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<document id="71F9ACCFA16C7BA94494375A814C570D" ID-CLB-Dataset="28961" ID-DOI="10.11646/zootaxa.4461.4.8" ID-GBIF-Dataset="852c1d58-1843-4263-a5b6-b13da89e1463" ID-ISSN="1175-5326" ID-Zenodo-Dep="1460324" ID-ZooBank="DE244009-3CA3-44F3-8D91-4CD35823204D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539338061151" checkinUser="plazi" docAuthor="Gong, Xuyun, Martens, Jochen &amp; Zhang, Chao" docDate="2018" docId="03805233FF9AA41B9AC5FB1EFD19FC71" docLanguage="en" docName="zootaxa.4461.4.8.pdf" docOrigin="Zootaxa 4461 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Biantes spatulatus Gong &amp; Martens &amp; Zhang 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="597" masterDocId="FFB92A4BFF9CA4119A52FFA1FFE8FF9C" masterDocTitle="Two new species of Biantes from China and Malaysia (Opiliones: Laniatores: Biantidae)" masterLastPageNumber="599" masterPageNumber="587" pageNumber="593" updateTime="1698580085747" updateUser="ExternalLinkService">
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<mods:title id="6449CA20AA290D242F7DE264A6B55D7F">Two new species of Biantes from China and Malaysia (Opiliones: Laniatores: Biantidae)</mods:title>
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<mods:namePart id="A188D3E7042D2A708113733A58BBC244">Gong, Xuyun</mods:namePart>
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<mods:namePart id="FD08D0FE8B4A23C3C1F96FD7961AAA20">Martens, Jochen</mods:namePart>
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<mods:namePart id="807F93C85D3F8234CAE2882A2BA92FFC">Zhang, Chao</mods:namePart>
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<mods:title id="2D22C89AAD08235CF5E7C10C93D3D0C7">Zootaxa</mods:title>
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<mods:date id="84D79648057286E588C2301805D2C160">2018</mods:date>
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<mods:number id="8295A35C3CD4C4CCEA1519FDD62FFC06">2018-08-23</mods:number>
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<treatment id="03805233FF9AA41B9AC5FB1EFD19FC71" ID-DOI="http://doi.org/10.5281/zenodo.5950776" ID-GBIF-Taxon="148702193" ID-Zenodo-Dep="5950776" LSID="urn:lsid:plazi:treatment:03805233FF9AA41B9AC5FB1EFD19FC71" httpUri="http://treatment.plazi.org/id/03805233FF9AA41B9AC5FB1EFD19FC71" lastPageId="10" lastPageNumber="597" pageId="6" pageNumber="593">
<subSubSection id="C333B0AEFF9AA4179AC5FB1EFE5BFB67" pageId="6" pageNumber="593" type="nomenclature">
<paragraph id="8B96E325FF9AA4179AC5FB1EFE31FB46" blockId="6.[151,473,1215,1275]" box="[151,473,1215,1242]" pageId="6" pageNumber="593">
<heading id="D0DE5449FF9AA4179AC5FB1EFE31FB46" bold="true" box="[151,473,1215,1242]" fontSize="11" level="1" pageId="6" pageNumber="593" reason="1">
<emphasis id="B95D3F37FF9AA4179AC5FB1EFE31FB46" bold="true" box="[151,473,1215,1242]" pageId="6" pageNumber="593">
<taxonomicName id="4C2998A6FF9AA4179AC5FB1EFE9DFB45" ID-CoL="7SLR9" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[151,373,1215,1241]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="spatulatus" status="sp. nov.">
<emphasis id="B95D3F37FF9AA4179AC5FB1EFE9DFB45" bold="true" box="[151,373,1215,1241]" italics="true" pageId="6" pageNumber="593">BIantes spatulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF9AA4179B2EFB61FE31FB46" box="[380,473,1216,1242]" pageId="6" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B96E325FF9AA4179AC5FB43FE5BFB67" blockId="6.[151,473,1215,1275]" box="[151,435,1250,1275]" pageId="6" pageNumber="593">
(
<figureCitation id="1312FFA0FF9AA4179ACDFB43FEEBFB67" box="[159,259,1250,1275]" captionStart="FIGURES 69" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[233,1361,294,1812]" captionTargetId="figure@3.[206,1393,196,1882]" captionTargetPageId="3" captionText="FIGURES 69. Body of holotype and paratype, dorsal view. 67. Biantes calyptroideus sp. nov. 6. Male. 7. Female. 89. Biantes spatulatus sp. nov. 8. Male. 9. Female. Scale bars: 1mm (67); 0.5 mm (89)." httpUri="https://zenodo.org/record/1460328/files/figure.png" pageId="6" pageNumber="593">Figs 89</figureCitation>
