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<document id="16389D91B074B1511563D6C920471B75" ID-DOI="10.11646/zootaxa.4098.3.6" ID-GBIF-Dataset="687e2f7e-8268-4707-9cd7-bf46f9753f3b" ID-ISSN="1175-5326" ID-Zenodo-Dep="257723" ID-ZooBank="A005DB6C-7389-4506-B2D5-392531F832EB" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460012672336" checkinUser="plazi" docAuthor="Phan, Quoc Toan &amp; Kompier, Tom" docDate="2016" docId="8A1EB74F8972FF99C7F360AFFE95903B" docLanguage="en" docName="zootaxa.4098.3.6.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protosticta khaosoidaoensis Asahina 1984" docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="7627CF37897FFF97C7646277FF99917B" masterDocTitle="A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)" masterLastPageNumber="544" masterPageNumber="529" pageNumber="542" updateTime="1698233463814" updateUser="plazi">
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<mods:title id="BCB012FE4F98A6133EC194F8E1B63B38">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="2034724AF5CD4FA8AFB7DDF291A3E4E8">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="32241C93079EC5B0AD85C7E1CB1BA457">Kompier, Tom</mods:namePart>
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<mods:date id="5E1962A2B8B4E9F6967645DED64ABD1B">2016</mods:date>
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<treatment id="8A1EB74F8972FF99C7F360AFFE95903B" ID-DOI="http://doi.org/10.5281/zenodo.6056391" ID-GBIF-Taxon="127620800" ID-Zenodo-Dep="6056391" LSID="urn:lsid:plazi:treatment:8A1EB74F8972FF99C7F360AFFE95903B" httpUri="http://treatment.plazi.org/id/8A1EB74F8972FF99C7F360AFFE95903B" lastPageId="14" lastPageNumber="543" pageId="13" pageNumber="542">
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<paragraph id="020806598972FF9AC7F360AFFC299389" blockId="13.[151,944,727,787]" box="[151,944,727,754]" pageId="13" pageNumber="542">
<heading id="5940B1358972FF9AC7F360AFFC299389" bold="true" box="[151,944,727,754]" fontSize="11" level="1" pageId="13" pageNumber="542" reason="1">
<emphasis id="30C3DA4B8972FF9AC7F360AFFC299389" bold="true" box="[151,944,727,754]" pageId="13" pageNumber="542">
Notes on the status of
<taxonomicName id="C5B77DDA8972FF9AC6CD60A0FC299389" ID-CoL="4N78H" authority="Asahina, 1984" authorityName="Asahina" authorityYear="1984" box="[425,944,727,754]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC6CD60A0FD61938A" bold="true" box="[425,760,727,753]" italics="true" pageId="13" pageNumber="542">Protosticta khaosoidaoensis</emphasis>
<bibRefCitation id="66267BA88972FF9AC46460AFFC299389" author="Asahina" box="[768,944,728,754]" pageId="13" pageNumber="542" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina, 1984</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28972FF9AC7F3608DFED09268" box="[151,329,762,787]" pageId="13" pageNumber="542" type="description">
<paragraph id="020806598972FF9AC7F3608DFED09268" blockId="13.[151,944,727,787]" box="[151,329,762,787]" pageId="13" pageNumber="542">
(
<figureCitation id="9A8C1ADC8972FF9AC7FA608DFE939268" box="[158,266,762,787]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Figure 10</figureCitation>
A–E)
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28972FF9AC7F36134FA0592CF" pageId="13" pageNumber="542" type="reference_group">
<paragraph id="020806598972FF9AC7F36134FBBF92ED" blockId="13.[151,1436,834,949]" pageId="13" pageNumber="542">
<taxonomicName id="C5B77DDA8972FF9AC7F36134FDFD9223" ID-CoL="4N78H" authority="Asahina (1984)" authorityName="Asahina" authorityYear="1984" box="[151,612,834,856]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC7F36134FE2C9223" box="[151,437,834,856]" italics="true" pageId="13" pageNumber="542">Protosticta khaosoidaoensis</emphasis>
:
<bibRefCitation id="66267BA88972FF9AC6A06135FDFD9223" author="Asahina" box="[452,612,834,856]" pageId="13" pageNumber="542" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina (1984)</bibRefCitation>
</taxonomicName>
, pp. 588–590,
<figureCitation id="9A8C1ADC8972FF9AC4646135FCDC9223" box="[768,837,834,856]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A – G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H – J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="13" pageNumber="542">Figs. 8</figureCitation>
–15.
