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<document id="112718A5B344E1A3B88884EAB7DAC29C" ID-CLB-Dataset="27715" ID-DOI="10.11646/zootaxa.4541.1.1" ID-GBIF-Dataset="9c3fecf1-4e11-4c79-a908-e0a5abbaf3ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617460" ID-ZooBank="B11E734C-218B-418C-84E6-719AB3C58AFF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927824231" checkinUser="plazi" docAuthor="Arachchige, Gayashan M., Jayakody, Sevvandi, Mooi, Rich &amp; Kroh, Andreas" docDate="2019" docId="038087B4FF848945FF02FF2372AA98C1" docLanguage="en" docName="zootaxa.4541.1.1.pdf" docOrigin="Zootaxa 4541 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Jacksonaster Lambert in Lambert &amp; Thiery 1914" docType="treatment" docVersion="7" lastPageNumber="49" masterDocId="FFB9FFCCFFAA8975FF95FFB47314996C" masterDocTitle="Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka" masterLastPageNumber="100" masterPageNumber="1" pageNumber="47" updateTime="1698715306146" updateUser="ExternalLinkService">
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<mods:title id="6848DD79F43D63185A76384E94F7CA79">Taxonomy and distribution of irregular echinoids (Echinoidea: Irregularia) from Sri Lanka</mods:title>
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<mods:namePart id="33072CA2181DA88FEB60B8BEE812632F">Arachchige, Gayashan M.</mods:namePart>
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<mods:namePart id="65C843C9A3D4BB501986F399AC383C78">Mooi, Rich</mods:namePart>
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<mods:namePart id="F5A1D65A26DE62FA62E67A4A656672F2">Kroh, Andreas</mods:namePart>
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<treatment id="038087B4FF848945FF02FF2372AA98C1" ID-DOI="http://doi.org/10.5281/zenodo.5935450" ID-GBIF-Taxon="154529664" ID-Zenodo-Dep="5935450" LSID="urn:lsid:plazi:treatment:038087B4FF848945FF02FF2372AA98C1" httpUri="http://treatment.plazi.org/id/038087B4FF848945FF02FF2372AA98C1" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="C3336529FF84895BFF02FF23726199DE" box="[151,373,151,178]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B9636A2FF84895BFF02FF23726199DE" blockId="46.[151,373,151,211]" box="[151,373,151,178]" pageId="46" pageNumber="47">
<heading id="D0DE81CEFF84895BFF02FF23726199DE" bold="true" box="[151,373,151,178]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B95DEAB0FF84895BFF02FF23726199DE" bold="true" box="[151,373,151,178]" pageId="46" pageNumber="47">
<taxonomicName id="4C294D21FF84895BFF02FF23722199DD" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[151,309,151,177]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="46" pageNumber="47" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF84895BFF02FF23722199DD" bold="true" box="[151,309,151,177]" italics="true" pageId="46" pageNumber="47">Jacksonaster</emphasis>
</taxonomicName>
sp. 2
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3336529FF84895BFF02FF0E726099BF" box="[151,372,186,211]" pageId="46" pageNumber="47" type="description">
<paragraph id="8B9636A2FF84895BFF02FF0E726099BF" blockId="46.[151,373,151,211]" box="[151,372,186,211]" pageId="46" pageNumber="47">
<figureCitation id="13122A27FF84895BFF02FF0E720499BF" box="[151,272,186,211]" captionStart="FIGURE 39" captionStartId="46.[151,250,1981,2003]" captionTargetBox="[154,1439,339,1945]" captionTargetId="figure@46.[154,1439,339,1945]" captionTargetPageId="46" captionText="FIGURE 39. Jacksonaster sp. 2 (AC, WUSL/EI/25; D, E, WUSL/EI/29): A, D, aboral view; B, E, oral view; C, lateral view." httpUri="https://zenodo.org/record/2617535/files/figure.png" pageId="46" pageNumber="47">Figures 39</figureCitation>
,