,
<figureCitation id="1312FFA0FF9AA4179B5DFB42FEB1FB67" box="[271,345,1251,1275]" captionStart-0="FIGURES 2837" captionStart-1="FIGURES 3846" captionStartId-0="7.[151,264,1873,1896]" captionStartId-1="8.[151,264,1917,1940]" captionTargetBox-0="[237,1435,303,1819]" captionTargetBox-1="[270,1353,252,1894]" captionTargetId-0="figure@7.[163,1448,193,1884]" captionTargetId-1="figure@8.[220,1373,193,1905]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." captionText-1="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri-0="https://zenodo.org/record/1460334/files/figure.png" httpUri-1="https://zenodo.org/record/1460336/files/figure.png" pageId="6" pageNumber="593">2843</figureCitation>
,
<figureCitation id="1312FFA0FF9AA4179B31FB43FE43FB67" box="[355,427,1250,1275]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="6" pageNumber="593">5052</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333B0AEFF9AA4179AC5FA8BFEBDFA17" pageId="6" pageNumber="593" type="materials_examined">
<paragraph id="8B96E325FF9AA4179AC5FA8BFEBDFA17" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
<materialsCitation id="3B41E978FF9AA4179AC5FA8BFEB9FA17" ID-GBIF-Occurrence="1919493524" collectingDate="2015-10-24" collectorName="Z. Z. Gao" country="Malaysia" elevation="306" latitude="5.5333333" location="Temenggor Lake" longLatPrecision="1305" longitude="101.35" pageId="6" pageNumber="593" specimenCount="4" stateProvince="Perak" typeStatus="holotype">
<emphasis id="B95D3F37FF9AA4179AC5FA8BFE52FADF" bold="true" box="[151,442,1322,1347]" pageId="6" pageNumber="593">
<typeStatus id="54925D87FF9AA4179AC5FA8BFF3AFADF" box="[151,210,1322,1347]" pageId="6" pageNumber="593">Type</typeStatus>
material.
<typeStatus id="54925D87FF9AA4179B1FFA8BFE52FADF" box="[333,442,1322,1347]" pageId="6" pageNumber="593" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<collectingCountry id="F33EA3B5FF9AA417985CFA8AFD4CFADF" box="[526,676,1323,1347]" name="Malaysia" pageId="6" pageNumber="593">MALAYSIA</collectingCountry>
:
<collectingRegion id="49ED2DC7FF9AA41798FDFA8BFD1AFADF" box="[687,754,1322,1347]" country="Malaysia" name="Perak" pageId="6" pageNumber="593">Perak</collectingRegion>
,
<location id="8EF6B5FEFF9AA41798ACFA8AFC55FADF" LSID="urn:lsid:plazi:treatment:03805233FF9AA41B9AC5FB1EFD19FC71:8EF6B5FEFF9AA41798ACFA8AFC55FADF" box="[766,957,1322,1347]" country="Malaysia" latitude="5.5333333" longLatPrecision="1305" longitude="101.35" name="Temenggor Lake" pageId="6" pageNumber="593" stateProvince="Perak">Temenggor Lake</location>
[
<geoCoordinate id="EE1D85E2FF9AA4179999FA8BFBD8FADF" box="[971,1072,1322,1347]" direction="north" orientation="latitude" pageId="6" pageNumber="593" precision="925" value="5.5333333">N05°32´</geoCoordinate>
,
<geoCoordinate id="EE1D85E2FF9AA4179E6FFA8AFB5CFADF" box="[1085,1204,1323,1347]" direction="east" orientation="longitude" pageId="6" pageNumber="593" precision="925" value="101.35">E 101°21´</geoCoordinate>
], alt.
<elevation id="00040416FF9AA4179EA1FA8BFADEFADF" box="[1267,1334,1322,1347]" metricMagnitude="2" metricUnit="m" metricValue="3.06" pageId="6" pageNumber="593" unit="m" value="306.0">
<quantity id="4CD14EC0FF9AA4179EA1FA8BFADEFADF" box="[1267,1334,1322,1347]" metricMagnitude="2" metricUnit="m" metricValue="3.06" pageId="6" pageNumber="593" unit="m" value="306.0">306m</quantity>
</elevation>
,
<collectingDate id="EFD33C0DFF9AA4179F10FA8BFF13FAFB" pageId="6" pageNumber="593" value="2015-10-24">
<date id="FF97C5E5FF9AA4179F10FA8BFF13FAFB" pageId="6" pageNumber="593" value="2015-10-24">October 24, 2015</date>
</collectingDate>
,
<collectorName id="26DC86F3FF9AA4179B54FAEEFE83FAFB" box="[262,363,1359,1383]" pageId="6" pageNumber="593">Z.Z. Gao</collectorName>
leg. (MHBU-Opi-15ZC1112).
