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<document id="FF2E6ECF15F5A84EC3A343C3A5EF9B39" ID-DOI="10.5281/zenodo.175929" ID-GBIF-Dataset="d280489c-4168-4888-ac6d-ea85f3b362e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="175929" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459824818497" checkinUser="plazi" docAuthor="Dijkstra, Klaas-Douwe B." docDate="2007" docId="0380878FFF8D9B44FF0AF8CFFB59FDE6" docLanguage="en" docName="zt01438p048.pdf" docOrigin="Zootaxa 1438" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hemicordulia atrovirens Dijkstra, 2007, n. sp." docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFB9FFF7FF8A9B4DFF9DFFCFFFF0FF8C" masterDocTitle="Gone with the wind: westward dispersal across the Indian Ocean and island speciation in Hemicordulia dragonflies (Odonata: Corduliidae)" masterLastPageNumber="48" masterPageNumber="27" pageNumber="34" updateTime="1698222116279" updateUser="plazi">
<mods:mods id="72AF81AC84F06FB641F6B80869B7ABEA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="84B0910A21B62B54AD61BDEB3B6B3958">Gone with the wind: westward dispersal across the Indian Ocean and island speciation in Hemicordulia dragonflies (Odonata: Corduliidae)</mods:title>
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<mods:name id="3A4ED680BB414972B910CCD503C0BEB7" type="personal">
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<mods:roleTerm id="317612E55AE831ABB74FA80A0C9E320E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="604AF9523952D93B878B5D4DDE30AA2E">Dijkstra, Klaas-Douwe B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F1DC6447579467764E42807C0DCDAE1D">text</mods:typeOfResource>
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<mods:title id="D9EFA044BA9F2A53E3CD4870E0530876">Zootaxa</mods:title>
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<mods:part id="C417C3B150E11D5E32965574BAB3DD02">
<mods:date id="62D767C6E7C18AB76914E05CD4432B04">2007</mods:date>
<mods:detail id="63C4B8F707CE03AA91B5118251237EB1" type="volume">
<mods:number id="4E11123FAC73B4EBD3890E7933626D87">1438</mods:number>
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<mods:classification id="298DC9EC611C203C8E75AA595734A95F">journal article</mods:classification>
<mods:identifier id="7761090DEA8CCA8E050C4E2CC89A790E" type="DOI">10.5281/zenodo.175929</mods:identifier>
<mods:identifier id="B00C9172384956AD9905720CE6BAB081" type="GBIF-Dataset">d280489c-4168-4888-ac6d-ea85f3b362e7</mods:identifier>
<mods:identifier id="07DFA984798DC10EBCE12212FD3FC911" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C34032CDBA88E6430BD7DC7E76B000A2" type="Zenodo-Dep">175929</mods:identifier>
</mods:mods>
<treatment id="0380878FFF8D9B44FF0AF8CFFB59FDE6" ID-DOI="http://doi.org/10.5281/zenodo.5688753" ID-GBIF-Taxon="119356531" ID-Zenodo-Dep="5688753" LSID="urn:lsid:plazi:treatment:0380878FFF8D9B44FF0AF8CFFB59FDE6" httpUri="http://treatment.plazi.org/id/0380878FFF8D9B44FF0AF8CFFB59FDE6" lastPageId="9" lastPageNumber="36" pageId="7" pageNumber="34">
