraph>
<paragraph id="8B9636C1FFF0FF8EFF7DFF78FD9933E8" blockId="24.[147,572,214,239]" box="[147,572,214,239]" pageId="24" pageNumber="261">A D</paragraph>
<caption id="DF566649FFF0FF8EFF6AFAAFFA143650" ID-DOI="http://doi.org/10.5281/zenodo.4535149" ID-Zenodo-Dep="4535149" httpUri="https://zenodo.org/record/4535149/files/figure.png" pageId="24" pageNumber="261" startId="24.[132,143,1281,1299]" targetBox="[135,1451,237,1242]" targetPageId="24">
<paragraph id="8B9636C1FFF0FF8EFF6AFAAFFA143650" blockId="24.[132,1457,1281,1367]" pageId="24" pageNumber="261">
FIG. 11. — Enamel hypoplasia on the tooth sample of
<taxonomicName id="4C294D42FFF0FF8EFDA3FAAFFC193614" authority="(Blumenbach, 1799)" authorityName="" baseAuthorityName="Blumenbach" baseAuthorityYear="1799" box="[589,956,1281,1299]" class="Mammalia" family="Rhinocerotidae" genus="Coelodonta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="24" pageNumber="261" phylum="Chordata" rank="species" species="antiquitatis">
<emphasis id="B95DEAD3FFF0FF8EFDA3FAAFFCAB3614" box="[589,782,1281,1299]" italics="true" pageId="24" pageNumber="261">Coelodonta antiquitatis</emphasis>
(Blumenbach, 1799)
</taxonomicName>
(woolly rhino) from Fouvent:
<emphasis id="B95DEAD3FFF0FF8EFB5DFAAFFB653614" bold="true" box="[1203,1216,1281,1299]" pageId="24" pageNumber="261">A</emphasis>
,
<emphasis id="B95DEAD3FFF0FF8EFB25FAAFFB7D3614" bold="true" box="[1227,1240,1281,1299]" pageId="24" pageNumber="261">B</emphasis>
, left d4 F11 C342, with a severe hypoplasia, characterized by the total lack of enamel on a part of the ectolophid (grey area).
<emphasis id="B95DEAD3FFF0FF8EFC2EFAB6FC6B362D" bold="true" box="[960,974,1304,1322]" pageId="24" pageNumber="261">C</emphasis>
,
<emphasis id="B95DEAD3FFF0FF8EFC36FAB6FC43362D" bold="true" box="[984,998,1304,1322]" pageId="24" pageNumber="261">D</emphasis>
, left M1 G9 D607;
<emphasis id="B95DEAD3FFF0FF8EFB6DFAB6FB2A362D" bold="true" box="[1155,1167,1304,1322]" pageId="24" pageNumber="261">E</emphasis>
,
<emphasis id="B95DEAD3FFF0FF8EFB74FAB6FB00362D" bold="true" box="[1178,1189,1304,1322]" pageId="24" pageNumber="261">F</emphasis>
, left M2 E11 B218. The last two teeth show less severe hypoplasia, visible through enamel constrictions on the lingual side (G9D607) and the ectoloph (F1 C243). These constrictions, parallel and remote, indicate the occurrence of two distinct episodes of physiological stress on each tooth. A, C, E, photographs. B, D, F, interpretative drawings (POA).