<emphasis id="B95D3F37FF9AA417989AFAEFFCD6FAFB" bold="true" box="[712,830,1358,1383]" pageId="6" pageNumber="593">
<typeStatus id="54925D87FF9AA417989AFAEFFCD1FAFB" box="[712,825,1358,1383]" pageId="6" pageNumber="593" type="paratype">Paratype</typeStatus>
:
</emphasis>
one female (MHBU-Opi-15ZC1113), same collecting data as
<typeStatus id="54925D87FF9AA4179AB9FAD3FEB9FA17" box="[235,337,1394,1419]" pageId="6" pageNumber="593" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C333B0AEFF9AA4179A95FA37FC15FA6B" pageId="6" pageNumber="593" type="diagnosis">
<paragraph id="8B96E325FF9AA4179A95FA37FC15FA6B" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
<emphasis id="B95D3F37FF9AA4179A95FA37FEA9FA33" bold="true" box="[199,321,1430,1455]" pageId="6" pageNumber="593">Diagnosis.</emphasis>
Patella of pedipalp with a meso-ventral setiferous tubercle, its length almost equal to the sum of the length of the tibia and tarsus. Conductor of penis smooth, without any apophyses; the tip conical, pointing downward; the titillator spoon-shaped; the ventral plate without spongy body.
</paragraph>
</subSubSection>
<subSubSection id="C333B0AEFF9AA4179A95F9A3FDA4F9FF" pageId="6" pageNumber="593" type="etymology">
<paragraph id="8B96E325FF9AA4179A95F9A3FDA4F9FF" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
<emphasis id="B95D3F37FF9AA4179A95F9A3FEA6F987" bold="true" box="[199,334,1538,1563]" pageId="6" pageNumber="593">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="4C2998A6FF9AA41799DFF9A2FBE8F987" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[909,1024,1539,1563]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="B95D3F37FF9AA41799DFF9A2FBE8F987" box="[909,1024,1539,1563]" italics="true" pageId="6" pageNumber="593">spatulatus</emphasis>
</taxonomicName>
” meaning “spoon-shaped”, referring to the shape of the titillator of the penis. It also alludes to the spathe in the aroid plant family surrounding the spadix inflorescence, which bears the flowers.
</paragraph>
</subSubSection>
<subSubSection id="C333B0AEFF9AA41B9A95F9CFFA6FFC54" lastPageId="10" lastPageNumber="597" pageId="6" pageNumber="593" type="description">
<paragraph id="8B96E325FF9AA4179A95F9CFFCDDF937" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
<emphasis id="B95D3F37FF9AA4179A95F9CFFDF7F91B" bold="true" box="[199,543,1646,1671]" pageId="6" pageNumber="593">
Description. Male (
<typeStatus id="54925D87FF9AA4179BFFF9CFFDFFF91B" box="[429,535,1646,1671]" pageId="6" pageNumber="593" type="holotype">holotype</typeStatus>
)
</emphasis>
: habitus as in
<figureCitation id="1312FFA0FF9AA4179894F9CFFCE5F91B" box="[710,781,1646,1671]" captionStart="FIGURES 69" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[233,1361,294,1812]" captionTargetId="figure@3.[206,1393,196,1882]" captionTargetPageId="3" captionText="FIGURES 69. Body of holotype and paratype, dorsal view. 67. Biantes calyptroideus sp. nov. 6. Male. 7. Female. 89. Biantes spatulatus sp. nov. 8. Male. 9. Female. Scale bars: 1mm (67); 0.5 mm (89)." httpUri="https://zenodo.org/record/1460328/files/figure.png" pageId="6" pageNumber="593">Figs 8</figureCitation>
,
<figureCitation id="1312FFA0FF9AA417994EF9CEFCD2F91B" box="[796,826,1647,1671]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="6" pageNumber="593">28</figureCitation>
,
<figureCitation id="1312FFA0FF9AA4179915F9CFFC79F91B" box="[839,913,1646,1671]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="6" pageNumber="593">5051</figureCitation>
. Coloration (
<figureCitation id="1312FFA0FF9AA4179E76F9CFFB41F91B" box="[1060,1193,1646,1671]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="6" pageNumber="593">Figs 5051</figureCitation>
): Entire body yellow, without reticulations, opisthosomal scutum light mahogany.