<subSubSection id="C3336512FF8D9B4AFF0AF8CFFEC0F8B0" pageId="7" pageNumber="34" type="nomenclature">
<paragraph id="8B963699FF8D9B4AFF0AF8CFFDF9F896" blockId="7.[151,521,1792,1852]" box="[151,521,1792,1818]" pageId="7" pageNumber="34">
<heading id="D0DE81F5FF8D9B4AFF0AF8CFFDF9F896" bold="true" box="[151,521,1792,1818]" fontSize="11" level="1" pageId="7" pageNumber="34" reason="1">
<emphasis id="B95DEA8BFF8D9B4AFF0AF8CFFDF9F896" bold="true" box="[151,521,1792,1818]" pageId="7" pageNumber="34">
<taxonomicName id="4C294D1AFF8D9B4AFF0AF8CFFE30F896" ID-CoL="3KMPL" box="[151,448,1792,1818]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="atrovirens" status="sp. nov.">
<emphasis id="B95DEA8BFF8D9B4AFF0AF8CFFE30F896" bold="true" box="[151,448,1792,1818]" italics="true" pageId="7" pageNumber="34">Hemicordulia atrovirens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57F0FF8D9B4AFE5AF8CFFDF9F896" box="[455,521,1792,1818]" pageId="7" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B963699FF8D9B4AFF0AF8EDFEC0F8B0" blockId="7.[151,521,1792,1852]" box="[151,304,1826,1852]" pageId="7" pageNumber="34">
<figureCitation id="13122A1CFF8D9B4AFF0AF8EDFF0FF8B0" box="[151,255,1826,1852]" captionStart="FIGURES 2 – 11. 2 – 6" captionStartId="5.[151,269,1923,1947]" captionTargetBox="[264,1285,480,1866]" captionTargetId="figure@5.[264,1323,469,1891]" captionTargetPageId="5" captionText="FIGURES 2 – 11. 2 – 6, Male appendages of Hemicordulia species in dorsal view. (2) H. asiatica; (3) H. virens; (4) H. atrovirens n. sp.; (5) H. similis; (6) H. africana n. sp.; 7 – 11, Male appendages of Hemicordulia species in lateral view. (7) H. asiatica; (8) H. virens; (9) H. atrovirens n. sp.; (10) H. similis; (11) H. africana n. sp." httpUri="https://zenodo.org/record/175931/files/figure.png" pageId="7" pageNumber="34">Figs 4, 9</figureCitation>
,
<figureCitation id="13122A1CFF8D9B4AFE96F8EDFEDCF8B0" box="[267,300,1826,1852]" captionStart="FIGURES 12 – 17" captionStartId="7.[151,269,1642,1666]" captionTargetBox="[265,1323,403,1602]" captionTargetId="figure@7.[263,1323,356,1636]" captionTargetPageId="7" captionText="FIGURES 12 – 17. Vulvar scale and S 9 of Hemicordulia species in ventral view. (12) H. asiatica; (13) H. virens; (14) H. atrovirens n. sp.; (15) aberrant H. similis (cf. Fraser 1949); (16) typical H. similis; (17) H. africana n. sp." httpUri="https://zenodo.org/record/175932/files/figure.png" pageId="7" pageNumber="34">14</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF8D9B4AFF0AF8BCFD6FF846" pageId="7" pageNumber="34" type="reference_group">
<paragraph id="8B963699FF8D9B4AFF0AF8BCFC23F806" blockId="7.[151,1436,1907,1994]" box="[151,979,1907,1930]" pageId="7" pageNumber="34">
<treatmentCitationGroup id="AB3911B7FF8D9B4AFF0AF8BCFC23F806" box="[151,979,1907,1930]" pageId="7" pageNumber="34">
