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<document id="3ED9780BA8B30B2546539E20CE1EAEB9" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan &amp; Kompier, Tom" docDate="2016" docId="8A1EB74F8975FF9CC7F361A3FB789183" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta spinosa Phan &amp; Kompier, 2016, spec. nov." docType="treatment" docUuid="1698D7F5-4699-40F0-B3D1-92F867ECA79D" docUuidSource="ZooBank" docVersion="8" lastPageNumber="539" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="539" updateTime="1698233463814" updateUser="plazi">
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<mods:title id="54D4FDFB23DF0D128A706263766D7ACC">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="32ADDFE46BF37C3B4E20FD863BBDAF8C">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="14BEF2715DAD2956EA30EDC660DDEE18">Kompier, Tom</mods:namePart>
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<mods:title id="DDD1D832ED2F3EE74CED0D973667CC47">Zootaxa</mods:title>
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<mods:date id="AABEB5EA5C500C2BC809BA543C450430">2016</mods:date>
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<treatment id="8A1EB74F8975FF9CC7F361A3FB789183" ID-DOI="http://doi.org/10.5281/zenodo.6056387" ID-GBIF-Taxon="127620798" ID-Zenodo-Dep="6056387" LSID="urn:lsid:zoobank.org:act:1698D7F5-4699-40F0-B3D1-92F867ECA79D" httpUri="http://treatment.plazi.org/id/8A1EB74F8975FF9CC7F361A3FB789183" lastPageId="11" lastPageNumber="539" pageId="10" pageNumber="539">
<subSubSection id="4AAD55D28975FF9DC7F361A3FD8E9295" box="[151,535,980,1006]" pageId="10" pageNumber="539" type="nomenclature">
<paragraph id="020806598975FF9DC7F361A3FD8E9295" blockId="10.[151,535,980,1039]" box="[151,535,980,1006]" pageId="10" pageNumber="539">
<heading id="5940B1358975FF9DC7F361A3FD8E9295" bold="true" box="[151,535,980,1006]" fontSize="11" level="1" pageId="10" pageNumber="539" reason="1">
<emphasis id="30C3DA4B8975FF9DC7F361A3FD8E9295" bold="true" box="[151,535,980,1006]" pageId="10" pageNumber="539">
8.
<taxonomicName id="C5B77DDA8975FF9DC7D061A3FE019295" ID-CoL="9WYZB" box="[180,408,980,1006]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="30C3DA4B8975FF9DC7D061A3FE019295" bold="true" box="[180,408,980,1006]" italics="true" pageId="10" pageNumber="539">Protosticta spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308975FF9DC6C461A3FD8E9295" box="[416,535,980,1006]" pageId="10" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="4AAD55D28975FF9DC7F36181FEAD9574" box="[151,308,1014,1039]" pageId="10" pageNumber="539" type="description">
<paragraph id="020806598975FF9DC7F36181FEAD9574" blockId="10.[151,535,980,1039]" box="[151,308,1014,1039]" pageId="10" pageNumber="539">
(
<figureCitation id="9A8C1ADC8975FF9DC7FA6181FF659574" box="[158,252,1014,1039]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Figure 8</figureCitation>
AJ)
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28975FF9DC7F36649FB679500" pageId="10" pageNumber="539" type="materials_examined">
<paragraph id="020806598975FF9DC7F36649FB679500" blockId="10.[151,1437,1086,2011]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7F36649FECB952C" bold="true" box="[151,338,1086,1111]" pageId="10" pageNumber="539">
<typeStatus id="DD0CB8FB8975FF9DC7F36649FF4B952C" box="[151,210,1086,1111]" pageId="10" pageNumber="539">Type</typeStatus>
specimens
</emphasis>
.
