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<document id="F6A6FA5348F5F4030745C85999F83320" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan &amp; Kompier, Tom" docDate="2016" docId="8A1EB74F8979FF90C7F36403FDD5931B" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta satoi Asahina 1997" docType="treatment" docVersion="7" lastPageNumber="536" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="535" updateTime="1698233463814" updateUser="plazi">
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<mods:title id="4476A7EDB41113DED0DAF786DF2A2728">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="95EC9753D6D269A69370D3ED177F8CB8">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="4FECEE6AC45A2B7AC31F864F76F8EF41">Kompier, Tom</mods:namePart>
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<mods:date id="E56D37B39BBADB9C0076993E7B487006">2016</mods:date>
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<treatment id="8A1EB74F8979FF90C7F36403FDD5931B" ID-DOI="http://doi.org/10.5281/zenodo.6056381" ID-GBIF-Taxon="127620795" ID-Zenodo-Dep="6056381" LSID="urn:lsid:plazi:treatment:8A1EB74F8979FF90C7F36403FDD5931B" httpUri="http://treatment.plazi.org/id/8A1EB74F8979FF90C7F36403FDD5931B" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535">
<subSubSection id="4AAD55D28979FF91C7F36403FD5597F5" box="[151,716,1652,1678]" pageId="6" pageNumber="535" type="nomenclature">
<paragraph id="020806598979FF91C7F36403FD5597F5" blockId="6.[151,716,1652,1712]" box="[151,716,1652,1678]" pageId="6" pageNumber="535">
<heading id="5940B1358979FF91C7F36403FD5597F5" bold="true" box="[151,716,1652,1678]" fontSize="11" level="1" pageId="6" pageNumber="535" reason="1">
<emphasis id="30C3DA4B8979FF91C7F36403FD5597F5" bold="true" box="[151,716,1652,1678]" pageId="6" pageNumber="535">
6.
<taxonomicName id="C5B77DDA8979FF91C7D06403FDB497F5" ID-CoL="4N794" authority="Asahina, 1997" authorityName="Asahina" authorityYear="1997" box="[180,557,1652,1678]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="30C3DA4B8979FF91C7D06403FEEF97F5" bold="true" box="[180,374,1652,1678]" italics="true" pageId="6" pageNumber="535">Protosticta satoi</emphasis>
Asahina, 1997
</taxonomicName>
—dark form
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28979FF91C7F364E0FBDE9650" pageId="6" pageNumber="535" type="description">
<paragraph id="020806598979FF91C7F364E0FEA797CB" blockId="6.[151,716,1652,1712]" box="[151,318,1687,1712]" pageId="6" pageNumber="535">
(
<figureCitation id="9A8C1ADC8979FF91C7FA64E0FF6597CB" box="[158,252,1687,1712]" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="6" pageNumber="535">Figure 5</figureCitation>
AK)
</paragraph>
<paragraph id="020806598979FF91C7F364AEFC219676" blockId="6.[151,1436,1752,1897]" pageId="6" pageNumber="535">
<taxonomicName id="C5B77DDA8979FF91C7F364AEFD029795" authority="Asahina (1997)" authorityName="Asahina" authorityYear="1997" box="[151,667,1752,1774]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="535" phylum="Arthropoda" rank="subSpecies" species="khaosoidaoensis" subSpecies="satoi">
<emphasis id="30C3DA4B8979FF91C7F364AEFE749795" box="[151,493,1752,1774]" italics="true" pageId="6" pageNumber="535">Protosticta khaosoidaoensis satoi</emphasis>
: Asahina (1997)
</taxonomicName>
, pp. 108109,
<figureCitation id="9A8C1ADC8979FF91C45664AFFC0D9795" box="[818,916,1752,1774]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1912,1934]" captionStartId-1="6.[151,250,1471,1493]" captionTargetBox-0="[169,1425,470,1883]" captionTargetBox-1="[156,1434,450,1445]" captionTargetId-0="figure@4.[150,1436,465,1891]" captionTargetId-1="figure@6.[151,1436,447,1449]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 3. Protosticta grandis, Cao Bang. [A E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." captionText-1="FIGURE 4. Protosticta spp. [A H] Protosticta ngoai spec. nov., holotype male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal view, (C) lateral view &amp; (D) oblique-ventral view; (E) paraprocts in ventral view; penile organ in (F) dorsal view &amp; (G) lateral view; (H) wings; [I J] Protosticta taipokauensis male—anal appendages in (I), lateral view &amp; (J), oblique-ventral view (rearranged from Asahina &amp; Dudgeon, 1987, p. 3, Figs. 6,7)." httpUri-0="https://zenodo.org/record/257726/files/figure.png" httpUri-1="https://zenodo.org/record/257727/files/figure.png" pageId="6" pageNumber="535">Figs. 34</figureCitation>
.
<typeStatus id="DD0CB8FB8979FF91C4FB64AEFC489795" box="[927,977,1753,1774]" pageId="6" pageNumber="535">Type</typeStatus>
locality: Tam Dao National Park, Vinh Phuc Province, northern
<collectingCountry id="7AA046C98979FF91C6EE6480FE789676" box="[394,481,1783,1805]" name="Vietnam" pageId="6" pageNumber="535">Vietnam</collectingCountry>
.—
<bibRefCitation id="66267BA88979FF91C69A6480FD1E9676" author="Tsuda" box="[510,647,1783,1805]" pageId="6" pageNumber="535" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately Published, Osaka, 430 pp." type="book" year="2000">Tsuda (2000)</bibRefCitation>
, p. 326 (catalogued
<collectingCountry id="7AA046C98979FF91C4376480FC379676" box="[851,942,1783,1805]" name="Vietnam" pageId="6" pageNumber="535">Vietnam</collectingCountry>
).
</paragraph>
<paragraph id="020806598979FF91C7F36560FBDE9650" blockId="6.[151,1436,1752,1897]" box="[151,1095,1813,1836]" pageId="6" pageNumber="535">
<taxonomicName id="C5B77DDA8979FF91C7F36560FE2D9650" box="[151,436,1813,1836]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8979FF91C7F36560FE2D9650" box="[151,436,1813,1836]" italics="true" pageId="6" pageNumber="535">Protosticta khaosoidaoensis</emphasis>
</taxonomicName>
nec Asahina, 1997:
<bibRefCitation id="66267BA88979FF91C5E76561FCDC9650" author="Do" box="[643,837,1813,1835]" pageId="6" pageNumber="535" refString="Do, M. C. &amp; Dang, T. T. H. (2007) Checklist of dragonfly from Vietnam. Vietnam National University Publisher, Hanoi, 182 pp." type="book" year="2007">Do &amp; Dang (2007)</bibRefCitation>
, p. 65 (distribution map).
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28979FF90C7F36542FDB89363" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535" type="reference_group">
<paragraph id="020806598979FF91C7F36542FC929612" blockId="6.[151,1436,1752,1897]" pageId="6" pageNumber="535">
<taxonomicName id="C5B77DDA8979FF91C7F36542FEA49631" ID-CoL="4N794" box="[151,317,1845,1866]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="30C3DA4B8979FF91C7F36542FEA49631" box="[151,317,1845,1866]" italics="true" pageId="6" pageNumber="535">Protosticta satoi</emphasis>
</taxonomicName>
: van
<bibRefCitation id="66267BA88979FF91C6106543FE469631" author="Tol" box="[372,479,1844,1866]" pageId="6" pageNumber="535" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
(new status), pp. 222227, Figs. 1116.—
<bibRefCitation id="66267BA88979FF91C4ED6543FC6A9631" author="Do" box="[905,1011,1844,1866]" pageId="6" pageNumber="535" refString="Do, M. C. (2011) Serial photographs of emergence of Protosticta satoi (Asahina, 1997) (dark form) in Tam Dao National Park, North Vietnam (Zygoptera: Platystictidae). Agrion, Newsletter of the worldwide dragonfly Association, 15 (1), 5 - 7." type="journal article" year="2011">Do (2011)</bibRefCitation>
(serial photographs of emergence), pp. 5 7,
<figureCitation id="9A8C1ADC8979FF91C7846524FEA29612" box="[224,315,1875,1897]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="1.[151,250,1915,1937]" captionStartId-1="3.[151,250,1687,1709]" captionStartId-2="4.[151,250,1912,1934]" captionStartId-3="6.[151,250,1471,1493]" captionStartId-4="7.[151,250,1584,1606]" captionStartId-5="8.[151,250,1858,1880]" captionStartId-6="9.[151,250,1699,1721]" captionStartId-7="11.[151,250,1179,1201]" captionStartId-8="12.[151,250,1246,1268]" captionTargetBox-0="[157,1432,706,1896]" captionTargetBox-1="[159,1432,476,1674]" captionTargetBox-2="[169,1425,470,1883]" captionTargetBox-3="[156,1434,450,1445]" captionTargetBox-4="[150,1433,668,1559]" captionTargetBox-5="[155,1430,564,1821]" captionTargetBox-6="[151,1432,481,1664]" captionTargetBox-7="[157,1428,303,1152]" captionTargetBox-8="[155,1429,273,1221]" captionTargetId-0="figure@1.[151,1435,701,1909]" captionTargetId-1="figure@3.[151,1437,469,1675]" captionTargetId-2="figure@4.[150,1436,465,1891]" captionTargetId-3="figure@6.[151,1436,447,1449]" captionTargetId-4="figure@7.[150,1436,663,1562]" captionTargetId-5="figure@8.[151,1435,555,1831]" captionTargetId-6="figure@9.[151,1436,479,1677]" captionTargetId-7="figure@11.[151,1436,303,1156]" captionTargetId-8="figure@12.[151,1436,265,1225]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="11" captionTargetPageId-8="12" captionText-0="FIGURE 1. Protosticta beaumonti, Mau Son. [A D] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus &amp; paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." captionText-1="FIGURE 2. Protosticta caroli, Ba Na. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H J] female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." captionText-2="FIGURE 3. Protosticta grandis, Cao Bang. [A E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." captionText-3="FIGURE 4. Protosticta spp. [A H] Protosticta ngoai spec. nov., holotype male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal view, (C) lateral view &amp; (D) oblique-ventral view; (E) paraprocts in ventral view; penile organ in (F) dorsal view &amp; (G) lateral view; (H) wings; [I J] Protosticta taipokauensis male—anal appendages in (I), lateral view &amp; (J), oblique-ventral view (rearranged from Asahina &amp; Dudgeon, 1987, p. 3, Figs. 6,7)." captionText-4="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." captionText-5="FIGURE 6. Protosticta satoi (light form), Xuan Son. [A B] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; [C E] female— (C) head &amp; thorax in lateral view; (D) prothorax in dorsal view; (E) abdomen tip in lateral view." captionText-6="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." captionText-7="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." captionText-8="FIGURE 9. Protosticta spp. [A M] Protosticta pseudocuriosa spec. nov. [A I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." httpUri-0="https://zenodo.org/record/257724/files/figure.png" httpUri-1="https://zenodo.org/record/257725/files/figure.png" httpUri-2="https://zenodo.org/record/257726/files/figure.png" httpUri-3="https://zenodo.org/record/257727/files/figure.png" httpUri-4="https://zenodo.org/record/257728/files/figure.png" httpUri-5="https://zenodo.org/record/257729/files/figure.png" httpUri-6="https://zenodo.org/record/257730/files/figure.png" httpUri-7="https://zenodo.org/record/257731/files/figure.png" httpUri-8="https://zenodo.org/record/257732/files/figure.png" pageId="6" pageNumber="535">Figs. 19</figureCitation>
.—von
<bibRefCitation id="66267BA88979FF91C6EE6524FDE69612" box="[394,639,1875,1897]" pageId="6" pageNumber="535" refString="von Ellenrieder, N., Hauser, M., Gaimari, S. &amp; Pham, T. (2015) First records of Macromia katae (Macromiidae) and Indothemis carnatica (Libellulidae) from Vietnam (Insecta: Odonata). Check List, 1514, 11 (1), 1 - 13." type="journal article">
Ellenrieder
<emphasis id="30C3DA4B8979FF91C5656523FDAD9612" box="[513,564,1875,1897]" italics="true" pageId="6" pageNumber="535">et al.</emphasis>
(2015)
</bibRefCitation>
, p. 8, Fig. 12.