<taxonomicName id="4C294D1AFF8D9B4AFF0AF8BCFDB6F806" authority="Selys, 1878" authorityName="Selys" authorityYear="1878" box="[151,582,1907,1930]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="asiatica" subSpecies="nec">
<emphasis id="B95DEA8BFF8D9B4AFF0AF8BCFE41F806" box="[151,433,1907,1930]" italics="true" pageId="7" pageNumber="34">Hemicordulia asiatica nec</emphasis>
(
<bibRefCitation id="EFB84B68FF8D9B4AFE22F8BCFDCEF806" author="Selys" box="[447,574,1907,1930]" pageId="7" pageNumber="47" refString="Selys-Longchamps, E. de (1878) Secondes additions au Synopsis des Cordulines. Bulletins de l'Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique (2), 45, 183 - 222 (bulletin); 5 - 44 (reprint)." type="journal article" year="1878">Selys, 1878</bibRefCitation>
)
</taxonomicName>
–
<treatmentCitation id="0A881088FF8D9B4AFDFDF8BCFC3FF806" author="Couteyen" box="[608,975,1907,1930]" page="107" pageId="7" pageNumber="34" year="2000">
<bibRefCitation id="EFB84B68FF8D9B4AFDFDF8BCFC3FF806" author="Couteyen" box="[608,975,1907,1930]" pageId="7" pageNumber="45" refString="Couteyen, S. &amp; Papazian, M. (2000) Contribution a la connaissance des Odonates de la Reunion 3. Hemicordulia asiatica Selys, 1878, une espece nouvelle pour l'ile (Odonata: Corduliidae). Martinia, 16, 107 - 110." type="journal article" year="2000">Couteyen &amp; Papazian (2000: 107)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B963699FF8D9B4AFF0AF85CFD6FF846" blockId="7.[151,1436,1907,1994]" pageId="7" pageNumber="34">
<taxonomicName id="4C294D1AFF8D9B4AFF0AF85CFE51F826" ID-CoL="3KMPL" box="[151,417,1939,1962]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="atrovirens" status="sp. nov.">
<emphasis id="B95DEA8BFF8D9B4AFF0AF85CFE51F826" box="[151,417,1939,1962]" italics="true" pageId="7" pageNumber="34">Hemicordulia atrovirens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E57F0FF8D9B4AFE30F85DFE1EF826" box="[429,494,1938,1962]" pageId="7" pageNumber="34" rank="species">
<emphasis id="B95DEA8BFF8D9B4AFE30F85DFE1EF826" bold="true" box="[429,494,1938,1962]" pageId="7" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
<typeStatus id="5492883BFF8D9B4AFE67F85CFDADF826" box="[506,605,1939,1962]" pageId="7" pageNumber="34" type="holotype">Holotype</typeStatus>
♂ and
<typeStatus id="5492883BFF8D9B4AFD24F85CFCE5F826" box="[697,789,1939,1962]" pageId="7" pageNumber="34" type="paratype">paratype</typeStatus>
Ψ:
<collectingCountry id="F33E7609FF8D9B4AFCD9F85CFC51F826" box="[836,929,1939,1962]" name="Réunion" pageId="7" pageNumber="34">Réunion</collectingCountry>
, Rivière de Sainte-Suzanne,
<quantity id="4CD19B7CFF8D9B4AFB74F85CFAC1F826" box="[1257,1329,1939,1962]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="34" unit="m" value="730.0">730 m</quantity>
a.s.l.,
<date id="FF971059FF8D9B4AFA1CF85CFEDCF846" pageId="7" pageNumber="34" value="1996-04-08" valueMax="1996-04-09" valueMin="1996-04-08">8– 9.iv.1996</date>
, A. Martens (RMNH) [examined].