<typeStatus id="DD0CB8FB8972FF9AC41E6134FC359223" box="[890,940,835,856]" pageId="13" pageNumber="542">Type</typeStatus>
locality: Khao Soi Dao mountains,
<collectingCountry id="7AA046C98972FF9AC2466135FAE79223" box="[1314,1406,834,856]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
.—
<bibRefCitation id="66267BA88972FF9AC7A36116FD88920C" author="Hamalainen" box="[199,529,865,887]" pageId="13" pageNumber="542" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the dragonflies of Thailand distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana (1999)</bibRefCitation>
, p. 60 (Chiang Mai, Chantaburi).—van
<bibRefCitation id="66267BA88972FF9AC4C86116FB83920C" author="Tol" box="[940,1050,865,887]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
, pp. 220–221,
<figureCitation id="9A8C1ADC8972FF9AC3D06116FABB920C" box="[1204,1314,865,887]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStart-5="FIGURE 10" captionStartId-0="7.[151,250,1584,1606]" captionStartId-1="8.[151,250,1858,1880]" captionStartId-2="9.[151,250,1699,1721]" captionStartId-3="11.[151,250,1179,1201]" captionStartId-4="12.[151,250,1246,1268]" captionStartId-5="14.[151,250,1195,1217]" captionTargetBox-0="[150,1433,668,1559]" captionTargetBox-1="[155,1430,564,1821]" captionTargetBox-2="[151,1432,481,1664]" captionTargetBox-3="[157,1428,303,1152]" captionTargetBox-4="[155,1429,273,1221]" captionTargetBox-5="[150,1436,378,1170]" captionTargetId-0="figure@7.[150,1436,663,1562]" captionTargetId-1="figure@8.[151,1435,555,1831]" captionTargetId-2="figure@9.[151,1436,479,1677]" captionTargetId-3="figure@11.[151,1436,303,1156]" captionTargetId-4="figure@12.[151,1436,265,1225]" captionTargetId-5="figure@14.[150,1437,375,1173]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="14" captionText-0="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." captionText-1="FIGURE 6. Protosticta satoi (light form), Xuan Son. [A – B] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; [C – E] female— (C) head &amp; thorax in lateral view; (D) prothorax in dorsal view; (E) abdomen tip in lateral view." captionText-2="FIGURE 7. Protosticta socculus spec. nov. [A – H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." captionText-3="FIGURE 8. Protosticta spinosa spec. nov., [A – G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H – J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." captionText-4="FIGURE 9. Protosticta spp. [A – M] Protosticta pseudocuriosa spec. nov. [A – I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N – Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." captionText-5="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri-0="https://zenodo.org/record/257728/files/figure.png" httpUri-1="https://zenodo.org/record/257729/files/figure.png" httpUri-2="https://zenodo.org/record/257730/files/figure.png" httpUri-3="https://zenodo.org/record/257731/files/figure.png" httpUri-4="https://zenodo.org/record/257732/files/figure.png" httpUri-5="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Figs. 5–10</figureCitation>
.
<bibRefCitation id="66267BA88972FF9AC24A6116FEB292ED" pageId="13" pageNumber="542" refString="Makbun, N. et al. (2011) Additional records of Odonata from Doi Suthep-Pui National Park, Chiang Mai Province, Thailand. Notulae Odonatologicae, 7 (7), 61 - 68." type="journal article">
Makbun
<emphasis id="30C3DA4B8972FF9AC2EE6115FF7B92ED" italics="true" pageId="13" pageNumber="542">et al.</emphasis>
(2011)
</bibRefCitation>
, p. 63 (Chiang Mai).—
<bibRefCitation id="66267BA88972FF9AC57061F7FCCA92ED" author="Yokoi" box="[532,851,896,918]" pageId="13" pageNumber="542" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A list of Lao Dragonflies. Kyoei Printing Co. Ltd. Koriyama, 110 pp, 28 pls." type="book" year="2014">Yokoi &amp; Souphanthong (2014)</bibRefCitation>
, p. 26, Pl. 3, Fig. 14.