<figureCitation id="13122A27FF84895BFE89FF0F725999BF" box="[284,333,187,211]" captionStart="FIGURE 40" captionStartId="47.[151,250,1098,1120]" captionTargetBox="[151,1436,193,1077]" captionTargetId="figure@47.[151,1436,193,1077]" captionTargetPageId="47" captionText="FIGURE 40. Comparison of apical system and petaloid area of Jacksonaster sp. 1 (A, WUSL/EI/26) and Jacksonaster sp. 2 (B, WUSL/EI/29)." httpUri="https://zenodo.org/record/2617537/files/figure.png" pageId="46" pageNumber="47">40B</figureCitation>
,
<figureCitation id="13122A27FF84895BFECDFF0F726099BF" box="[344,372,187,211]" captionStart="FIGURE 41" captionStartId="48.[151,250,1811,1833]" captionTargetBox="[166,1395,514,1671]" captionTargetId="figure@48.[151,1436,483,1789]" captionTargetPageId="48" captionText="FIGURE 41. Distribution map of Jacksonaster sp. 2 recorded in this study." httpUri="https://zenodo.org/record/2617539/files/figure.png" pageId="46" pageNumber="47">41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3336529FF84895BFF02FEB6762B9877" box="[151,1343,258,283]" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B9636A2FF84895BFF02FEB6762B9877" blockId="46.[151,1343,258,283]" box="[151,1343,258,283]" pageId="46" pageNumber="47">
<emphasis id="B95DEAB0FF84895BFF02FEB672709877" bold="true" box="[151,356,258,283]" pageId="46" pageNumber="47">Material studied.</emphasis>
Two denuded tests: WUSL/EI/25, EI/29 from Silavathurai, northern coast of
<collectingCountry id="F33E7632FF84895BFB5FFEB676289877" box="[1226,1340,258,283]" name="Sri Lanka" pageId="46" pageNumber="47">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF56662AFF84895BFF02F809768D9EBF" box="[151,1433,1981,2003]" httpUri="https://zenodo.org/record/2617535/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1981,2003]" targetBox="[154,1439,339,1945]" targetPageId="46">
<paragraph id="8B9636A2FF84895BFF02F809768D9EBF" blockId="46.[151,1433,1981,2003]" box="[151,1433,1981,2003]" pageId="46" pageNumber="47">
<emphasis id="B95DEAB0FF84895BFF02F80972349EBF" bold="true" box="[151,288,1981,2003]" pageId="46" pageNumber="47">FIGURE 39.</emphasis>
<taxonomicName id="4C294D21FF84895BFEB0F80972BD9EBF" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[293,425,1981,2003]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="46" pageNumber="47" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF84895BFEB0F80972BD9EBF" box="[293,425,1981,2003]" italics="true" pageId="46" pageNumber="47">Jacksonaster</emphasis>
</taxonomicName>
sp. 2 (AC, WUSL/EI/25; D, E, WUSL/EI/29): A, D, aboral view; B, E, oral view; C, lateral view.
</paragraph>
</caption>
<caption id="DF56662AFF85895AFF02FBFE72409D13" httpUri="https://zenodo.org/record/2617537/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1098,1120]" targetBox="[151,1436,193,1077]" targetPageId="47">
<paragraph id="8B9636A2FF85895AFF02FBFE72409D13" blockId="47.[151,1436,1098,1151]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF02FBFE72359D33" bold="true" box="[151,289,1098,1120]" pageId="47" pageNumber="48">FIGURE 40.</emphasis>
Comparison of apical system and petaloid area of
<taxonomicName id="4C294D21FF85895AFCBBFBFE70A69D0C" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[814,946,1098,1120]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="47" pageNumber="48" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF85895AFCBBFBFE70A69D0C" box="[814,946,1098,1120]" italics="true" pageId="47" pageNumber="48">Jacksonaster</emphasis>
</taxonomicName>
sp. 1 (A, WUSL/EI/26) and
<taxonomicName id="4C294D21FF85895AFB4AFBFE76779D0C" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[1247,1379,1098,1120]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="47" pageNumber="48" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF85895AFB4AFBFE76779D0C" box="[1247,1379,1098,1120]" italics="true" pageId="47" pageNumber="48">Jacksonaster</emphasis>
</taxonomicName>
sp. 2 (B, WUSL/EI/29).