</paragraph>
<paragraph id="8B96E325FF9AA4179A95F916FDC8F8C3" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
Dorsum (
<figureCitation id="1312FFA0FF9AA4179B63F917FE92F953" box="[305,378,1718,1743]" captionStart="FIGURES 69" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[233,1361,294,1812]" captionTargetId="figure@3.[206,1393,196,1882]" captionTargetPageId="3" captionText="FIGURES 69. Body of holotype and paratype, dorsal view. 67. Biantes calyptroideus sp. nov. 6. Male. 7. Female. 89. Biantes spatulatus sp. nov. 8. Male. 9. Female. Scale bars: 1mm (67); 0.5 mm (89)." httpUri="https://zenodo.org/record/1460328/files/figure.png" pageId="6" pageNumber="593">Figs 8</figureCitation>
,
<figureCitation id="1312FFA0FF9AA4179BDAF917FE4FF953" box="[392,423,1718,1743]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="6" pageNumber="593">51</figureCitation>
). Dorsal scutum granular and trapezoid in shape, widest portion of body at scutal area V. Anterior margin of carapace with two shallow cheliceral sockets and five small tubercles laterally. Eyes widely separated from each other with circum-ocular tubercles, adjacent to the latero-posterior region of carapace; the distance between eyes shorter than the width of scutal area I. The mesotergal sulci inconspicuous. Scutum and free tergites without enlarged tubercles.
</paragraph>
<paragraph id="8B96E325FF9AA4179A95F8CAFAC3F857" blockId="6.[151,1437,1322,1995]" pageId="6" pageNumber="593">
Venter. Genital operculum a rounded pentagon (
<figureCitation id="1312FFA0FF9AA417988CF8CBFCD9F81F" box="[734,817,1898,1923]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="6" pageNumber="593">Fig. 35</figureCitation>
). Coxae tuberculate. Coxa I with conspicuous tubercles with setae on top (
<figureCitation id="1312FFA0FF9AA4179B3EF82FFE2AF83B" box="[364,450,1934,1959]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="6" pageNumber="593">Fig. 29</figureCitation>
). Coxae II, III and IV only with minute tubercles. Genital operculum with a few setatipped granules. Free sternites with a row of minute tubercles with setae on top. Spiracles clearly visible.
</paragraph>
<caption id="DF56B3ADFF9BA4169AC5F8F0FCCFF87F" httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="7" pageNumber="594" startId="7.[151,264,1873,1896]" targetBox="[237,1435,303,1819]" targetPageId="7">
<paragraph id="8B96E325FF9BA4169AC5F8F0FCCFF87F" blockId="7.[151,1436,1873,2019]" pageId="7" pageNumber="594">
<emphasis id="B95D3F37FF9BA4169AC5F8F0FEBBF8FB" bold="true" box="[151,339,1873,1896]" pageId="7" pageNumber="594">FIGURES 2837.</emphasis>
<taxonomicName id="4C2998A6FF9BA4169B0BF8F2FDFBF8F4" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[345,531,1874,1896]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="spatulatus" status="sp. nov.">
<emphasis id="B95D3F37FF9BA4169B0BF8F2FDFBF8F4" box="[345,531,1874,1896]" italics="true" pageId="7" pageNumber="594">Biantes spatulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF9BA416984BF8F3FD81F8FB" box="[537,617,1874,1895]" pageId="7" pageNumber="594" rank="species">
<emphasis id="B95D3F37FF9BA416984BF8F3FD81F8FB" bold="true" box="[537,617,1874,1895]" pageId="7" pageNumber="594">sp. nov.</emphasis>
</taxonomicNameLabel>
male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537).