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF829B45FF0AFF57FC7AFE46" pageId="8" pageNumber="35" type="materials_examined">
<paragraph id="8B963699FF829B45FF0AFF57FC7AFE46" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF0AFF57FE9CFF3D" box="[151,364,152,177]" italics="true" pageId="8" pageNumber="35">Further material:</emphasis>
<collectingCountry id="F33E7609FF829B45FEE5FF57FDF1FF3E" box="[376,513,152,178]" name="Réunion" pageId="8" pageNumber="35">RÉUNION</collectingCountry>
: 1 Ψ (Coll. Martin), “
<collectingCountry id="F33E7609FF829B45FC8EFF57FC8CFF3E" box="[787,892,152,178]" name="Réunion" pageId="8" pageNumber="35">Réunion</collectingCountry>
” (
<collectionCode id="ED38AE5CFF829B45FC06FF57FC0BFF3E" LSID="urn:lsid:biocol.org:col:34988" box="[923,1019,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="35">MNHN</collectionCode>
); 2 ď, Bras Cabot,
<quantity id="4CD19B7CFF829B45FB69FF57FAA2FF3E" box="[1268,1362,152,178]" metricMagnitude="3" metricUnit="m" metricValue="1.325" pageId="8" pageNumber="35" unit="m" value="1325.0">1325 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FF0AFF0FFEF6FF56" box="[151,262,192,218]" pageId="8" pageNumber="35" value="1996-04-04">4.iv.1996</date>
, A. Martens (Coll. A. Martens); 2 ď, Rivière des Marsouins,
<quantity id="4CD19B7CFF829B45FC71FF0FFBB7FF56" box="[1004,1095,192,218]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="8" pageNumber="35" unit="m" value="1330.0">1330 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FB05FF0FFAF7FF56" box="[1176,1287,192,218]" pageId="8" pageNumber="35" value="1996-04-04">4.iv.1996</date>
, A. Martens (Coll. A. Martens); 3 ď, ravine near Rivière Langevin,
<quantity id="4CD19B7CFF829B45FC85FF27FC93FE8E" box="[792,867,232,258]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="35" unit="m" value="350.0">350 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FC03FF27FBD6FE8E" box="[926,1062,232,258]" pageId="8" pageNumber="35" value="1996-04-14">14.iv.1996</date>
, A. Martens (Coll. A. Martens); 1 ď, Chemin-de-Ceinture, forêt de ravenales,
<date id="FF971059FF829B45FD24FEDFFCC3FEA6" box="[697,819,272,298]" pageId="8" pageNumber="35" value="1999-02-12">12.ii.1999</date>
, J. &amp; Cl. Pierre (
<collectionCode id="ED38AE5CFF829B45FB9CFEDFFB91FEA6" LSID="urn:lsid:biocol.org:col:34988" box="[1025,1121,272,298]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="35">MNHN</collectionCode>
); 1 Ψ, commune de Saint- Philippe, Ravine Basse Vallée, altitude
<quantity id="4CD19B7CFF829B45FDF7FEF7FD46FEDE" box="[618,694,312,338]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="35" unit="m" value="280.0">280 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FC98FEF7FC8BFEDE" box="[773,891,312,338]" pageId="8" pageNumber="35" value="1999-05-22">22.v.1999</date>
, S. Couteyen (Coll. M. Papazian); 1 ď, Saint- Paul, old watermill,
<quantity id="4CD19B7CFF829B45FE15FEAFFE46FEF6" box="[392,438,352,378]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="35" unit="m" value="2.0">2 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FD98FEAFFD9CFEF6" box="[517,620,352,378]" pageId="8" pageNumber="35" value="2000-05-01">1.v.2000</date>
, A. Martens (Coll. A. Martens); 2 ď, Saint-Benoit, Bras Cabot,
<quantity id="4CD19B7CFF829B45FAFFFEAFFF5EFE2E" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="35" unit="m" value="1100.0">1100 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FF67FE47FEADFE2E" box="[250,349,392,418]" pageId="8" pageNumber="35" value="2001-12">xii.2001</date>
, S. Couteyen (Coll. M. Papazian); 1 ď, La Plaine des Palmistes, Forêt de Bébour, Rivière des Marsouins,
<quantity id="4CD19B7CFF829B45FEBCFE7FFE8AFE46" box="[289,378,432,458]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="8" pageNumber="35" unit="m" value="1330.0">1330 m</quantity>
a.s.l.,
<date id="FF971059FF829B45FE5BFE7FFDB0FE46" box="[454,576,432,458]" pageId="8" pageNumber="35" value="2003-02-14">14.ii.2003</date>
, D. Grand (Coll. D. Grand).