</paragraph>
</caption>
<paragraph id="8B9636C1FFF0FF8EFF6AFA1DFEB036CC" blockId="24.[131,775,1459,2028]" box="[132,277,1459,1485]" pageId="24" pageNumber="261">DESCRIPTION</paragraph>
<paragraph id="8B9636C1FFF0FF8EFF6AFA7DFEB434EB" blockId="24.[131,775,1459,2028]" pageId="24" pageNumber="261">
One hundred and thirteen bone remains (both cranial and postcranial elements) have been referred to as the reindeer,
<taxonomicName id="4C294D42FFF0FF8EFF6DF9BDFEA5352B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[131,256,1554,1580]" class="Mammalia" family="Cervidae" genus="Rangifer" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="261" phylum="Chordata" rank="species" species="tarandus">
<emphasis id="B95DEAD3FFF0FF8EFF6DF9BDFEA5352B" box="[131,256,1554,1580]" italics="true" pageId="24" pageNumber="261">R. tarandus</emphasis>
</taxonomicName>
. Their stratigraphical location at Fouvent suggests a minimal number of 16 individuals. All the typical reindeer morphological features (see
<bibRefCitation id="EFB84B30FFF0FF8EFDDBF9FCFD79356B" author="MAGNIEZ P." box="[565,732,1618,1644]" pageId="24" pageNumber="261" refId="ref26543" refString="MAGNIEZ P. 2010. - Etude paleontologique des Artiodactyles de la grotte Tournal (Bize-Minervois, Aude, France). Etude taphonomique, archeozoologique et paleoecologique des grands Mammiferes dans leur cadre biostratigraphique et paleoenvironnemental. These de Doctorat, Universite de Perpignan Via Domitia, 916 p." type="book" year="2010">Magniez 2010</bibRefCitation>
for a review) could be observed on the material (including teeth, antler and postcranial elements). Unfortunately, this material is too much altered and the sample is not sufficiently abundant to meet the conditions for any population structure analysis (age-structure, sex-ratio...). Be as it may, a left basilar fragment of shed antler (FVT.1842.106) can be associated to a male adult (
<figureCitation id="13122A44FFF0FF8EFE19F89FFDF0344B" box="[503,597,1841,1868]" captionStart="FIG" captionStartId="20.[132,143,1917,1935]" captionTargetBox="[264,1349,228,1810]" captionTargetId="figure-15@20.[206,1382,173,1836]" captionTargetPageId="20" captionText="FIG. 9.— Ungulates (other than rhinoceros) remains from Fouvent: A, B, giant deer Megaloceros giganteus Blumenbach,1799:right 1842.14 (A) and left 1842.13 (B) mandibles; C-E, red deer Cervus elaphus Linnaeus, 1758: basal part of right antler 1842.109 (C) and ingested left talus 1842.97 (D, palmar view; E, dorsal view); F-H, horse Equus germanicus Nehring,1884:right metacarpal 1842.2 (F), first 1842.6 (G) and third H9.A.1 (H) phalanges;I-L, large bovid Bovidae:right calcaneum 1842.78 (I), first phalange G10.E/A.9 (J, dorsal view; K, lateral view) and metapodial distal portion F11.A.50 (L); M-O reindeer Rangifer tarandus (Linnaeus, 1758): basal part of left antler 1842.106 (M), portion of right antler F10.B.329 (N) and right metatarsal G9.24 (O). Scale bar: 20 mm. Pictures by JBF." figureDoi="http://doi.org/10.5281/zenodo.4535145" httpUri="https://zenodo.org/record/4535145/files/figure.png" pageId="24" pageNumber="261">Fig. 9M</figureCitation>
). Another small fragment (FVT.1992.F10.B.329) could match a female or a young male (Averbouh pers. comm.;
<figureCitation id="13122A44FFF0FF8EFDACF8DFFD3B348C" box="[578,670,1905,1931]" captionStart="FIG" captionStartId="20.[132,143,1917,1935]" captionTargetBox="[264,1349,228,1810]" captionTargetId="figure-15@20.[206,1382,173,1836]" captionTargetPageId="20" captionText="FIG. 9.— Ungulates (other than rhinoceros) remains from Fouvent: A, B, giant deer Megaloceros giganteus Blumenbach,1799:right 1842.14 (A) and left 1842.13 (B) mandibles; C-E, red deer Cervus elaphus Linnaeus, 1758: basal part of right antler 1842.109 (C) and ingested left talus 1842.97 (D, palmar view; E, dorsal view); F-H, horse Equus germanicus Nehring,1884:right metacarpal 1842.2 (F), first 1842.6 (G) and third H9.A.1 (H) phalanges;I-L, large bovid Bovidae:right calcaneum 1842.78 (I), first phalange G10.E/A.9 (J, dorsal view; K, lateral view) and metapodial distal portion F11.A.50 (L); M-O reindeer Rangifer tarandus (Linnaeus, 1758): basal part of left antler 1842.106 (M), portion of right antler F10.B.329 (N) and right metatarsal G9.24 (O). Scale bar: 20 mm. Pictures by JBF." figureDoi="http://doi.org/10.5281/zenodo.4535145" httpUri="https://zenodo.org/record/4535145/files/figure.png" pageId="24" pageNumber="261">Fig. 9N</figureCitation>
). Finally, distinct tooth wear stages as observed on isolated teeth suggest various age classes (young adults, adults, old adults, and seniles).
</paragraph>
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