<emphasis id="30C3DA4B8975FF9DC6046649FE54952C" bold="true" box="[352,461,1086,1111]" pageId="10" pageNumber="539">
<typeStatus id="DD0CB8FB8975FF9DC6046649FE54952C" box="[352,461,1086,1111]" pageId="10" pageNumber="539" type="holotype">Holotype</typeStatus>
</emphasis>
: male and female, Nam Bung, Van Chan district, Yen Bai Province, northern
<collectingCountry id="7AA046C98975FF9DC2526649FA01952C" box="[1334,1432,1086,1111]" name="Vietnam" pageId="10" pageNumber="539">Vietnam</collectingCountry>
,
<date id="760920998975FF9DC7F36614FE839500" box="[151,282,1123,1147]" pageId="10" pageNumber="539" value="2014-06-01">01.VI.2014</date>
. Tom Kompier leg.;
<emphasis id="30C3DA4B8975FF9DC5656615FDF79500" bold="true" box="[513,622,1122,1147]" pageId="10" pageNumber="539">
<typeStatus id="DD0CB8FB8975FF9DC5656615FDF79500" box="[513,622,1122,1147]" pageId="10" pageNumber="539" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="14B1CDD08975FF9DC5186614FD539500" box="[636,714,1122,1147]" pageId="10" pageNumber="539" type="male">1 male</specimenCount>
, same date, location and collector as the
<typeStatus id="DD0CB8FB8975FF9DC3F06615FB639500" box="[1172,1274,1122,1147]" pageId="10" pageNumber="539" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28975FF9DC7A366F1FBEB95E5" box="[199,1138,1158,1183]" pageId="10" pageNumber="539" type="etymology">
<paragraph id="020806598975FF9DC7A366F1FBEB95E5" blockId="10.[151,1437,1086,2011]" box="[199,1138,1158,1183]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A366F1FEDF95E4" bold="true" box="[199,326,1158,1183]" pageId="10" pageNumber="539">Etymology</emphasis>
. The specific name &quot;
<taxonomicName id="C5B77DDA8975FF9DC55066F0FD1095E5" box="[564,649,1159,1182]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="30C3DA4B8975FF9DC55066F0FD1095E5" box="[564,649,1159,1182]" italics="true" pageId="10" pageNumber="539">spinosa</emphasis>
</taxonomicName>
&quot; refers to the large spine on the paraprocts.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28975FF9CC7A366DDFB789183" lastPageId="11" lastPageNumber="540" pageId="10" pageNumber="539" type="description">
<paragraph id="020806598975FF9DC7A366DDFB9097EC" blockId="10.[151,1437,1086,2011]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A366DDFDBF95B8" bold="true" box="[199,550,1194,1219]" pageId="10" pageNumber="539">
Description of male
<typeStatus id="DD0CB8FB8975FF9DC6A466DDFDBF95B8" box="[448,550,1194,1219]" pageId="10" pageNumber="539" type="holotype">holotype</typeStatus>
</emphasis>
. A small and colorful
<taxonomicName id="C5B77DDA8975FF9DC45566DBFC3495B8" box="[817,941,1196,1219]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="genus">
<emphasis id="30C3DA4B8975FF9DC45566DBFC3495B8" box="[817,941,1196,1219]" italics="true" pageId="10" pageNumber="539">Protosticta</emphasis>
</taxonomicName>
with relatively short abdomen.
<emphasis id="30C3DA4B8975FF9DC24466DDFAC495B8" box="[1312,1373,1194,1219]" italics="true" pageId="10" pageNumber="539">Head</emphasis>
(
<figureCitation id="9A8C1ADC8975FF9DC20A66DDFF3C959C" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
A). Anteclypeus, genae whitish; labrum blueish yellow, anterior margin black. Frons and top of head are glossy greenish black. The first and second segments of antennae pale yellow, the remainder black.
<emphasis id="30C3DA4B8975FF9DC3DD6685FAB29470" box="[1209,1323,1266,1291]" italics="true" pageId="10" pageNumber="539">Prothorax</emphasis>
(
<figureCitation id="9A8C1ADC8975FF9DC2586685FAE69470" box="[1340,1407,1266,1291]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
B) yellowish, anterior lobe of pronotum yellow; middle and posterior lobes yellow laterally, but with clear-cut black mark over middle line and contrasting with black propleuron.
<emphasis id="30C3DA4B8975FF9DC407674DFC4C9428" box="[867,981,1338,1363]" italics="true" pageId="10" pageNumber="539">Synthorax</emphasis>
glossy greenish black with pale yellow stripe over metepisternum. Lower half of metepimeron and venter pale yellow.
<emphasis id="30C3DA4B8975FF9DC3406728FBC0940D" box="[1060,1113,1375,1398]" italics="true" pageId="10" pageNumber="539">Legs</emphasis>
and
<emphasis id="30C3DA4B8975FF9DC3F76728FB4D940D" box="[1171,1236,1375,1398]" italics="true" pageId="10" pageNumber="539">coxae</emphasis>
pale yellow with grey stripe over flexor surface and black spines.