</paragraph>
<paragraph id="020806598979FF91C7F365F8FBFF968B" blockId="6.[151,1436,1935,2032]" pageId="6" pageNumber="535">
<emphasis id="30C3DA4B8979FF91C7F365F8FE0B96D3" bold="true" box="[151,402,1935,1960]" pageId="6" pageNumber="535">Specimens examined</emphasis>
.
<specimenCount id="14B1CDD08979FF91C6C765F8FE6696D3" box="[419,511,1935,1960]" pageId="6" pageNumber="535" type="male">5 males</specimenCount>
,
<specimenCount id="14B1CDD08979FF91C56965F8FDED96D3" box="[525,628,1935,1960]" pageId="6" pageNumber="535" type="female">1 female</specimenCount>
, Pia Oac Nature Reserve, Cao Bang Province,
<date id="760920998979FF91C3FE65F8FA9696D3" box="[1178,1295,1935,1960]" pageId="6" pageNumber="535" value="2015-05-18">18.V.2015</date>
, Quoc Toan Phan leg.;
<specimenCount id="14B1CDD08979FF91C66B65C4FEC796B7" box="[271,350,1971,1996]" pageId="6" pageNumber="535" type="male">1 male</specimenCount>
, Tam Dao National Park, Vinh Phuc Province,
<date id="760920998979FF91C41665C4FB9196B0" box="[882,1032,1971,1995]" pageId="6" pageNumber="535" value="2013-08-18">18.VIII.2013</date>
;
<specimenCount id="14B1CDD08979FF91C37065C4FBF496B7" box="[1044,1133,1971,1996]" pageId="6" pageNumber="535" type="male">4 males</specimenCount>
, Ba Be National Park, Bac Kan Province,
<date id="760920998979FF91C65A65A0FE5E968B" box="[318,455,2007,2032]" pageId="6" pageNumber="535" value="2013-07-26">26.VII.2013</date>
and
<specimenCount id="14B1CDD08979FF91C69965A0FDCF968B" box="[509,598,2007,2032]" pageId="6" pageNumber="535" type="male">4 males</specimenCount>
, same location,
<date id="760920998979FF91C46E65A0FC159694" box="[778,908,2007,2031]" pageId="6" pageNumber="535" value="2014-06-22">22.VI.2014</date>
, Tom Kompier leg.
</paragraph>
<paragraph id="020806598978FF90C7A362E0FEC0901E" blockId="7.[151,1437,151,608]" pageId="7" pageNumber="536">
<emphasis id="30C3DA4B8978FF90C7A362E0FE9391CB" bold="true" box="[199,266,151,176]" pageId="7" pageNumber="536">Notes</emphasis>
. Asahina (1997) described the subspecies
<taxonomicName id="C5B77DDA8978FF90C58262EEFC6191CB" ID-CoL="4N794" box="[742,1016,151,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="536" phylum="Arthropoda" rank="subSpecies" species="khaosoidaoensis" subSpecies="satoi">
<emphasis id="30C3DA4B8978FF90C58262EEFC6191CB" box="[742,1016,151,176]" italics="true" pageId="7" pageNumber="536">P. khaosoidaoensis satoi</emphasis>
</taxonomicName>
(=
<taxonomicName id="C5B77DDA8978FF90C37B62EEFBE891CB" ID-CoL="4N794" box="[1055,1137,153,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="30C3DA4B8978FF90C37B62EEFBE891CB" box="[1055,1137,153,176]" italics="true" pageId="7" pageNumber="536">P. satoi</emphasis>
</taxonomicName>
light form) on the basis of a single female from Tam Dao National Park (Vinh Phuc Province, northern
<collectingCountry id="7AA046C98978FF90C30562CBFB5C91AE" box="[1121,1221,188,213]" name="Vietnam" pageId="7" pageNumber="536">Vietnam</collectingCountry>
). Van
<bibRefCitation id="66267BA88978FF90C27862CBFA0591AF" author="Tol" box="[1308,1436,188,213]" pageId="7" pageNumber="536" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
subsequently elevated
<emphasis id="30C3DA4B8978FF90C6F16296FE7D9183" box="[405,484,225,248]" italics="true" pageId="7" pageNumber="536">P. s a t o i</emphasis>
as a distinct species from
<taxonomicName id="C5B77DDA8978FF90C4756296FC7F9183" ID-CoL="4N78H" box="[785,998,223,248]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8978FF90C4756296FC7F9183" box="[785,998,223,248]" italics="true" pageId="7" pageNumber="536">P. khaosoidaoensis</emphasis>
</taxonomicName>
based on specimens from Cuc Phuong National Park (Ninh Binh Province) and Tam Dao. As in
<taxonomicName id="C5B77DDA8978FF90C4446372FC2B9066" ID-CoL="4N77Y" box="[800,946,260,285]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="beaumonti">
<emphasis id="30C3DA4B8978FF90C4446372FC2B9066" box="[800,946,260,285]" italics="true" pageId="7" pageNumber="536">P. beaumonti</emphasis>
</taxonomicName>
, there are no obvious structural differences between the dark and light froms of
<emphasis id="30C3DA4B8978FF90C548635EFDE3903B" box="[556,634,297,320]" italics="true" pageId="7" pageNumber="536">P. s at oi</emphasis>
, but the forms are easily separated by their thoracic color pattern (
<figureCitation id="9A8C1ADC8978FF90C2076350FF3C901E" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="7" pageNumber="536">Figs. 5</figureCitation>
A, 5H, 6A, 6C).
</paragraph>
<paragraph id="020806598978FF90C7A36307FDB89363" blockId="7.[151,1437,151,608]" pageId="7" pageNumber="536">
Van
<bibRefCitation id="66267BA88978FF90C7996318FEEE90F3" author="Tol" box="[253,375,367,392]" pageId="7" pageNumber="536" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
described the color pattern of the prothorax of the male
<emphasis id="30C3DA4B8978FF90C3706306FBFC90F3" box="[1044,1125,369,392]" italics="true" pageId="7" pageNumber="536">P. s a t o i</emphasis>
dark form from Tam Dao National Park as “anterior lobe black, median lobe with paired protuberance brownish black, medially darker”. This character is consistent with all dark form males of
<emphasis id="30C3DA4B8978FF90C47563CEFCFB90AB" box="[785,866,441,464]" italics="true" pageId="7" pageNumber="536">
<taxonomicName id="C5B77DDA8978FF90C47563CEFCD290AB" box="[785,843,441,464]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="sat">P. sat</taxonomicName>
oi
</emphasis>
from several locations in northern
<collectingCountry id="7AA046C98978FF90C39163C0FACD90AB" box="[1269,1364,439,464]" name="Vietnam" pageId="7" pageNumber="536">Vietnam</collectingCountry>
(
<figureCitation id="9A8C1ADC8978FF90C20063C0FF3C908E" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="7" pageNumber="536">Figs. 5</figureCitation>
B,
<date id="760920998978FF90C7A163ABFF46908E" box="[197,223,476,501]" pageId="7" pageNumber="536">5I</date>
), apart from one male from the same location (Tam Dao), which has the median lobe yellowish, anterior and posterior lobes darkened (
<figureCitation id="9A8C1ADC8978FF90C6DD6388FE659363" box="[441,508,511,536]" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="7" pageNumber="536">Fig. 5</figureCitation>
K).
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28978FF90C7A36053FDD5931B" pageId="7" pageNumber="536" type="description">
<paragraph id="020806598978FF90C7A36053FDD5931B" blockId="7.[151,1437,151,608]" pageId="7" pageNumber="536">
<emphasis id="30C3DA4B8978FF90C7A36053FEEF9346" bold="true" box="[199,374,548,573]" pageId="7" pageNumber="536">Measurements</emphasis>
. Male. Hindwing 1922.5; abdomen including appendages 3442. Female. Hindwing 22, abdomen including appendages 3334.
</paragraph>
</subSubSection>
</treatment>
</document>