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF829B45FF5BFE18FD48FD96" pageId="8" pageNumber="35" type="diagnosis">
<paragraph id="8B963699FF829B45FF5BFE18FD48FD96" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF5BFE18FEB6FE7D" bold="true" box="[198,326,471,497]" pageId="8" pageNumber="35">Diagnosis.</emphasis>
Largest and darkest species, with the most extensive and deepest metallic green markings of all species under consideration (see
<taxonomicName id="4C294D1AFF829B45FDA0FDCFFD57FD95" box="[573,679,512,537]" class="Insecta" family="Corduliidae" genus="Cordulia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="B95DEA8BFF829B45FDA0FDCFFD57FD95" box="[573,679,512,537]" italics="true" pageId="8" pageNumber="35">H. virens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF829B45FF5BFDE8FEE1F97E" pageId="8" pageNumber="35" type="description">
<paragraph id="8B963699FF829B45FF5BFDE8FB5DFAEE" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF5BFDE8FEAFFDCD" bold="true" box="[198,351,551,577]" pageId="8" pageNumber="35">Description.</emphasis>
<typeStatus id="5492883BFF829B45FEF5FDE7FE25FDCE" box="[360,469,552,578]" pageId="8" pageNumber="35" type="holotype">Holotype</typeStatus>
male. Measurements (mm): entire length: 51, abdomen length (without appendages): 34, Fw length: 35, Hw length: 34.5, Fw Pt: 2.0. Labium beige, darker anteriorly (discoloured?). Mandibles, genae, labrum, clypeus and narrow area on lower frons brownish yellow; darker on edges. Vertex and dorsum of frons deep metallic green-blue, merging via a deep brown band into yellow area on antefrons (latter is not visible in dorsal view). Antennae, occipital triangle, occiput and postgenae black; paired rounded swellings on occiput lower than in
<taxonomicName id="4C294D1AFF829B45FE6CFD3FFD87FC85" box="[497,631,752,777]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B95DEA8BFF829B45FE6CFD3FFD87FC85" box="[497,631,752,777]" italics="true" pageId="8" pageNumber="35">H. africana</emphasis>
</taxonomicName>
. Labrum, clypeus, frons, vertex and occipital triangle with long bristly black hairs. Labium, genae and occiput with finer whitish hairs. Prothorax dark brown. Synthorax deep metallic green with weakly contrasting brownish yellow areas on anterior half of mesepisternum and centres of mesepimeron, metepisternum and metepimeron: metallic areas are much wider than the pale areas between them making entire synthorax almost uniformly green. Mesokatepisternum, metakatepisternum and synthoracic venter brownish yellow with metallic green lustre; poststernum posteriorly dark. Synthorax covered with dense pale long hairs, especially long on mesepisternum; shorter, darker and denser hairs on antealar sinus. Legs black save for yellow coxae and a hint at base of fore femora. Keels present on anterior face of apical half of fore tibiae and almost whole length of hind tibiae (just falling short of their bases), but absent on middle tibiae. Venation blackish. Wing membrane evenly but very lightly tinted brown, tinged yellower at extreme bases. Membranule dark brown, slightly paler at extreme base. Pt dark brown. Venation as for genus. 8 Ax in both Fw,
<quantity id="4CD19B7CFF829B45FEFCFB67FE7DFB4E" box="[353,397,1192,1218]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="35" unit="in" value="6.0">6 in</quantity>
Hw. 7–8 Px in Fw,
<quantity id="4CD19B7CFF829B45FDEFFB67FD6FFB4E" box="[626,671,1192,1218]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="35" unit="in" value="9.0">9 in</quantity>