<emphasis id="30C3DA4B8975FF9DC5B467F3FC8A94E0" box="[720,787,1412,1435]" italics="true" pageId="10" pageNumber="539">Wings</emphasis>
(
<figureCitation id="9A8C1ADC8975FF9DC44367F5FCF294E0" box="[807,875,1410,1435]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
G) hyaline with 16 postnodal veins. Pterostigma brown.
<emphasis id="30C3DA4B8975FF9DC78867D1FECF94C4" box="[236,342,1446,1471]" italics="true" pageId="10" pageNumber="539">Abdomen</emphasis>
relatively short, blackish brown with yellow markings as follows: S1 ventral half pale yellow; S2 dorsally with pale yellow stripe widest at proximal side, lower half pale yellow; S37 with prominent pale yellow basal ring, becoming more yellow at ventral side on S7; S8 black with large bright yellow ventral mark, widest at base, S9 black with indistinct yellow line along venter; S10 black.
<emphasis id="30C3DA4B8975FF9DC4CC6465FBF79750" box="[936,1134,1554,1579]" italics="true" pageId="10" pageNumber="539">Anal appendages</emphasis>
(
<figureCitation id="9A8C1ADC8975FF9DC3E46465FB569750" box="[1152,1231,1554,1579]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Figs. 8</figureCitation>
CE) black; cerci yellowish at base, only one median protuberance ventrally; tip smooth. Basal half of paraprocts black, apical half amber colored; a large spine directed inward at midpoint; tips directed inward, nearly touching, then curving out again, their outer sides hollowed out (
<figureCitation id="9A8C1ADC8975FF9DC55A6409FD1997EC" box="[574,640,1662,1687]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
E).
<emphasis id="30C3DA4B8975FF9DC5CC6409FCA097ED" box="[680,825,1662,1687]" italics="true" pageId="10" pageNumber="539">Penile organ</emphasis>
shaped as
<figureCitation id="9A8C1ADC8975FF9DC4D76409FC6A97EC" box="[947,1011,1662,1687]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">
Fig.
<specimenCount id="14B1CDD08975FF9DC4826408FC6A97EC" box="[998,1011,1663,1687]" pageId="10" pageNumber="539" type="generic">8</specimenCount>
</figureCitation>
F.
</paragraph>
<paragraph id="020806598975FF9DC7A364D5FBFE9678" blockId="10.[151,1437,1086,2011]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A364D5FE8797C0" bold="true" box="[199,286,1698,1723]" pageId="10" pageNumber="539">Female</emphasis>
. Color pattern as
<typeStatus id="DD0CB8FB8975FF9DC68064D5FDDF97C0" box="[484,582,1698,1723]" pageId="10" pageNumber="539" type="holotype">holotype</typeStatus>
male (
<figureCitation id="9A8C1ADC8975FF9DC5F564D5FD4497C0" box="[657,733,1698,1723]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Figs. 8</figureCitation>
HI). Wings hyaline with 16 postnodal veins. Abdominal color pattern as follows: S1 with a large oval spot at base, S27 with prominent pale yellow basal ring; S810 black, but S8 with large pale yellow mark on ventral half laterally. Ovipositor structure as
<figureCitation id="9A8C1ADC8975FF9DC370649DFBCF9678" box="[1044,1110,1770,1795]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
J.
</paragraph>
<paragraph id="020806598975FF9DC7A36579FE2C9630" blockId="10.[151,1437,1086,2011]" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A36579FEEF965C" bold="true" box="[199,374,1806,1831]" pageId="10" pageNumber="539">Measurements</emphasis>
. Male. Hindwing 22; abdomen including appendages 35. Female.
<collectingCountry id="7AA046C98975FF9DC3E66579FB23965C" box="[1154,1210,1806,1831]" name="India" pageId="10" pageNumber="539">Hind</collectingCountry>
wing 25; abdomen including appendages 36.