</paragraph>
</caption>
<subSubSection id="C3336529FF85895AFF52FB7172BC9EC7" pageId="47" pageNumber="48" type="description">
<paragraph id="8B9636A2FF85895AFF52FB7173F79C25" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52FB71724D9DB2" bold="true" box="[199,345,1221,1246]" pageId="47" pageNumber="48">Description.</emphasis>
<emphasis id="B95DEAB0FF85895AFEF7FB71711B9DB1" box="[354,527,1221,1246]" italics="true" pageId="47" pageNumber="48">Shape and size</emphasis>
—Test elliptical to rounded; medium-sized,
<quantity id="4CD19B47FF85895AFB98FB7277609DB1" box="[1037,1140,1222,1246]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" pageId="47" pageNumber="48" unit="mm" value="32.1">32.1 mm</quantity>
and 43.0 mm TL, longer than broad, width 87% and 89% TL; greatest width coinciding with anterior paired petals; TH 18% TL; oral side flat; edge thick, anterior edge thickness approximately 12% TL, posterior edge slightly thinner than anterior edge of test.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52FAE172FF9CFE" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52FAE172739C01" box="[199,359,1365,1390]" italics="true" pageId="47" pageNumber="48">Apical system</emphasis>
—Subcentral, 49% and 50% TL from anterior margin of test; with five circular gonopores; hydropores in sinuous groove.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52FA2977CB9F05" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52FA2972579CDA" box="[199,323,1437,1462]" italics="true" pageId="47" pageNumber="48">Ambulacra</emphasis>
—Petaloid area large, 73% and 77% TL, slightly elevated; petals narrow, closed distally; outer pore of each pair more elongate than inner pore; pores conjugate, but conjugating furrows inconspicuous; anterior unpaired petal slightly longer than others, 35% and 38% TL; anterior paired petals shortest, 31% and 32% TL; posterior paired petals slightly shorter than petal
<date id="FF971062FF85895AFD2EF9BE71EF9F4D" box="[699,763,1545,1569]" pageId="47" pageNumber="48" value="1934-03">III 34</date>
% and 35% TL; all petal widths similar, approximately 12% TL; width of poriferous zone IIIa 128% and 103% of interporiferous zone at 2/3 of the petal length; food grooves simple, unbranched, reaching up to 64% and 71% of corresponding test radius of ambulacrum III.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52F9C176629FE2" blockId="47.[151,1436,1221,2032]" box="[199,1398,1653,1678]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F9C172639FE2" box="[199,375,1653,1678]" italics="true" pageId="47" pageNumber="48">Interambulacra</emphasis>
—On oral side, interambulacra narrow, forming straight bands separating broad ambulacra.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52F92D772E9FBA" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F92D72779FDE" box="[199,355,1689,1714]" italics="true" pageId="47" pageNumber="48">Tuberculation</emphasis>
—Aboral primary tubercles small and densely distributed among numerous miliary tubercles; on oral surface, primary tubercles larger and sparse relative to those on aboral surface.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52F95670D89E71" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F956722D9F95" box="[199,313,1762,1785]" italics="true" pageId="47" pageNumber="48">Peristome</emphasis>
—Small, length approximately 7% TL, width approximately 9% TL; rounded to pentagonal; situated slightly anterior of centre, 46% and 48% TL from anterior margin of test.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52F89E71BB9E09" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F89E72259E2D" box="[199,305,1834,1857]" italics="true" pageId="47" pageNumber="48">Periproct</emphasis>
—Small, length 5% and 7% TL; transversely elongate; width 6% and 8% TL; lying on oral surface 13% and 14% TL from posterior margin of test.