Hw. Fw and Hw triangles with single cross-veins. Discoidal field of 2 rows of cells at base. Anal loops of 18 cells. Abdomen black with green- and blue-purple gloss, tergites unmarked except for an indistinct brownish yellow lateral spot on S2, S2–3 intersegmental ring and narrow streaks on lateral carinae S3. Sternites black. Cerci and epiproct slender, black (
<figureCitation id="13122A1CFF829B45FBD2FAEFFB4DFAB6" box="[1103,1213,1312,1338]" captionStart="FIGURES 2 – 11. 2 – 6" captionStartId="5.[151,269,1923,1947]" captionTargetBox="[264,1285,480,1866]" captionTargetId="figure@5.[264,1323,469,1891]" captionTargetPageId="5" captionText="FIGURES 2 – 11. 2 – 6, Male appendages of Hemicordulia species in dorsal view. (2) H. asiatica; (3) H. virens; (4) H. atrovirens n. sp.; (5) H. similis; (6) H. africana n. sp.; 7 – 11, Male appendages of Hemicordulia species in lateral view. (7) H. asiatica; (8) H. virens; (9) H. atrovirens n. sp.; (10) H. similis; (11) H. africana n. sp." httpUri="https://zenodo.org/record/175931/files/figure.png" pageId="8" pageNumber="35">Figs 4, 9</figureCitation>
). Cerci with thick, blunt tips. Secondary genitalia similar to
<taxonomicName id="4C294D1AFF829B45FDE4FA87FCF0FAED" box="[633,768,1352,1377]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B95DEA8BFF829B45FDE4FA87FCF0FAED" box="[633,768,1352,1377]" italics="true" pageId="8" pageNumber="35">H. africana</emphasis>
</taxonomicName>
(cf.
<figureCitation id="13122A1CFF829B45FCAAFA87FC8AFAEE" box="[823,890,1352,1378]" captionStart="FIGURE 1" captionStartId="4.[151,254,2011,2035]" captionTargetBox="[456,1126,1707,1972]" captionTargetId="figure@4.[456,1130,1707,2001]" captionTargetPageId="4" captionText="FIGURE 1. Secondary genitalia of Hemicordulia africana n. sp. in lateral view." httpUri="https://zenodo.org/record/175930/files/figure.png" pageId="8" pageNumber="35">Fig. 1</figureCitation>
), hamule blackish brown.
</paragraph>
<paragraph id="8B963699FF829B45FF5BFABFFF3CF92E" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
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female. Measurements (mm): entire length: 55, abdomen length (without appendages): 39, Fw length: 39, Hw length: 38, Fw Pt: 2.5. Larger and slightly paler than
<typeStatus id="5492883BFF829B45FC4EFA57FBCEFA3E" box="[979,1086,1432,1458]" pageId="8" pageNumber="35" type="holotype">holotype</typeStatus>
. Paired rounded swellings on occiput more strongly swollen than in male, only with long pale hairs. Pale areas on synthorax larger and brighter, appearing as distinct spots. Fore femora ventrally largely pale. Basal yellow in wings more extensive than in male, to Cux and almost to Ax
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Fw, and just beyond Cux and to Ax
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Hw. 8–9 Ax in Fw,
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Hw. 7 Px in Fw,
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Hw. Anal loops of 19–21 cells. Vulvar scale appressed, dark brown, extending over less than a third of sternite S9, semi-circular with deep cleft (
<figureCitation id="13122A1CFF829B45FC9DF9AFFCAAF9F6" box="[768,858,1632,1658]" captionStart="FIGURES 12 – 17" captionStartId="7.[151,269,1642,1666]" captionTargetBox="[265,1323,403,1602]" captionTargetId="figure@7.[263,1323,356,1636]" captionTargetPageId="7" captionText="FIGURES 12 – 17. Vulvar scale and S 9 of Hemicordulia species in ventral view. (12) H. asiatica; (13) H. virens; (14) H. atrovirens n. sp.; (15) aberrant H. similis (cf. Fraser 1949); (16) typical H. similis; (17) H. africana n. sp." httpUri="https://zenodo.org/record/175932/files/figure.png" pageId="8" pageNumber="35">Fig. 14</figureCitation>
). Cerci black, long and slender, 2.5x longer than S10.