</paragraph>
<paragraph id="020806598975FF9CC7A36521FB789183" blockId="10.[151,1437,1086,2011]" lastBlockId="11.[151,1437,151,248]" lastPageId="11" lastPageNumber="540" pageId="10" pageNumber="539">
<emphasis id="30C3DA4B8975FF9DC7A36521FE509614" bold="true" box="[199,457,1878,1903]" pageId="10" pageNumber="539">Differential diagnosis</emphasis>
.
<taxonomicName id="C5B77DDA8975FF9DC6BE6520FDD69615" box="[474,591,1879,1902]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="30C3DA4B8975FF9DC6BE6520FDD69615" box="[474,591,1879,1902]" italics="true" pageId="10" pageNumber="539">P. spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308975FF9DC53D6520FD529615" box="[601,715,1879,1902]" pageId="10" pageNumber="539" rank="species">spec. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="C5B77DDA8975FF9DC40D6520FA019614" authority="Kiran, Kalesh and Kunte, 2015" authorityName="Kiran, Kalesh and Kunte" authorityYear="2015" box="[873,1432,1878,1903]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="ponmudiensis">
<emphasis id="30C3DA4B8975FF9DC40D6520FBBB9614" box="[873,1058,1878,1903]" italics="true" pageId="10" pageNumber="539">P. ponmudiensis</emphasis>
<bibRefCitation id="66267BA88975FF9DC3486521FA019614" author="Kiran" box="[1068,1432,1878,1903]" pageId="10" pageNumber="539" refString="Kiran, C. G., Kalesh, S. &amp; Kunte, K. (2015) A new species of damselfly, Protosticta ponmudiensis (Odonata: Zygoptera: Platystictidae) from Ponmudi Hills in the Western Ghats of India. Journal of Threatened Taxa, 7 (5), 7146 - 7151. http: // dx. doi. org / 10.11609 / JoTT. o 4145.7146 - 51" type="journal article" year="2015">Kiran, Kalesh and Kunte, 2015</bibRefCitation>
</taxonomicName>
, from the Western Ghats of
<collectingCountry id="7AA046C98975FF9DC6A7650DFE6496E8" box="[451,509,1914,1939]" name="India" pageId="10" pageNumber="539">India</collectingCountry>
by the paraprocts having a large spine directed inward (
<figureCitation id="9A8C1ADC8975FF9DC30F650DFB3296E8" box="[1131,1195,1914,1939]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
E &amp;
<figureCitation id="9A8C1ADC8975FF9DC385650DFABB96E8" box="[1249,1314,1914,1939]" captionStart="FIGURE 3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[169,1425,470,1883]" captionTargetId="figure@4.[150,1436,465,1891]" captionTargetPageId="4" captionText="FIGURE 3. Protosticta grandis, Cao Bang. [A E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257726/files/figure.png" pageId="10" pageNumber="539">Fig. 3</figureCitation>
C, p. 7149,
<bibRefCitation id="66267BA88975FF9DC7F365E9FEC496CC" author="Kiran" box="[151,349,1950,1975]" pageId="10" pageNumber="539" refString="Kiran, C. G., Kalesh, S. &amp; Kunte, K. (2015) A new species of damselfly, Protosticta ponmudiensis (Odonata: Zygoptera: Platystictidae) from Ponmudi Hills in the Western Ghats of India. Journal of Threatened Taxa, 7 (5), 7146 - 7151. http: // dx. doi. org / 10.11609 / JoTT. o 4145.7146 - 51" type="journal article" year="2015">
Kiran
<emphasis id="30C3DA4B8975FF9DC7BB65E8FE8196CC" box="[223,280,1950,1975]" italics="true" pageId="10" pageNumber="539">et al.</emphasis>
2015
</bibRefCitation>
). However,
<emphasis id="30C3DA4B8975FF9DC68165E8FDCC96CD" box="[485,597,1951,1974]" italics="true" pageId="10" pageNumber="539">P. s pi no s a</emphasis>
spec. nov. differs from
<taxonomicName id="C5B77DDA8975FF9DC40C65E8FB8796CC" box="[872,1054,1950,1975]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="ponmudiensis">
<emphasis id="30C3DA4B8975FF9DC40C65E8FB8796CC" box="[872,1054,1950,1975]" italics="true" pageId="10" pageNumber="539">P. ponmudiensis</emphasis>
</taxonomicName>
by the difference in color pattern and by the structure of the cerci, which are stout with a ventral protuberance in