</paragraph>
<paragraph id="020806598972FF9AC7F361D7FA0592CF" blockId="13.[151,1436,834,949]" box="[151,1436,926,949]" pageId="13" pageNumber="542">
<taxonomicName id="C5B77DDA8972FF9AC7F361D7FC0892CF" ID-CoL="4N78H" authority="Kitagawa &amp; Katatani (2002)" authorityName="Kitagawa &amp; Katatani" authorityYear="2002" box="[151,913,926,949]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="subSpecies" species="khaosoidaoensis" subSpecies="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC7F361D7FDF892CF" box="[151,609,926,949]" italics="true" pageId="13" pageNumber="542">Protosticta khaosoidaoensis khaosoidaoensis</emphasis>
:
<bibRefCitation id="66267BA88972FF9AC50A61E9FC0892CF" author="Kitagawa" box="[622,913,926,948]" pageId="13" pageNumber="542" refString="Kitagawa, K. &amp; Katatani, N. (2002) Notes on Thai Odonata, part 1. The Odonata from Chantaburi in March, 1998. Aeschna, 39, 33 - 42." type="journal article" year="2002">Kitagawa &amp; Katatani (2002)</bibRefCitation>
</taxonomicName>
, pp. 33–34,
<figureCitation id="9A8C1ADC8972FF9AC36861E9FBDE92CF" box="[1036,1095,926,948]" captionStart="FIGURE 1" captionStartId="1.[151,250,1915,1937]" captionTargetBox="[157,1432,706,1896]" captionTargetId="figure@1.[151,1435,701,1909]" captionTargetPageId="1" captionText="FIGURE 1. Protosticta beaumonti, Mau Son. [A – D] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus &amp; paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E – F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." httpUri="https://zenodo.org/record/257724/files/figure.png" pageId="13" pageNumber="542">Fig. 1</figureCitation>
(Pong Nam Ron, Khao Soi Dao).
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28972FF9AC7F36196FBC695D5" pageId="13" pageNumber="542" type="materials_examined">
<paragraph id="020806598972FF9AC7F36196FAC2953A" blockId="13.[151,1436,993,1090]" pageId="13" pageNumber="542">
<emphasis id="30C3DA4B8972FF9AC7F36196FACD9539" italics="true" pageId="13" pageNumber="542">
The material identified in van
<bibRefCitation id="66267BA88972FF9AC68A6196FDFA9281" author="Tol" box="[494,611,993,1018]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
as this species, but no longer considered to belong to it:
<specimenCount id="14B1CDD08972FF9AC38F6195FADD9281" box="[1259,1348,993,1018]" pageId="13" pageNumber="542" type="male">2 males</specimenCount>
, Huong Son district, Ha Tinh Province,
<date id="760920998972FF9AC69D6671FDEB9565" box="[505,626,1030,1054]" pageId="13" pageNumber="542" value="1990-07-02">2.VII.1990</date>
;
<specimenCount id="14B1CDD08972FF9AC5E56671FD569565" box="[641,719,1029,1054]" pageId="13" pageNumber="542" type="male">1 male</specimenCount>
, Ky Son district, Nghe An Province,
<date id="760920998972FF9AC30D6671FB7B9565" box="[1129,1250,1030,1054]" pageId="13" pageNumber="542" value="1990-07-08">8.VII.1990</date>
;
<specimenCount id="14B1CDD08972FF9AC3956671FAD09565" box="[1265,1353,1029,1054]" pageId="13" pageNumber="542" type="male">2 males</specimenCount>
, Thanh Chuong district, Nghe An Province,
<date id="760920998972FF9AC54A665DFD409539" box="[558,729,1066,1090]" pageId="13" pageNumber="542" value="1990-07-11" valueMax="1990-07-12" valueMin="1990-07-11">11-12.VII.1990</date>
;
<specimenCount id="14B1CDD08972FF9AC583665DFCAC9539" box="[743,821,1065,1090]" pageId="13" pageNumber="542" type="male">1 male</specimenCount>
, Doi Suthep, Chiang Mai,
<collectingCountry id="7AA046C98972FF9AC305665EFB5F9539" box="[1121,1222,1065,1090]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
(JvT 19422)
</emphasis>
.
