127 lines
11 KiB
XML
127 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.72.744" ID-GBIF-Dataset="9b029a3f-f31b-421f-a62e-fb4dd8ff6731" ID-PMC="PMC3083004" ID-Pensoft-Pub="1313-2970-72-23" ID-PubMed="21594103" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-72-23" ModsDocOrigin="ZooKeys 72" ModsDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451250861919" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak" docDate="2010" docId="E959CB544DC34197A1EBE22C3A763F76" docLanguage="en" docName="ZooKeys 72: 23-68" docOrigin="ZooKeys 72" docSource="http://dx.doi.org/10.3897/zookeys.72.744" docTitle="Tylopus bispinosus Likhitrakarn, Golovatch, Prateepasen & Panha, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="27" masterDocId="D7146C3215665021FFDFFFACFF8CFFD0" masterDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="68" masterPageNumber="23" pageNumber="26" updateTime="1668165214761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prateepasen, Rujiporn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>72</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.72.744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.72.744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-72-23</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360926" LSID="urn:lsid:zoobank.org:act:D501889C-39AB-427B-A47D-0903412C651C" httpUri="http://treatment.plazi.org/id/E959CB544DC34197A1EBE22C3A763F76" lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">
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<subSubSection pageId="3" pageNumber="26" type="nomenclature">
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<paragraph pageId="3" pageNumber="26">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:D501889C-39AB-427B-A47D-0903412C651C" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="26">sp. n.</taxonomicNameLabel>
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Figs 13
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="26" type="holotype">
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<paragraph pageId="3" pageNumber="26">Holotype</paragraph>
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<paragraph pageId="3" pageNumber="26">♂ (CUMZ), Thailand, Tak Province, Umphang District, near Umphang City, ca 490 m, 16°2'20N, 98°52E, 6.07.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="26" type="paratypes">
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<paragraph pageId="3" pageNumber="26">Paratypes:</paragraph>
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<paragraph pageId="3" pageNumber="26">1 ♂, 1 ♀, 2 juv. (CUMZ), same locality, together with holotype. 6 ♂, 4 ♀ (CUMZ), 3 ♂ (ZMUC), 3 ♂ (ZMUM), Tak Province, Umphang District, Doi Hua Mod, 900 m, 16°3'14N, 98°49'16E, 5.06.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn. 6 ♂, 1 ♀ (CUMZ), same Province, same District, Cave Ta Ko Bi, ca 530 m, 16°03'14N, 98°49'14E, 5.07.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="26" type="name">
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<paragraph pageId="3" pageNumber="26">Name:</paragraph>
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<paragraph pageId="3" pageNumber="26">To emphasize the spiniform processes h and z of the gonopod.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="26" type="diagnosis">
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<paragraph pageId="3" pageNumber="26">Diagnosis:</paragraph>
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<paragraph pageId="3" pageNumber="26">Differs from congeners in both processes h and z of the gonopod being spiniform.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26" type="description">
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<paragraph pageId="3" pageNumber="26">Description:</paragraph>
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<paragraph lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">
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Length 26 mm (holotype), 25-29 mm (♂), 33-38 mm (♀), width of midbody pro- and metazona 2.0 and 2.9 mm (holotype), 1.8-2.4 and 3.1-3.2 mm (♂), 2.4-2.7 and 3.3-3.8 mm (♀), respectively. Coloration of live animals black
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<pageBreakToken pageId="4" pageNumber="27" start="start">-</pageBreakToken>
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brown (Fig. 1A): calluses of paraterga, venter and legs only slightly lighter, dark brown, but turning light brown in alcohol (Fig. 1
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<normalizedToken originalValue="A–K">A-K</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="4" pageNumber="27">Clypeolabral region of head very densely, vertigial region sparsely setose. Epicranial suture distinct. Antennae long and slender, reaching behind segment 4 (♂) or 3 (♀) dorsally. In width, head <collum <segments 3-4 <2 <5-16 (♂), or head = segment 3 <4 <collum <segments 5-16 (♀); thereafter body gradually and gently tapering towards telson (Fig. 1B).</paragraph>
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<paragraph pageId="4" pageNumber="27">
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Tegument generally rather smooth and shining, but prozona very finely rugulose, metaterga often rugose (Fig. 1
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<normalizedToken originalValue="B–G">B-G</normalizedToken>
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); surface below paraterga finely microgranular (Fig. 1E, F). Collum with three transverse rows of setae: 5+5 in anterior, 2+2 in middle, and 4+4 in posterior row; paraterga evident, rounded, flap-shaped (Fig. 1B, C). Metaterga with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points. Axial line at most barely visible only on metaterga. Paraterga strongly developed (Fig. 1
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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), lying high (at 1/3-1/4 midbody height), only slightly inclined laterally, pointed caudally and acutangular already from segment 2, especially strongly so on caudal segments; calluses very thin on poreless segments, slightly thicker on pore-bearing ones; anterior 1/3 of poreless calluses with two evident (anterior larger, posterior smaller), lateral, setigerous incisions, but with only a single strong one (anterior) on pore-bearing calluses (Fig. 1
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<normalizedToken originalValue="B–G">B-G</normalizedToken>
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); paraterga more strongly developed in ♂. Ozopores entirely lateral, lying in an ovoid groove about 1/3 in front of caudal corner, the latter always surpassing rear tergal contour (Fig. 1
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<normalizedToken originalValue="B–H">B-H</normalizedToken>
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). Transverse sulcus evident on metaterga 4-18, reaching base of paraterga, evident and rather deep, faintly rugulose at bottom. Stricture between pro- and metazona very clearly ribbed (Fig. 1D). Epiproct tip faintly concave to subtruncate, pre-apical papillae evident (Fig. 1G). Hypoproct roundly subtrapeziform, caudal setae strongly separated (Fig. 1H). Pleurosternal carinae well-developed on segments 2-17 (♂) or 2-7 (♀), mostly as low bulges anteriorly and a distinct spine posteriorly (Fig. 1C, E, F).
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</paragraph>
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<paragraph pageId="4" pageNumber="27">Sterna moderately setose, without modifications; a deeply notched sternal lobe between ♂coxae 4 (Fig. 1I, J). Legs long and slender (Fig. 1B, C, H), especially so in ♂ compared to ♀ (1.3-1.5 versus 0.9-1.1 times as long as midbody height); prefemora distinctly bulged laterally (Fig. 1K), acropodites with particularly dense, nearly adpressed setae, including tarsal brushes.</paragraph>
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<paragraph pageId="4" pageNumber="27">Gonopods (Figs 2, 3) with lobe l well-demarcated; spine h very small; spine z considerably more prominent.</paragraph>
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<caption pageId="4" pageNumber="27">
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<paragraph pageId="4" pageNumber="27">
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Figure 1.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., ♂ paratype from near Umphang City (A) and ♂ paratype from Doi Hua Mod (
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<normalizedToken originalValue="B–K">B-K</normalizedToken>
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). A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively. D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively K midbody leg.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="27">
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<paragraph pageId="4" pageNumber="27">
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Figure 2.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., ♂ paratype from Doi Hua Mod. A, B right gonopod, mesal and lateral views, respectively
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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distal part of right gonopod, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="27">
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<paragraph pageId="4" pageNumber="27">
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Figure 3.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., ♂ paratype from Doi Hua Mod. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |