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<document id="579AB3D2F2FE09C495E938C1181A5DB7" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan &amp; Kompier, Tom" docDate="2016" docId="8A1EB74F8976FF9DC7F362EFFC8B92FB" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta socculus Phan &amp; Kompier, 2016, spec. nov." docType="treatment" docUuid="80D9D205-80E7-4D90-8F7D-BB52BCFA7CC4" docUuidSource="ZooBank" docVersion="8" lastPageNumber="539" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="538" updateTime="1698233463814" updateUser="plazi">
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<mods:title id="5EB0FC3902631E31A198FB8C05EE038F">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:name id="C25CCA6FB0576782E066D4E3DE96FF7F" type="personal">
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<mods:namePart id="F217662FF942C35924B62F5C2EF6D23F">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="233117DF322141D6595827FEBAE2BCF9">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="C4AB1ADD859B8C4DEE9B5998488A21AA">text</mods:typeOfResource>
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<mods:title id="B7DFFBDC2E56CBAE949473706D4BEDB4">Zootaxa</mods:title>
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<mods:date id="26A3E133292206AB65676C0BB630F67D">2016</mods:date>
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<mods:number id="36A9763E84F0410EC6140ABFC64F764C">4098</mods:number>
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<treatment id="8A1EB74F8976FF9DC7F362EFFC8B92FB" ID-DOI="http://doi.org/10.5281/zenodo.6056385" ID-GBIF-Taxon="127620797" ID-Zenodo-Dep="6056385" LSID="urn:lsid:zoobank.org:act:80D9D205-80E7-4D90-8F7D-BB52BCFA7CC4" httpUri="http://treatment.plazi.org/id/8A1EB74F8976FF9DC7F362EFFC8B92FB" lastPageId="10" lastPageNumber="539" pageId="9" pageNumber="538">
<subSubSection id="4AAD55D28976FF9EC7F362EFFDBC91C9" box="[151,549,151,178]" pageId="9" pageNumber="538" type="nomenclature">
<paragraph id="020806598976FF9EC7F362EFFDBC91C9" blockId="9.[151,549,151,211]" box="[151,549,151,178]" pageId="9" pageNumber="538">
<heading id="5940B1358976FF9EC7F362EFFDBC91C9" bold="true" box="[151,549,151,178]" fontSize="11" level="1" pageId="9" pageNumber="538" reason="1">
<emphasis id="30C3DA4B8976FF9EC7F362EFFDBC91C9" bold="true" box="[151,549,151,178]" pageId="9" pageNumber="538">
7.
<taxonomicName id="C5B77DDA8976FF9EC7D062E0FE3C91CA" ID-CoL="9WYZ9" box="[180,421,151,177]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="socculus" status="sp. nov.">
<emphasis id="30C3DA4B8976FF9EC7D062E0FE3C91CA" bold="true" box="[180,421,151,177]" italics="true" pageId="9" pageNumber="538">Protosticta socculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308976FF9EC6C962EFFDBC91C9" box="[429,549,152,178]" pageId="9" pageNumber="538" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28976FF9EC7F362CDFEA791A8" box="[151,318,186,211]" pageId="9" pageNumber="538" type="description">
<paragraph id="020806598976FF9EC7F362CDFEA791A8" blockId="9.[151,549,151,211]" box="[151,318,186,211]" pageId="9" pageNumber="538">
(
<figureCitation id="9A8C1ADC8976FF9EC7FA62CDFF6491A8" box="[158,253,186,211]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="9" pageNumber="538">Figure 7</figureCitation>
AK)
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28976FF9EC7F36375FC2E9018" pageId="9" pageNumber="538" type="materials_examined">
<paragraph id="020806598976FF9EC7F36375FC2E9018" blockId="9.[151,1437,258,427]" pageId="9" pageNumber="538">
<emphasis id="30C3DA4B8976FF9EC7F36375FECD9060" bold="true" box="[151,340,258,283]" pageId="9" pageNumber="538">
<typeStatus id="DD0CB8FB8976FF9EC7F36375FF4B9060" box="[151,210,258,283]" pageId="9" pageNumber="538">Type</typeStatus>
specimens
</emphasis>
.
<emphasis id="30C3DA4B8976FF9EC6006375FE489060" bold="true" box="[356,465,258,283]" pageId="9" pageNumber="538">
<typeStatus id="DD0CB8FB8976FF9EC6006375FE489060" box="[356,465,258,283]" pageId="9" pageNumber="538" type="holotype">Holotype</typeStatus>
</emphasis>
: male, Aso, Dong Giang district, Quang Nam Province, central
<collectingCountry id="7AA046C98976FF9EC3D06375FA8F9060" box="[1204,1302,258,283]" name="Vietnam" pageId="9" pageNumber="538">Vietnam</collectingCountry>
,
<date id="760920998976FF9EC2476375FA019060" box="[1315,1432,258,283]" pageId="9" pageNumber="538" value="2015-05-28">28.V.2015</date>
, Quoc Toan Phan leg.;
<emphasis id="30C3DA4B8976FF9EC6F66351FD939044" bold="true" box="[402,522,294,319]" pageId="9" pageNumber="538">
<typeStatus id="DD0CB8FB8976FF9EC6F66351FD939044" box="[402,522,294,319]" pageId="9" pageNumber="538" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="14B1CDD08976FF9EC57D6350FDEB9044" box="[537,626,294,319]" pageId="9" pageNumber="538" type="male">4 males</specimenCount>
, same date, location and collector as the
<typeStatus id="DD0CB8FB8976FF9EC3246351FB3C9044" box="[1088,1189,294,319]" pageId="9" pageNumber="538" type="holotype">holotype</typeStatus>
;
<specimenCount id="14B1CDD08976FF9EC3D56350FA929044" box="[1201,1291,294,319]" pageId="9" pageNumber="538" type="male">4 males</specimenCount>
, Bhalee, Tay Giang district, Quang Nam Province,
<date id="760920998976FF9EC55B633DFD589018" box="[575,705,330,355]" pageId="9" pageNumber="538" value="2015-09-19">19.IX.2015</date>
, Quoc Toan Phan leg.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28976FF9EC7A36319FC1290D1" pageId="9" pageNumber="538" type="etymology">
<paragraph id="020806598976FF9EC7A36319FC1290D1" blockId="9.[151,1437,258,427]" pageId="9" pageNumber="538">
<emphasis id="30C3DA4B8976FF9EC7A36319FEDF90FC" bold="true" box="[199,326,366,391]" pageId="9" pageNumber="538">Etymology</emphasis>
. The specific name
<taxonomicName id="C5B77DDA8976FF9EC5586319FD0590FC" box="[572,668,366,391]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="socculus">
<emphasis id="30C3DA4B8976FF9EC5586319FD0590FC" box="[572,668,366,391]" italics="true" pageId="9" pageNumber="538">socculus</emphasis>
</taxonomicName>
is derived from the Latin word “
<taxonomicName id="C5B77DDA8976FF9EC3496319FB0B90FC" box="[1069,1170,366,391]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="socculus">socculus</taxonomicName>
” (meaning slipper), in reference to the slipper-shaped structure at the tip of the paraprocts.
</paragraph>
</subSubSection>
<caption id="56C856D18976FF9EC7F364D4FC4A966E" httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="9" pageNumber="538" targetBox="[151,1432,481,1664]" targetPageId="9">
<paragraph id="020806598976FF9EC7F364D4FC4A966E" blockId="9.[151,1436,1699,1813]" pageId="9" pageNumber="538">
<emphasis id="30C3DA4B8976FF9EC7F364D4FE9497C2" bold="true" box="[151,269,1699,1721]" pageId="9" pageNumber="538">FIGURE 7</emphasis>
.
<taxonomicName id="C5B77DDA8976FF9EC67E64D3FE7197C2" box="[282,488,1699,1721]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="socculus" status="sp. nov.">
<emphasis id="30C3DA4B8976FF9EC67E64D3FE7197C2" box="[282,488,1699,1721]" italics="true" pageId="9" pageNumber="538">Protosticta socculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308976FF9EC68A64D3FDCD97C2" box="[494,596,1700,1721]" pageId="9" pageNumber="538" rank="species">spec. nov.</taxonomicNameLabel>
[AH] holotype male—(
<emphasis id="30C3DA4B8976FF9EC43264D4FCF197C3" bold="true" box="[854,872,1699,1720]" pageId="9" pageNumber="538">A</emphasis>
) head &amp; thorax in lateral view; (
<emphasis id="30C3DA4B8976FF9EC3DE64D3FB5297C2" bold="true" box="[1210,1227,1700,1721]" pageId="9" pageNumber="538">B</emphasis>
) prothorax in dorsal view; anal appendages in (
<emphasis id="30C3DA4B8976FF9EC6C764B5FE2C97A3" bold="true" box="[419,437,1730,1752]" pageId="9" pageNumber="538">C</emphasis>
) dorsal &amp; (
<emphasis id="30C3DA4B8976FF9EC54D64B5FDA297AC" bold="true" box="[553,571,1730,1751]" pageId="9" pageNumber="538">D</emphasis>
) lateral view; (
<emphasis id="30C3DA4B8976FF9EC5B164B5FD7F97AC" bold="true" box="[725,742,1730,1751]" pageId="9" pageNumber="538">E</emphasis>
) right cercus and paraproct in oblique-dorsal view; (
<emphasis id="30C3DA4B8976FF9EC39C64B5FA9E97AC" bold="true" box="[1272,1287,1730,1751]" pageId="9" pageNumber="538">F</emphasis>
) tip of cerci in posteroventral view; (
<emphasis id="30C3DA4B8976FF9EC6116497FE10978D" bold="true" box="[373,393,1760,1782]" pageId="9" pageNumber="538">G</emphasis>
) paraprocts in ventral view; (
<emphasis id="30C3DA4B8976FF9EC5D26496FD53978D" bold="true" box="[694,714,1761,1782]" pageId="9" pageNumber="538">H</emphasis>
) penile organ in dorsal view; (
<emphasis id="30C3DA4B8976FF9EC3676496FB94978D" bold="true" box="[1027,1037,1761,1782]" pageId="9" pageNumber="538">I</emphasis>
) wings; [JK] paratype male—(
<emphasis id="30C3DA4B8976FF9EC2326496FAFB978D" bold="true" box="[1366,1378,1761,1782]" pageId="9" pageNumber="538">J</emphasis>
) anal appendages in lateral view; (
<emphasis id="30C3DA4B8976FF9EC6DE6488FE57966F" bold="true" box="[442,462,1791,1812]" pageId="9" pageNumber="538">K</emphasis>
) right cercus and paraproct in oblique-dorsal view.
</paragraph>
</caption>
<subSubSection id="4AAD55D28976FF9DC7A3654BFC8B92FB" lastPageId="10" lastPageNumber="539" pageId="9" pageNumber="538" type="description">
<paragraph id="020806598976FF9EC7A3654BFA05969E" blockId="9.[151,1436,1852,2021]" pageId="9" pageNumber="538">
<emphasis id="30C3DA4B8976FF9EC7A3654BFDB1962E" bold="true" box="[199,552,1852,1877]" pageId="9" pageNumber="538">
Description of male
<typeStatus id="DD0CB8FB8976FF9EC6A5654BFDB1962E" box="[449,552,1852,1877]" pageId="9" pageNumber="538" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="30C3DA4B8976FF9EC55C654BFDED962E" box="[568,628,1852,1877]" italics="true" pageId="9" pageNumber="538">Head</emphasis>
(
<figureCitation id="9A8C1ADC8976FF9EC5EC654BFD57962E" box="[648,718,1852,1877]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="9" pageNumber="538">Fig. 7</figureCitation>
A). Anteclypeus and labrum pale blue with inferior margin of labrum black. Antennae black, basal half of second segment pale yellow; postclypeus, genae, frons, vertex and occiput entirely black.
<emphasis id="30C3DA4B8976FF9EC6C565F3FD8A96E6" box="[417,531,1924,1949]" italics="true" pageId="9" pageNumber="538">Prothorax</emphasis>
(
<figureCitation id="9A8C1ADC8976FF9EC54265F3FDF596E6" box="[550,620,1924,1949]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="9" pageNumber="538">Fig. 7</figureCitation>
B) almost entirely pale yellow, with anterior and posterior lobes black dorsally.
<emphasis id="30C3DA4B8976FF9EC79A65DFFEF696BA" box="[254,367,1960,1985]" italics="true" pageId="9" pageNumber="538">Synthorax</emphasis>
dull black with pale yellow stripe over metepisternum, black line over interpleural suture and mostly pale yellow metepimeron.
<emphasis id="30C3DA4B8976FF9EC57265B9FDD2969E" box="[534,587,1998,2021]" italics="true" pageId="9" pageNumber="538">Legs</emphasis>
pale yellow with black stripe over flexor surface; spines black.
<emphasis id="30C3DA4B8976FF9EC27E65BAFAC7969F" box="[1306,1374,1997,2020]" italics="true" pageId="9" pageNumber="538">Wings</emphasis>
(
<figureCitation id="9A8C1ADC8976FF9EC20A65BBFA05969E" box="[1390,1436,1996,2021]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="9" pageNumber="538">Fig.</figureCitation>
</paragraph>
<paragraph id="020806598975FF9DC7F362EFFA1D901F" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
<date id="760920998975FF9DC7F362EFFF3691CB" box="[151,175,152,176]" pageId="10" pageNumber="539">7I</date>
) hyaline, 1315 postnodal veins in both wings. Pterostigma brown.
<emphasis id="30C3DA4B8975FF9DC4AF62E0FBAC91CB" box="[971,1077,151,176]" italics="true" pageId="10" pageNumber="539">Abdomen</emphasis>
long and thin, black with pale yellow markings as follows: S12 short, ventral half pale yellow; S37 very long, blackish brown with whitish anterior ring and indistinct brown sup-apical ring; S89 shorter, dorsum of S8 blackish brown, large part of basal segment bluish white; S9 bluish white with anterior and posterior margin black; S10 entirely black.
<emphasis id="30C3DA4B8975FF9DC2036373FE84903B" italics="true" pageId="10" pageNumber="539">Anal appendages</emphasis>
(
<figureCitation id="9A8C1ADC8975FF9DC6496350FEE2903B" box="[301,379,295,320]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Figs. 7</figureCitation>
CG) black; cerci strongly curved downward; paraprocts with elongated tip, tapering, with a prominent slipper-shaped cup-like structure bearing a long spine on its distal margin.
<emphasis id="30C3DA4B8975FF9DC330633BFB7C901F" box="[1108,1253,332,357]" italics="true" pageId="10" pageNumber="539">Penile organ</emphasis>
as in
<figureCitation id="9A8C1ADC8975FF9DC243633BFAF1901F" box="[1319,1384,332,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Fig. 7</figureCitation>
H.
</paragraph>
<paragraph id="020806598975FF9DC7A36318FC2890F3" blockId="10.[151,1437,151,896]" box="[199,945,367,392]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A36318FEEF90F3" bold="true" box="[199,374,367,392]" pageId="10" pageNumber="539">Measurements</emphasis>
. Hindwing 19; abdomen including appendages 44.
</paragraph>
<paragraph id="020806598975FF9DC7A363E3FE3B90D6" blockId="10.[151,1437,151,896]" box="[199,418,404,429]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A363E3FE8790D6" bold="true" box="[199,286,404,429]" pageId="10" pageNumber="539">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="020806598975FF9DC7A363C0FEA49363" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A363C0FD8490AB" bold="true" box="[199,541,439,464]" pageId="10" pageNumber="539">
Variation in
<typeStatus id="DD0CB8FB8975FF9DC60763C0FE5790AB" box="[355,462,439,464]" pageId="10" pageNumber="539" type="paratype">paratype</typeStatus>
males
</emphasis>
. In the
<typeStatus id="DD0CB8FB8975FF9DC51863CFFD4590AB" box="[636,732,440,464]" pageId="10" pageNumber="539" type="paratype">paratype</typeStatus>
males, the slipper-shaped cup-like structure at the tip of the paraprocts is not as prominent as in the
<typeStatus id="DD0CB8FB8975FF9DC53363ABFD20908E" box="[599,697,476,501]" pageId="10" pageNumber="539" type="holotype">holotype</typeStatus>
(
<figureCitation id="9A8C1ADC8975FF9DC5AC63ABFC8F908F" box="[712,790,476,501]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Figs. 7</figureCitation>
JK), but the spine at the distal margin can be seen easily in lateral view.
</paragraph>
<paragraph id="020806598975FF9DC7A36053FB7B9347" blockId="10.[151,1437,151,896]" box="[199,1250,548,573]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A36053FDCB9346" bold="true" box="[199,594,548,573]" pageId="10" pageNumber="539">
Measurements of
<typeStatus id="DD0CB8FB8975FF9DC6FF6053FD9E9346" box="[411,519,548,573]" pageId="10" pageNumber="539" type="paratype">paratype</typeStatus>
males
</emphasis>
. Hindwing 1920; abdomen including appendages 4344.
</paragraph>
<paragraph id="020806598975FF9DC7A36030FBEB93D3" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A36030FE5E931B" bold="true" box="[199,455,583,608]" pageId="10" pageNumber="539">Differential diagnosis</emphasis>
.
<emphasis id="30C3DA4B8975FF9DC6BC603EFDC8931B" box="[472,593,583,608]" italics="true" pageId="10" pageNumber="539">P. s o c cu l us</emphasis>
spec. nov. appears to be closely related to
<taxonomicName id="C5B77DDA8975FF9DC327603EFA80931B" box="[1091,1305,583,608]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8975FF9DC327603EFA80931B" box="[1091,1305,583,608]" italics="true" pageId="10" pageNumber="539">P. khaosoidaoensis</emphasis>
</taxonomicName>
judging by the elongated shape of the paraprocts, which both have a very large cup-like structure with a spine at the distal margin on their apex on the dorsal side. It can be separated from that species as follows:
</paragraph>
<paragraph id="020806598975FF9DC7A360C3FD36938B" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
(1) In
<taxonomicName id="C5B77DDA8975FF9DC67260C2FE0E93B6" box="[278,407,692,717]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="socculus" status="sp. nov.">
<emphasis id="30C3DA4B8975FF9DC67260C2FE0E93B6" box="[278,407,692,717]" italics="true" pageId="10" pageNumber="539">P. socculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308975FF9DC6C660C2FD8093B7" box="[418,537,693,716]" pageId="10" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
, cerci with only one protuberance ventrally at midpoint (
<figureCitation id="9A8C1ADC8975FF9DC3A460C3FA8B93B7" box="[1216,1298,692,717]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Figs. 7</figureCitation>
C, 7E), two protuberances in
<taxonomicName id="C5B77DDA8975FF9DC63C60AEFDB5938B" box="[344,556,727,752]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8975FF9DC63C60AEFDB5938B" box="[344,556,727,752]" italics="true" pageId="10" pageNumber="539">P. khaosoidaoensis</emphasis>
</taxonomicName>
(
<figureCitation id="9A8C1ADC8975FF9DC55860A0FD17938B" box="[572,654,727,752]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="10" pageNumber="539">Fig. 10</figureCitation>
B).
</paragraph>
<paragraph id="020806598975FF9DC7A3608BFE4A9243" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
(2) In
<taxonomicName id="C5B77DDA8975FF9DC66B608AFE15926E" box="[271,396,764,789]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="socculus" status="sp. nov.">
<emphasis id="30C3DA4B8975FF9DC66B608AFE15926E" box="[271,396,764,789]" italics="true" pageId="10" pageNumber="539">P. socculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308975FF9DC6F0608AFD9E926F" box="[404,519,765,788]" pageId="10" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
, the tip of the cerci is structurally complex (
<figureCitation id="9A8C1ADC8975FF9DC364608BFBDD926F" box="[1024,1092,764,789]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Fig. 7</figureCitation>
F), flattened and rounded in
<taxonomicName id="C5B77DDA8975FF9DC2E3608AFEC99243" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8975FF9DC2E3608AFEC99243" italics="true" pageId="10" pageNumber="539">P. khaosoidaoensis</emphasis>
</taxonomicName>
(
<figureCitation id="9A8C1ADC8975FF9DC63B6168FE2B9243" box="[351,434,799,824]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="10" pageNumber="539">Fig. 10</figureCitation>
C).
</paragraph>
<paragraph id="020806598975FF9DC7A36133FC8B92FB" blockId="10.[151,1437,151,896]" pageId="10" pageNumber="539">
(3) In
<taxonomicName id="C5B77DDA8975FF9DC6766132FE099226" box="[274,400,836,861]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="socculus" status="sp. nov.">
<emphasis id="30C3DA4B8975FF9DC6766132FE099226" box="[274,400,836,861]" italics="true" pageId="10" pageNumber="539">P. socculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308975FF9DC6FE6132FD969227" box="[410,527,837,860]" pageId="10" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
, the dorsum of the synthorax is completely black (
<figureCitation id="9A8C1ADC8975FF9DC33F6133FB399227" box="[1115,1184,836,861]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="10" pageNumber="539">Fig. 7</figureCitation>
A), a pale yellow line along the dorsal carina in
<taxonomicName id="C5B77DDA8975FF9DC6DD611EFD1492FB" box="[441,653,871,896]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8975FF9DC6DD611EFD1492FB" box="[441,653,871,896]" italics="true" pageId="10" pageNumber="539">P. khaosoidaoensis</emphasis>
</taxonomicName>
(
<figureCitation id="9A8C1ADC8975FF9DC5F86110FD7792FB" box="[668,750,871,896]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="10" pageNumber="539">Fig. 10</figureCitation>
A).
</paragraph>
</subSubSection>
</treatment>
</document>