</paragraph>
</caption>
<caption id="DF56B3ADFF94A4199AC5F8DCFEF4F86C" httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="8" pageNumber="595" startId="8.[151,264,1917,1940]" targetBox="[270,1353,252,1894]" targetPageId="8">
<paragraph id="8B96E325FF94A4199AC5F8DCFEF4F86C" blockId="8.[151,1436,1917,2032]" pageId="8" pageNumber="595">
<emphasis id="B95D3F37FF94A4199AC5F8DCFEBFF80F" bold="true" box="[151,343,1917,1940]" pageId="8" pageNumber="595">FIGURES 3846.</emphasis>
<taxonomicName id="4C2998A6FF94A4199B32F8DEFDF6F808" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[352,542,1918,1940]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="595" phylum="Arthropoda" rank="species" species="spatulatus" status="sp. nov.">
<emphasis id="B95D3F37FF94A4199B32F8DEFDF6F808" box="[352,542,1918,1940]" italics="true" pageId="8" pageNumber="595">Biantes spatulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF94A4199875F8DEFD93F808" box="[551,635,1919,1940]" pageId="8" pageNumber="595" rank="species">
<emphasis id="B95D3F37FF94A4199875F8DEFD93F808" bold="true" box="[551,635,1919,1940]" pageId="8" pageNumber="595">sp. nov.</emphasis>
</taxonomicNameLabel>
genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41.
<emphasis id="B95D3F37FF94A419982FF83FFD15F82F" box="[637,765,1949,1971]" italics="true" pageId="8" pageNumber="595">Pars distalis</emphasis>
of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146).
</paragraph>
</caption>
<caption id="DF56B3ADFF95A4189AC5F9D2FC96F898" httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="9" pageNumber="596" startId="9.[151,264,1651,1674]" targetBox="[151,1434,194,1631]" targetPageId="9">
<paragraph id="8B96E325FF95A4189AC5F9D2FC96F898" blockId="9.[151,1436,1651,1796]" pageId="9" pageNumber="596">
<emphasis id="B95D3F37FF95A4189AC5F9D2FEBCF915" bold="true" box="[151,340,1651,1674]" pageId="9" pageNumber="596">FIGURES 4752.</emphasis>
Photographs of male holotype and female paratype of
<taxonomicName id="4C2998A6FF95A418992CF9D4FBB2F916" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[894,1114,1652,1674]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="calyptroideus" status="sp. nov.">
<emphasis id="B95D3F37FF95A418992CF9D4FBB2F916" box="[894,1114,1652,1674]" italics="true" pageId="9" pageNumber="596">Biantes calyptroideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF95A4189E32F9D5FB58F915" box="[1120,1200,1652,1673]" pageId="9" pageNumber="596" rank="species">
<emphasis id="B95D3F37FF95A4189E32F9D5FB58F915" bold="true" box="[1120,1200,1652,1673]" pageId="9" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2998A6FF95A4189EB0F9D4FA74F916" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[1250,1436,1652,1674]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="spatulatus" status="sp. nov.">
<emphasis id="B95D3F37FF95A4189EB0F9D4FA74F916" box="[1250,1436,1652,1674]" italics="true" pageId="9" pageNumber="596">Biantes spatulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF95A4189AC5F932FF0FF934" box="[151,231,1683,1704]" pageId="9" pageNumber="596" rank="species">
<emphasis id="B95D3F37FF95A4189AC5F932FF0FF934" bold="true" box="[151,231,1683,1704]" pageId="9" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
. 4749.
<taxonomicName id="4C2998A6FF95A4189B6DF935FDF2F935" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[319,538,1683,1705]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="calyptroideus" status="sp. nov.">
<emphasis id="B95D3F37FF95A4189B6DF935FDF2F935" box="[319,538,1683,1705]" italics="true" pageId="9" pageNumber="596">Biantes calyptroideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF95A418984DF932FD87F934" box="[543,623,1683,1704]" pageId="9" pageNumber="596" rank="species">
<emphasis id="B95D3F37FF95A418984DF932FD87F934" bold="true" box="[543,623,1683,1704]" pageId="9" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052.
<taxonomicName id="4C2998A6FF95A4189E70F913FB34F95B" authority="Gong &amp; Martens &amp; Zhang, 2018" authorityName="Gong &amp; Martens &amp; Zhang" authorityYear="2018" box="[1058,1244,1713,1735]" class="Arachnida" family="Biantidae" genus="Biantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="596" phylum="Arthropoda" rank="species" species="spatulatus" status="sp. nov.">
<emphasis id="B95D3F37FF95A4189E70F913FB34F95B" box="[1058,1244,1713,1735]" italics="true" pageId="9" pageNumber="596">Biantes spatulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26E824CFF95A4189EB1F913FADCF95B" box="[1251,1332,1714,1735]" pageId="9" pageNumber="596" rank="species">
<emphasis id="B95D3F37FF95A4189EB1F913FADCF95B" bold="true" box="[1251,1332,1714,1735]" pageId="9" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052).
</paragraph>
</caption>
<paragraph id="8B96E325FF95A4189A95F890FB0BF80E" blockId="9.[151,1437,1841,2010]" pageId="9" pageNumber="596">
Chelicerae (
<figureCitation id="1312FFA0FF95A4189B1FF890FE27F8D6" box="[333,463,1841,1866]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="9" pageNumber="596">Figs 3032</figureCitation>
). Basal segment of chelicera with distinct dorsal bulla, no prominent armaments except a few small setiferous tubercles. Cheliceral claw unarmed, with sparse setae only. Fingers relatively short, inner edges toothed as illustrated (
<figureCitation id="1312FFA0FF95A4189B84F8D8FDC3F80E" box="[470,555,1913,1938]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="9" pageNumber="596">Fig. 32</figureCitation>
): moveable finger with five teeth, fixed finger with four teeth.
</paragraph>
<paragraph id="8B96E325FF95A41B9A95F83CFD6FFF6B" blockId="9.[151,1437,1841,2010]" lastBlockId="10.[151,1437,151,1005]" lastPageId="10" lastPageNumber="597" pageId="9" pageNumber="596">
Pedipalps (
<figureCitation id="1312FFA0FF95A4189B14F83CFE74F82A" box="[326,412,1949,1974]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="9" pageNumber="596">Fig. 33</figureCitation>
). Coxa with one dorsal setiferous tubercle. Trochanter smooth. Femur straight, unarmed, with only a few ventral setae. Patella slim and markedly elongate, the length almost equal to the sum of the length of the tibia and tarsus, also with a meso-ventral setiferous tubercle. Tibia with two ventromesal setiferous tubercles and three ventroectal ones. Tarsus with two setiferous tubercles on each side of ventral surface. Tarsal claw curved, approximately the same length as the tarsus.
</paragraph>
<paragraph id="8B96E325FF96A41B9A95FEA4FE06FEF9" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="597">
Legs (
<figureCitation id="1312FFA0FF96A41B9B5FFEA5FE89FE80" box="[269,353,260,285]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="10" pageNumber="597">Fig. 34</figureCitation>
). All segments smooth, with only scarce short setae. Femur IV slightly curved. Distitarsus I with two tarsomeres, distitarsus II with three tarsomeres. Distitarsi IIIIV with scopulae (
<figureCitation id="1312FFA0FF96A41B9E1AFE86FB75FEDC" box="[1096,1181,295,320]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="10" pageNumber="597">Fig. 34</figureCitation>
). Tarsal claws smooth. Tarsal formula (IIV): 3/5/5/5.
</paragraph>
<paragraph id="8B96E325FF96A41B9A95FECEFAD3FD34" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="597">
Penis (
<figureCitation id="1312FFA0FF96A41B9B46FECEFE72FE14" box="[276,410,367,392]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Figs 3843</figureCitation>
). The basal third of the shaft slender and slightly broadened, the distal half more strongly enlarged but nearly parallel-sided until apical portion (
<emphasis id="B95D3F37FF96A41B997EFE34FC55FE30" box="[812,957,404,428]" italics="true" pageId="10" pageNumber="597">pars distalis</emphasis>
;
<figureCitation id="1312FFA0FF96A41B9980FE35FB8AFE30" box="[978,1122,404,429]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Figs 38, 40</figureCitation>
). The ventral plate nearly rectangular, with slightly indented apical margin. Two spoon-shaped titillators enclose nearly two thirds of
<emphasis id="B95D3F37FF96A41B9F17FE19FF00FE68" italics="true" pageId="10" pageNumber="597">capsula interna</emphasis>
in penis resting state (
<figureCitation id="1312FFA0FF96A41B9BB8FE7DFDD6FE68" box="[490,574,476,501]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Fig. 38</figureCitation>
). In the everted state, titillators bent perpendicular to truncus shaft, forming two characteristic spoon-shaped structures (“spathes”, see above). The tips of the two conductors closely associated to each other, in the resting state surrounded by both titillators, just like the nib of fountain pen (
<figureCitation id="1312FFA0FF96A41B9F6EFD85FA67FDA0" box="[1340,1423,548,573]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Fig. 38</figureCitation>
). On top of the inner side of both conductors a triangular, conical proximal pointing tip (
<figureCitation id="1312FFA0FF96A41B9E31FDE6FB0DFDFC" box="[1123,1253,583,608]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Figs 41, 43</figureCitation>
). Stylus slender, hidden between the conductors.