</paragraph>
<paragraph id="8B963699FF829B45FF5BF960FEE1F97E" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF5BF960FEB1F945" bold="true" box="[198,321,1711,1737]" pageId="8" pageNumber="35">Variation.</emphasis>
Limited. Pale markings are slightly more extensive in females; teneral specimens were not examined.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF829B44FF5BF930FAA6FE7E" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="etymology">
<paragraph id="8B963699FF829B45FF5BF930FD9BF8CE" blockId="8.[151,1437,151,2018]" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF5BF930FEA3F895" bold="true" box="[198,339,1791,1817]" pageId="8" pageNumber="35">Etymology.</emphasis>
The Latin
<taxonomicName id="4C294D1AFF829B45FE4BF8CFFDBBF895" box="[470,587,1792,1817]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="atrovirens">
<emphasis id="B95DEA8BFF829B45FE4BF8CFFDBBF895" box="[470,587,1792,1817]" italics="true" pageId="8" pageNumber="35">atrovirens</emphasis>
</taxonomicName>
(being dark green) emphasises both the close relationship and the main difference with its Mauritian neighbour.
</paragraph>
<paragraph id="8B963699FF829B44FF5BF880FAA6FE7E" blockId="8.[151,1437,151,2018]" lastBlockId="9.[151,1436,152,618]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
<emphasis id="B95DEA8BFF829B45FF5BF880FE47F8E5" bold="true" box="[198,439,1871,1897]" pageId="8" pageNumber="35">Range and ecology.</emphasis>
Endemic to
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. The larval habitat has been said to range from torrents to stagnant water under closed cover, but is probably principally pools and calm sections of shady fast-flowing streams (
<bibRefCitation id="EFB84B68FF829B45FF62F86FFDB4F836" author="Couteyen" box="[255,580,1952,1978]" pageId="8" pageNumber="45" refString="Couteyen, S. &amp; Papazian, M. (2000) Contribution a la connaissance des Odonates de la Reunion 3. Hemicordulia asiatica Selys, 1878, une espece nouvelle pour l'ile (Odonata: Corduliidae). Martinia, 16, 107 - 110." type="journal article" year="2000">Couteyen &amp; Papazian 2000</bibRefCitation>
;
<bibRefCitation id="EFB84B68FF829B45FDCDF86FFD2EF836" author="Grand" box="[592,734,1952,1978]" pageId="8" pageNumber="46" refString="Grand, D. (2004) Compte rendu odonatologique d'un voyage a l'ile de la Reunion. Martinia, 20, 67 - 75." type="journal article" year="2004">Grand 2004</bibRefCitation>
; A. Martens in litt.).
<taxonomicName id="4C294D1AFF829B45FC4FF86FFB9BF835" box="[978,1131,1952,1977]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="atrovirens">
<emphasis id="B95DEA8BFF829B45FC4FF86FFB9BF835" box="[978,1131,1952,1977]" italics="true" pageId="8" pageNumber="35">H. atrovirens</emphasis>
</taxonomicName>
seldom cohabits with any of the ten other anisopterans found on
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(
<bibRefCitation id="EFB84B68FF829B45FD4DF807FC7AF86E" author="Couteyen" box="[720,906,1992,2018]" pageId="8" pageNumber="45" refString="Couteyen, S. (2006) Etude de l'explotation des sites de reproduction par les Anisopteres a l'ile de la Reunion (Odonata). Bulletin de la Societe entomologique de France, 111, 65 - 71." type="journal article" year="2006">Couteyen 2006</bibRefCitation>
): overlap (number of sites where two species co-occur as percentage of sites where either or both occur) with each of the nine species shared with mainland Africa was at most 9%, while that with
<taxonomicName id="4C294D1AFF839B44FDF5FF0FFCA4FF55" box="[616,852,192,217]" class="Insecta" family="Aeshnidae" genus="Gynacantha" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="bispina">
<emphasis id="B95DEA8BFF839B44FDF5FF0FFCA4FF55" box="[616,852,192,217]" italics="true" pageId="9" pageNumber="36">Gynacantha bispina</emphasis>