</paragraph>
<paragraph id="020806598972FF9AC7A36606FBC695D5" blockId="13.[151,1437,1137,2026]" pageId="13" pageNumber="542">
<emphasis id="30C3DA4B8972FF9AC7A36606FE5D95F1" bold="true" box="[199,452,1137,1162]" pageId="13" pageNumber="542">Specimens examined</emphasis>
.
<specimenCount id="14B1CDD08972FF9AC6B36605FDB395F1" box="[471,554,1137,1162]" pageId="13" pageNumber="542" type="male">1 male</specimenCount>
, Krating waterfall, Chantaburi Province,
<collectingCountry id="7AA046C98972FF9AC3766606FBE195F1" box="[1042,1144,1137,1162]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
,
<date id="760920998972FF9AC3EC6605FA8D95F1" box="[1160,1300,1138,1162]" pageId="13" pageNumber="542" value="2011-07-10">10.VII.2011</date>
;
<specimenCount id="14B1CDD08972FF9AC2406605FA0195F1" box="[1316,1432,1137,1162]" pageId="13" pageNumber="542" type="female">2 females</specimenCount>
, Krating waterfall, Chantaburi Province,
<collectingCountry id="7AA046C98972FF9AC53E66E2FD5995D5" box="[602,704,1173,1198]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
,
<date id="760920998972FF9AC5AF66E2FCD395D5" box="[715,842,1173,1198]" pageId="13" pageNumber="542" value="2015-04-03">03.IV.2015</date>
, Noppadon Makbun leg.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28972FF99C7A366CEFDF79183" lastPageId="14" lastPageNumber="543" pageId="13" pageNumber="542" type="discussion">
<paragraph id="020806598972FF9AC7A366CEFCEE94D1" blockId="13.[151,1437,1137,2026]" pageId="13" pageNumber="542">
<emphasis id="30C3DA4B8972FF9AC7A366CEFE9395A9" bold="true" box="[199,266,1209,1234]" pageId="13" pageNumber="542">Notes</emphasis>
. Van
<bibRefCitation id="66267BA88972FF9AC63666CEFE5595A9" author="Tol" box="[338,460,1209,1234]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
recorded
<taxonomicName id="C5B77DDA8972FF9AC52066CDFC8595A9" box="[580,796,1209,1234]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC52066CDFC8595A9" box="[580,796,1209,1234]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
from Huong Son district, Ha Tinh Province of central
<collectingCountry id="7AA046C98972FF9AC7F366AAFF6F958D" box="[151,246,1245,1270]" name="Vietnam" pageId="13" pageNumber="542">Vietnam</collectingCountry>
based on a comparison with a male of
<taxonomicName id="C5B77DDA8972FF9AC5A866A9FC3C958D" box="[716,933,1245,1270]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC5A866A9FC3C958D" box="[716,933,1245,1270]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
from Doi Suthep in Chiang Mai Province,
<collectingCountry id="7AA046C98972FF9AC7F36776FF679461" box="[151,254,1281,1306]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
. Although he recognized that the Vietnamese specimen resembled
<emphasis id="30C3DA4B8972FF9AC37B6775FB089462" box="[1055,1169,1282,1305]" italics="true" pageId="13" pageNumber="542">P. c ur i o s a</emphasis>
, he identified it as
<taxonomicName id="C5B77DDA8972FF9AC2E36775FEC99445" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC2E36775FEC99445" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
based on the similarity of the color pattern of the prothorax to the original description (
<bibRefCitation id="66267BA88972FF9AC25A6752FF4A9419" author="Asahina" pageId="13" pageNumber="542" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina 1984</bibRefCitation>
). To ascertain the status of
<taxonomicName id="C5B77DDA8972FF9AC565673DFD4C9419" box="[513,725,1353,1378]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC565673DFD4C9419" box="[513,725,1353,1378]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