<emphasis id="B95D3F37FF96A41B9855FDCCFD7DFD18" box="[519,661,620,644]" italics="true" pageId="10" pageNumber="597">Pars distalis</emphasis>
of truncus with 12 ventral setae: four setae on the ventral plate, six setae near the base of the ventral plate, two additional setae placed slightly more proximally (
<figureCitation id="1312FFA0FF96A41B9EFEFD2EFAC7FD34" box="[1196,1327,655,680]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Figs 40, 43</figureCitation>
).
</paragraph>
<paragraph id="8B96E325FF96A41B9A95FD15FE3AFCA4" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="597">
<emphasis id="B95D3F37FF96A41B9A95FD15FE74FD51" bold="true" box="[199,412,692,717]" pageId="10" pageNumber="597">
Female
<typeStatus id="54925D87FF96A41B9B75FD15FE7FFD51" box="[295,407,692,717]" pageId="10" pageNumber="597" type="paratype">paratype</typeStatus>
:
</emphasis>
(
<figureCitation id="1312FFA0FF96A41B9BFFFD15FE10FD50" box="[429,504,692,717]" captionStart="FIGURES 69" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[233,1361,294,1812]" captionTargetId="figure@3.[206,1393,196,1882]" captionTargetPageId="3" captionText="FIGURES 69. Body of holotype and paratype, dorsal view. 67. Biantes calyptroideus sp. nov. 6. Male. 7. Female. 89. Biantes spatulatus sp. nov. 8. Male. 9. Female. Scale bars: 1mm (67); 0.5 mm (89)." httpUri="https://zenodo.org/record/1460328/files/figure.png" pageId="10" pageNumber="597">Figs 9</figureCitation>
,
<figureCitation id="1312FFA0FF96A41B9854FD15FDA6FD51" box="[518,590,692,717]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="10" pageNumber="597">3637</figureCitation>
,
<figureCitation id="1312FFA0FF96A41B980EFD15FD4DFD51" box="[604,677,692,717]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">4446</figureCitation>
,
<figureCitation id="1312FFA0FF96A41B98E1FD15FD3BFD51" box="[691,723,692,717]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="10" pageNumber="597">52</figureCitation>
). In general, appearance similar to the male (
<figureCitation id="1312FFA0FF96A41B9E8EFD15FACFFD50" box="[1244,1319,692,717]" captionStart="FIGURES 69" captionStartId="3.[151,264,1914,1937]" captionTargetBox="[233,1361,294,1812]" captionTargetId="figure@3.[206,1393,196,1882]" captionTargetPageId="3" captionText="FIGURES 69. Body of holotype and paratype, dorsal view. 67. Biantes calyptroideus sp. nov. 6. Male. 7. Female. 89. Biantes spatulatus sp. nov. 8. Male. 9. Female. Scale bars: 1mm (67); 0.5 mm (89)." httpUri="https://zenodo.org/record/1460328/files/figure.png" pageId="10" pageNumber="597">Figs 9</figureCitation>
,
<figureCitation id="1312FFA0FF96A41B9F67FD15FABCFD51" box="[1333,1364,692,717]" captionStart="FIGURES 4752" captionStartId="9.[151,264,1651,1674]" captionTargetBox="[151,1434,194,1631]" captionTargetId="figure@9.[151,1436,193,1631]" captionTargetPageId="9" captionText="FIGURES 4752. Photographs of male holotype and female paratype of Biantes calyptroideus sp. nov. and Biantes spatulatus sp. nov.. 4749. Biantes calyptroideus sp. nov.. 47. Male body and parts of appendages, dorsal view. 48. Female body and parts of appendages, dorsal view. 49. Male body and parts of appendages, lateral view. 5052. Biantes spatulatus sp. nov.. 50. Male body and parts of appendages, lateral view. 51. Male body and parts of appendages, dorsal view. 52. Female body and parts of appendages, dorsal view. Scale bars: 1 mm (49); 0.5 mm (4748; 5052)." httpUri="https://zenodo.org/record/1460338/files/figure.png" pageId="10" pageNumber="597">52</figureCitation>
), with only a slight difference in inner edges of finger of chelicera (
<figureCitation id="1312FFA0FF96A41B996BFD76FC62FD6C" box="[825,906,727,752]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="10" pageNumber="597">Fig. 37</figureCitation>
) and in genital operculum (