</paragraph>
<paragraph id="8B9636A2FF85895AFF52F8C572BC9EC7" blockId="47.[151,1436,1221,2032]" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F8C571609EE6" bold="true" box="[199,628,1905,1930]" pageId="47" pageNumber="48">
Observed occurrence in
<collectingCountry id="F33E7632FF85895AFE67F8C5717B9EE6" box="[498,623,1905,1930]" name="Sri Lanka" pageId="47" pageNumber="48">Sri Lanka</collectingCountry>
.
</emphasis>
Denuded tests were collected from Silavathurai, northern coast of Sri Lanka at
<quantity id="4CD19B47FF85895AFF6AF827722C9EC6" box="[255,312,1939,1963]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="47" pageNumber="48" unit="m" value="28.0">28 m</quantity>
(
<figureCitation id="13122A27FF85895AFED2F826728F9EC7" box="[327,411,1938,1963]" captionStart="FIGURE 41" captionStartId="48.[151,250,1811,1833]" captionTargetBox="[166,1395,514,1671]" captionTargetId="figure@48.[151,1436,483,1789]" captionTargetPageId="48" captionText="FIGURE 41. Distribution map of Jacksonaster sp. 2 recorded in this study." httpUri="https://zenodo.org/record/2617539/files/figure.png" pageId="47" pageNumber="48">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336529FF858945FF52F80072AA98C1" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8B9636A2FF858945FF52F80071C899B9" blockId="47.[151,1436,1221,2032]" lastBlockId="48.[151,1436,151,429]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B95DEAB0FF85895AFF52F800722F9EA1" bold="true" box="[199,315,1972,1997]" pageId="47" pageNumber="48">Remarks.</emphasis>
The two specimens of
<taxonomicName id="4C294D21FF85895AFDA9F80071DA9EA1" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[572,718,1972,1997]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="47" pageNumber="48" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF85895AFDA9F80071DA9EA1" box="[572,718,1972,1997]" italics="true" pageId="47" pageNumber="48">Jacksonaster</emphasis>
</taxonomicName>
sp. 2 were collected from a different locality to
<taxonomicName id="4C294D21FF85895AFB71F80076629EA1" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[1252,1398,1972,1997]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="47" pageNumber="48" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF85895AFB71F80076629EA1" box="[1252,1398,1972,1997]" italics="true" pageId="47" pageNumber="48">Jacksonaster</emphasis>
</taxonomicName>
sp. 1. They can be distinguished from
<taxonomicName id="4C294D21FF85895AFDB4F86371A79E9C" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[545,691,2007,2032]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="47" pageNumber="48" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF85895AFDB4F86371A79E9C" box="[545,691,2007,2032]" italics="true" pageId="47" pageNumber="48">Jacksonaster</emphasis>
</taxonomicName>
sp.
<quantity id="4CD19B47FF85895AFD76F86C701B9E9C" box="[739,783,2007,2032]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="48" unit="in" value="1.0">1 in</quantity>
having a more elliptical, rounded test outline, thicker test margin, relatively larger petaloid area, relatively closed petals, wider interporiferous zones (
<figureCitation id="13122A27FF9A8945FB5FFF23762799DC" box="[1226,1331,151,176]" captionStart="FIGURE 40" captionStartId="47.[151,250,1098,1120]" captionTargetBox="[151,1436,193,1077]" captionTargetId="figure@47.[151,1436,193,1077]" captionTargetPageId="47" captionText="FIGURE 40. Comparison of apical system and petaloid area of Jacksonaster sp. 1 (A, WUSL/EI/26) and Jacksonaster sp. 2 (B, WUSL/EI/29)." httpUri="https://zenodo.org/record/2617537/files/figure.png" pageId="48" pageNumber="49">Fig. 40B</figureCitation>
) and the periproct situated further from the posterior margin.