</taxonomicName>
, the only other Mascarene endemic on the island, was 12%. Moreover, within its usual altitudinal range of
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(one record provided here is near sea level), below
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only forested habitats were occupied. Because all nine widespread species avoid forest and six are confined to lower altitudes,
<bibRefCitation id="EFB84B68FF839B44FDECFEF7FCC8FEDE" author="Couteyen" box="[625,824,312,338]" pageId="9" pageNumber="45" refString="Couteyen, S. (2006) Etude de l'explotation des sites de reproduction par les Anisopteres a l'ile de la Reunion (Odonata). Bulletin de la Societe entomologique de France, 111, 65 - 71." type="journal article" year="2006">Couteyen (2006)</bibRefCitation>
suggested that both endemics are out-competed by the mainland species. However, the distribution may also be explained by different habitat preferences (A. Martens in litt.). Males fly along streams, both under forest cover and in full sun. In cloudy weather and towards dusk, adults appear in clearings and on the forest edge, making prolonged flights close to the vegetation (
<bibRefCitation id="EFB84B68FF839B44FF49FE17FD14FE7E" author="Couteyen" box="[212,740,472,498]" pageId="9" pageNumber="45" refString="Couteyen, S. &amp; Papazian, M. (2000) Contribution a la connaissance des Odonates de la Reunion 3. Hemicordulia asiatica Selys, 1878, une espece nouvelle pour l'ile (Odonata: Corduliidae). Martinia, 16, 107 - 110." type="journal article" year="2000">Couteyen &amp; Papazian 2000; D. Grand in litt.</bibRefCitation>
). Adults have been observed from December to May.
</paragraph>
</subSubSection>
<subSubSection id="C3336512FF839B44FF5BFE30FB59FDE6" pageId="9" pageNumber="36" type="discussion">
<paragraph id="8B963699FF839B44FF5BFE30FB59FDE6" blockId="9.[151,1436,152,618]" pageId="9" pageNumber="36">
<emphasis id="B95DEA8BFF839B44FF5BFE30FECFFD95" bold="true" box="[198,319,511,537]" pageId="9" pageNumber="36">Remarks.</emphasis>
See
<taxonomicName id="4C294D1AFF839B44FEE0FDCFFE19FD95" box="[381,489,512,537]" class="Insecta" family="Corduliidae" genus="Cordulia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="B95DEA8BFF839B44FEE0FDCFFE19FD95" box="[381,489,512,537]" italics="true" pageId="9" pageNumber="36">H. virens</emphasis>
</taxonomicName>
. Paulson &amp; Buden (2003) observed increasing size of
<taxonomicName id="4C294D1AFF839B44FBE3FDCFFAAEFD96" authority="Asahina" authorityName="Asahina" box="[1150,1374,512,538]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="haluco">
<emphasis id="B95DEA8BFF839B44FBE3FDCFFB05FD95" box="[1150,1269,512,537]" italics="true" pageId="9" pageNumber="36">H. haluco</emphasis>
Asahina
</taxonomicName>
with increasing altitude on Pohnpei. The slight size difference seen between
<taxonomicName id="4C294D1AFF839B44FC72FDE7FB7BFDCD" box="[1007,1163,552,577]" class="Insecta" family="Corduliidae" genus="Hemicordulia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="atrovirens">
<emphasis id="B95DEA8BFF839B44FC72FDE7FB7BFDCD" box="[1007,1163,552,577]" italics="true" pageId="9" pageNumber="36">H. atrovirens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C294D1AFF839B44FB55FDE7FAC4FDCD" box="[1224,1332,552,577]" class="Insecta" family="Corduliidae" genus="Cordulia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="B95DEA8BFF839B44FB55FDE7FAC4FDCD" box="[1224,1332,552,577]" italics="true" pageId="9" pageNumber="36">H. virens</emphasis>
</taxonomicName>
, and the darker colour of the latter, may be linked to the greater elevation of the
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habitats.
</paragraph>
</subSubSection>
</treatment>
</document>
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