<figureCitation id="1312FFA0FF96A41B9EECFD76FAF9FD6C" box="[1214,1297,727,752]" captionStart="FIGURES 2837" captionStartId="7.[151,264,1873,1896]" captionTargetBox="[237,1435,303,1819]" captionTargetId="figure@7.[163,1448,193,1884]" captionTargetPageId="7" captionText="FIGURES 2837. Biantes spatulatus sp. nov. male holotype and female paratype. 28. Male body, lateral view. 29. Right coxae IIV of male, lateral view. 30. Left chelicera of male, medial view. 31. Same, ectal view. 32. Cheliceral fingers of male, frontal view. 33. Left pedipalp of male, medial view. 34. Right tarsus IV of male, retrolateral view. 35. Genital operculum of male, ventral view. 36. Genital operculum of female, ventral view. 37. Cheliceral fingers of female, frontal view. Scale bars: 1 mm (28); 0.5 mm (29, 33); 0.25 mm (3032, 34); 0.125 mm (3537)." httpUri="https://zenodo.org/record/1460334/files/figure.png" pageId="10" pageNumber="597">Fig. 36</figureCitation>
). Ovipositor (
<figureCitation id="1312FFA0FF96A41B9ACDFD5DFEF7FC89" box="[159,287,764,789]" captionStart="FIGURES 3846" captionStartId="8.[151,264,1917,1940]" captionTargetBox="[270,1353,252,1894]" captionTargetId="figure@8.[220,1373,193,1905]" captionTargetPageId="8" captionText="FIGURES 3846. Biantes spatulatus sp. nov. genitalia of male holotype and female paratype. 38. Penis, dorsal view. 39. Same, lateral view. 40. Same, ventral view. 41. Pars distalis of penis (everted), dorsal view. 42. Same, lateral view. 43. Same, ventral view. 44. Ovipositor, dorsal view. 45. Same, ventral view. 46. Same, lateral view. Scale bars: 0.25 mm (3840); 0.125 mm (4146)." httpUri="https://zenodo.org/record/1460336/files/figure.png" pageId="10" pageNumber="597">Figs 4446</figureCitation>
) conical. Distal tip divided into two external lobes. Each lobe with five setae: two on the ventral side and three on the dorsal side.
</paragraph>
<paragraph id="8B96E325FF96A41B9A95FCE5FBDEFC38" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="597">
<emphasis id="B95D3F37FF96A41B9A95FCE5FE9EFCC1" bold="true" box="[199,374,836,861]" pageId="10" pageNumber="597">Measurements</emphasis>
. Male (female
<typeStatus id="54925D87FF96A41B9879FCE5FD79FCC0" box="[555,657,836,860]" pageId="10" pageNumber="597" type="paratype">paratype</typeStatus>
): body 1.47(1.58) long; scutum 1.32 (1.26) long, 1.15 (1.10) wide; carapace 0.51 (0.46) long, 0.81 (0.73) wide. Interocular distance 0.46 (0.43) (not including the eye). Pedipalp claw 0.32(0.31) long. Penis 0.65 long. Measurements of pedipalp and legs as in
<tableCitation id="C6ABD69EFF96A41B998EFC2DFBDAFC38" box="[988,1074,908,933]" captionStart="TABLE 2" captionStartId="10.[151,244,1052,1077]" captionText="TABLE 2. Pedipalp and leg measurements of Biantes spatulatus sp. nov., holotype/female" pageId="10" pageNumber="597">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8B96E325FF96A41B9A95FC0EFA6FFC54" blockId="10.[151,1437,151,1005]" box="[199,1415,943,968]" pageId="10" pageNumber="597">
<emphasis id="B95D3F37FF96A41B9A95FC0EFECBFC54" bold="true" box="[199,291,943,968]" pageId="10" pageNumber="597">Habitat</emphasis>
. The specimens were collected by leaf litter sifting in dark, moist forest habitat under dense canopy.
</paragraph>
</subSubSection>
<subSubSection id="C333B0AEFF96A41B9A95FC75FD19FC71" box="[199,753,980,1005]" pageId="10" pageNumber="597" type="distribution">
<paragraph id="8B96E325FF96A41B9A95FC75FD19FC71" blockId="10.[151,1437,151,1005]" box="[199,753,980,1005]" pageId="10" pageNumber="597">
<emphasis id="B95D3F37FF96A41B9A95FC75FEB1FC71" bold="true" box="[199,345,980,1005]" pageId="10" pageNumber="597">Distribution</emphasis>
. Known only from the
<typeStatus id="54925D87FF96A41B9832FC74FD78FC71" box="[608,656,981,1005]" pageId="10" pageNumber="597">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>