</paragraph>
<paragraph id="8B9636A2FF9A8945FF52FF6B7074982C" blockId="48.[151,1436,151,429]" pageId="48" pageNumber="49">
Using the key of
<bibRefCitation id="EFB84B53FF9A8945FE00FF6B71BD9994" author="Clark, A. M. G. &amp; Rowe, F. W. E." box="[405,681,223,248]" pageId="48" pageNumber="49" refId="ref34228" refString="Clark, A. M. G. &amp; Rowe, F. W. E. (1971) Monograph of Shallow-water Indo-West Pacific Echinoderms. British Museum Natural History Publications, London, 238 pp." type="book" year="1971">Clark and Rowe (1971)</bibRefCitation>
, these specimens resemble
<taxonomicName id="4C294D21FF9A8945FC7CFF6B77E99994" baseAuthorityName="L. Agassiz" baseAuthorityYear="1841" box="[1001,1277,223,248]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="B95DEAB0FF9A8945FC7CFF6B77E99994" box="[1001,1277,223,248]" italics="true" pageId="48" pageNumber="49">Jacksonaster depressum</emphasis>
</taxonomicName>
. However, as stated in
<bibRefCitation id="EFB84B53FF9A8945FE9FFEB172FF9871" author="Mortensen, T." box="[266,491,260,285]" pageId="48" pageNumber="49" refId="ref35386" refString="Mortensen, T. (1948 b) A Monograph of the Echinoidea IV. 2. Clypeasteroida. Clypeasteridae, Arachnoidae, Fibulariidae, Laganidae and Scutellidae. C. A. Reitzel, Copenhagen, 471 pp." type="book" year="1948">Mortensen (1948b)</bibRefCitation>
, the posterior end of
<taxonomicName id="4C294D21FF9A8945FD69FEB077009871" baseAuthorityName="L. Agassiz" baseAuthorityYear="1841" box="[764,1044,260,285]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="depressum">
<emphasis id="B95DEAB0FF9A8945FD69FEB077009871" box="[764,1044,260,285]" italics="true" pageId="48" pageNumber="49">Jacksonaster depressum</emphasis>
</taxonomicName>
is usually “reenteringly curved” (indented). This feature was not observed in
<taxonomicName id="4C294D21FF9A8945FD19FE93700A982C" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[652,798,295,320]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF9A8945FD19FE93700A982C" box="[652,798,295,320]" italics="true" pageId="48" pageNumber="49">Jacksonaster</emphasis>
</taxonomicName>
sp. 2.
</paragraph>
<paragraph id="8B9636A2FF9A8945FF52FEF872AA98C1" blockId="48.[151,1436,151,429]" pageId="48" pageNumber="49">
These specimens are similar in outline to
<taxonomicName id="4C294D21FF9A8945FD32FEF9707D9809" authorityName="Koehler" authorityYear="1922" box="[679,873,332,357]" class="Echinoidea" family="Laganidae" genus="Laganum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="joubini">
<emphasis id="B95DEAB0FF9A8945FD32FEF9707D9809" box="[679,873,332,357]" italics="true" pageId="48" pageNumber="49">Laganum joubini</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C294D21FF9A8945FC33FEF977019809" authorityName="Koehler" authorityYear="1922" box="[934,1045,332,357]" class="Echinoidea" family="Laganidae" genus="Laganum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="species" species="joubini">
<emphasis id="B95DEAB0FF9A8945FC33FEF977019809" box="[934,1045,332,357]" italics="true" pageId="48" pageNumber="49">L. joubini</emphasis>
</taxonomicName>
has a more anterior periproct and more anterior apical system.
<taxonomicName id="4C294D21FF9A8945FE49FEDB717A98E4" authorityName="Lambert in Lambert &amp; Thiery" authorityYear="1914" box="[476,622,367,392]" class="Echinoidea" family="Laganidae" genus="Jacksonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Clypeasteroida" pageId="48" pageNumber="49" phylum="Echinodermata" rank="genus">
<emphasis id="B95DEAB0FF9A8945FE49FEDB717A98E4" box="[476,622,367,392]" italics="true" pageId="48" pageNumber="49">Jacksonaster</emphasis>
</taxonomicName>
in
<collectingCountry id="F33E7632FF9A8945FD06FEDB701598E4" box="[659,769,367,392]" name="Sri Lanka" pageId="48" pageNumber="49">Sri Lanka</collectingCountry>
requires further examination and revision, perhaps through future molecular analyses.
</paragraph>
</subSubSection>
</treatment>
</document>