<emphasis id="30C3DA4B8975FF9DC37165B3FB1A96A0" box="[1045,1155,1988,2011]" italics="true" pageId="10" pageNumber="539">P. s p i n os a</emphasis>
spec. nov. (
<figureCitation id="9A8C1ADC8975FF9DC26965B5FAD796A0" box="[1293,1358,1986,2011]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="10" pageNumber="539">Fig. 8</figureCitation>
E), but slender with a small inner spine in
<taxonomicName id="C5B77DDA8974FF9CC57962EEFD4B91CB" box="[541,722,151,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="540" phylum="Arthropoda" rank="species" species="ponmudiensis">
<emphasis id="30C3DA4B8974FF9CC57962EEFD4B91CB" box="[541,722,151,176]" italics="true" pageId="11" pageNumber="540">P. ponmudiensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="66267BA88974FF9CC58562E0FC3D91CB" author="Kiran" box="[737,932,151,176]" pageId="11" pageNumber="540" refString="Kiran, C. G., Kalesh, S. &amp; Kunte, K. (2015) A new species of damselfly, Protosticta ponmudiensis (Odonata: Zygoptera: Platystictidae) from Ponmudi Hills in the Western Ghats of India. Journal of Threatened Taxa, 7 (5), 7146 - 7151. http: // dx. doi. org / 10.11609 / JoTT. o 4145.7146 - 51" type="journal article" year="2015">
Kiran
<emphasis id="30C3DA4B8974FF9CC44D62EEFCF891CB" box="[809,865,151,176]" italics="true" pageId="11" pageNumber="540">et al.</emphasis>
2015
</bibRefCitation>
: p. 7150,
<figureCitation id="9A8C1ADC8974FF9CC37462E0FBCD91CB" box="[1040,1108,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1687,1709]" captionTargetBox="[159,1432,476,1674]" captionTargetId="figure@3.[151,1437,469,1675]" captionTargetPageId="3" captionText="FIGURE 2. Protosticta caroli, Ba Na. [A G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H J] female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." httpUri="https://zenodo.org/record/257725/files/figure.png" pageId="11" pageNumber="540">Fig. 2</figureCitation>
J); the tip of the paraprocts of
<emphasis id="30C3DA4B8974FF9CC7F362CAFE9D91AF" box="[151,260,189,212]" italics="true" pageId="11" pageNumber="540">P. s p i n os a</emphasis>
spec. nov. are curved abruptly inward, then outward and hollowed out, almost like a slender spoon (
<figureCitation id="9A8C1ADC8974FF9CC20062CBFF3D9183" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="11" pageNumber="540">Figs. 8</figureCitation>
DE), but the tips are smooth and broad in
<taxonomicName id="C5B77DDA8974FF9CC5EC6296FCA79183" box="[648,830,223,248]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="540" phylum="Arthropoda" rank="species" species="ponmudiensis">
<emphasis id="30C3DA4B8974FF9CC5EC6296FCA79183" box="[648,830,223,248]" italics="true" pageId="11" pageNumber="540">P. ponmudiensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="66267BA88974FF9CC42862A8FB889183" author="Kiran" box="[844,1041,223,248]" pageId="11" pageNumber="540" refString="Kiran, C. G., Kalesh, S. &amp; Kunte, K. (2015) A new species of damselfly, Protosticta ponmudiensis (Odonata: Zygoptera: Platystictidae) from Ponmudi Hills in the Western Ghats of India. Journal of Threatened Taxa, 7 (5), 7146 - 7151. http: // dx. doi. org / 10.11609 / JoTT. o 4145.7146 - 51" type="journal article" year="2015">
Kiran
<emphasis id="30C3DA4B8974FF9CC4F16296FC549183" box="[917,973,223,248]" italics="true" pageId="11" pageNumber="540">et al.</emphasis>
2015
</bibRefCitation>
: p. 7149,
<figureCitation id="9A8C1ADC8974FF9CC31962A8FB279183" box="[1149,1214,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[169,1425,470,1883]" captionTargetId="figure@4.[150,1436,465,1891]" captionTargetPageId="4" captionText="FIGURE 3. Protosticta grandis, Cao Bang. [A E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257726/files/figure.png" pageId="11" pageNumber="540">Fig. 3</figureCitation>
C).
</paragraph>
</subSubSection>
</treatment>
</document>