in
<collectingCountry id="7AA046C98972FF9AC59D673EFCC29419" box="[761,859,1353,1378]" name="Vietnam" pageId="13" pageNumber="542">Vietnam</collectingCountry>
, we examined the
<typeStatus id="DD0CB8FB8972FF9AC34D673DFBFD9419" box="[1065,1124,1354,1378]" pageId="13" pageNumber="542">types</typeStatus>
and other specimens of this species from its
<typeStatus id="DD0CB8FB8972FF9AC6066719FE0B94FD" box="[354,402,1390,1414]" pageId="13" pageNumber="542">type</typeStatus>
locality (Chantaburi,
<collectingCountry id="7AA046C98972FF9AC5F8671AFC9894FD" box="[668,769,1389,1414]" name="Thailand" pageId="13" pageNumber="542">Thailand</collectingCountry>
) and compared these with the Vietnamese and
<collectingCountry id="7AA046C98972FF9AC223671AFAE394FD" box="[1351,1402,1389,1414]" name="Thailand" pageId="13" pageNumber="542">Thai</collectingCountry>
<taxonomicName id="C5B77DDA8972FF9AC2E36719FEC994D1" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC2E36719FEC994D1" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
illustrated by van
<bibRefCitation id="66267BA88972FF9AC54467E6FCEA94D1" author="Tol" box="[544,883,1425,1450]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008" yearSuffix="p">
Tol (2008: p. 221,
<figureCitation id="9A8C1ADC8972FF9AC59467E6FCF394D1" box="[752,874,1425,1450]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStart-5="FIGURE 10" captionStartId-0="7.[151,250,1584,1606]" captionStartId-1="8.[151,250,1858,1880]" captionStartId-2="9.[151,250,1699,1721]" captionStartId-3="11.[151,250,1179,1201]" captionStartId-4="12.[151,250,1246,1268]" captionStartId-5="14.[151,250,1195,1217]" captionTargetBox-0="[150,1433,668,1559]" captionTargetBox-1="[155,1430,564,1821]" captionTargetBox-2="[151,1432,481,1664]" captionTargetBox-3="[157,1428,303,1152]" captionTargetBox-4="[155,1429,273,1221]" captionTargetBox-5="[150,1436,378,1170]" captionTargetId-0="figure@7.[150,1436,663,1562]" captionTargetId-1="figure@8.[151,1435,555,1831]" captionTargetId-2="figure@9.[151,1436,479,1677]" captionTargetId-3="figure@11.[151,1436,303,1156]" captionTargetId-4="figure@12.[151,1436,265,1225]" captionTargetId-5="figure@14.[150,1437,375,1173]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="14" captionText-0="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." captionText-1="FIGURE 6. Protosticta satoi (light form), Xuan Son. [A – B] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; [C – E] female— (C) head &amp; thorax in lateral view; (D) prothorax in dorsal view; (E) abdomen tip in lateral view." captionText-2="FIGURE 7. Protosticta socculus spec. nov. [A – H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." captionText-3="FIGURE 8. Protosticta spinosa spec. nov., [A – G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H – J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." captionText-4="FIGURE 9. Protosticta spp. [A – M] Protosticta pseudocuriosa spec. nov. [A – I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N – Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." captionText-5="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri-0="https://zenodo.org/record/257728/files/figure.png" httpUri-1="https://zenodo.org/record/257729/files/figure.png" httpUri-2="https://zenodo.org/record/257730/files/figure.png" httpUri-3="https://zenodo.org/record/257731/files/figure.png" httpUri-4="https://zenodo.org/record/257732/files/figure.png" httpUri-5="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Figs. 5–10</figureCitation>
)
</bibRefCitation>
.
</paragraph>
<paragraph id="020806598972FF9AC7A367C2FECE97DE" blockId="13.[151,1437,1137,2026]" pageId="13" pageNumber="542">
Surprisingly, the male &quot;
<taxonomicName id="C5B77DDA8972FF9AC6AB67C1FD3A94B5" box="[463,675,1461,1486]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC6AB67C1FD3A94B5" box="[463,675,1461,1486]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
&quot; from Doi Suthep illustrated by van
<bibRefCitation id="66267BA88972FF9AC32667C2FA1B94B5" author="Tol" box="[1090,1410,1461,1486]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008" yearSuffix="p">
Tol (2008: p. 221,
<figureCitation id="9A8C1ADC8972FF9AC27567C2FAE394B5" box="[1297,1402,1461,1486]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,250,1584,1606]" captionStartId-1="8.[151,250,1858,1880]" captionStartId-2="9.[151,250,1699,1721]" captionTargetBox-0="[150,1433,668,1559]" captionTargetBox-1="[155,1430,564,1821]" captionTargetBox-2="[151,1432,481,1664]" captionTargetId-0="figure@7.[150,1436,663,1562]" captionTargetId-1="figure@8.[151,1435,555,1831]" captionTargetId-2="figure@9.[151,1436,479,1677]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." captionText-1="FIGURE 6. Protosticta satoi (light form), Xuan Son. [A – B] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; [C – E] female— (C) head &amp; thorax in lateral view; (D) prothorax in dorsal view; (E) abdomen tip in lateral view." captionText-2="FIGURE 7. Protosticta socculus spec. nov. [A – H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri-0="https://zenodo.org/record/257728/files/figure.png" httpUri-1="https://zenodo.org/record/257729/files/figure.png" httpUri-2="https://zenodo.org/record/257730/files/figure.png" pageId="13" pageNumber="542">Figs. 5–7</figureCitation>
)
</bibRefCitation>
is different from
<taxonomicName id="C5B77DDA8972FF9AC65B67ADFD8F9489" box="[319,534,1497,1522]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC65B67ADFD8F9489" box="[319,534,1497,1522]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
in several characters: (1) in
<taxonomicName id="C5B77DDA8972FF9AC40567ADFBA19489" box="[865,1080,1497,1522]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC40567ADFBA19489" box="[865,1080,1497,1522]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
, the paraprocts have elongated tips, extending as far as the cerci (
<figureCitation id="9A8C1ADC8972FF9AC54C678AFDE2976D" box="[552,635,1533,1558]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Fig. 10</figureCitation>
B and Fig. 13, p. 589,
<bibRefCitation id="66267BA88972FF9AC4E3678AFBB1976D" author="Asahina" box="[903,1064,1533,1558]" pageId="13" pageNumber="542" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina 1984</bibRefCitation>
), but are very short in van Tol's specimen (2008: p. 221,
<figureCitation id="9A8C1ADC8972FF9AC6CA6456FE6B9741" box="[430,498,1569,1594]" captionStart="FIGURE 6" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[155,1430,564,1821]" captionTargetId="figure@8.[151,1435,555,1831]" captionTargetPageId="8" captionText="FIGURE 6. Protosticta satoi (light form), Xuan Son. [A – B] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; [C – E] female— (C) head &amp; thorax in lateral view; (D) prothorax in dorsal view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257729/files/figure.png" pageId="13" pageNumber="542">Fig. 6</figureCitation>
); (2) in
<taxonomicName id="C5B77DDA8972FF9AC52A6455FCBD9741" box="[590,804,1569,1594]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC52A6455FCBD9741" box="[590,804,1569,1594]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
the paraprocts have at their dorsal apex a large cup-like structure with a small spine (
<figureCitation id="9A8C1ADC8972FF9AC6B96432FDB69725" box="[477,559,1605,1630]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Fig. 10</figureCitation>
B) that is absent in van Tol's male (van
<bibRefCitation id="66267BA88972FF9AC4896432FBCC9725" author="Tol" box="[1005,1109,1605,1630]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol 2008</bibRefCitation>
: p. 221,
<figureCitation id="9A8C1ADC8972FF9AC3D26432FB639725" box="[1206,1274,1605,1630]" captionStart="FIGURE 7" captionStartId="9.[151,250,1699,1721]" captionTargetBox="[151,1432,481,1664]" captionTargetId="figure@9.[151,1436,479,1677]" captionTargetPageId="9" captionText="FIGURE 7. Protosticta socculus spec. nov. [A – H] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – K] paratype male— (J) anal appendages in lateral view; (K) right cercus and paraproct in oblique-dorsal view." httpUri="https://zenodo.org/record/257730/files/figure.png" pageId="13" pageNumber="542">Fig. 7</figureCitation>
). Therefore, it appears that the male
<taxonomicName id="C5B77DDA8972FF9AC6EB641DFDFD97F9" box="[399,612,1641,1666]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC6EB641DFDFD97F9" box="[399,612,1641,1666]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
from Doi Suthep illustrated by van
<bibRefCitation id="66267BA88972FF9AC49B641EFBEF97F9" author="Tol" box="[1023,1142,1641,1666]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
does not represent true
<taxonomicName id="C5B77DDA8972FF9AC2E3641DFEC997DD" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC2E3641DFEC997DD" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="020806598972FF9AC7A364C5FC569605" blockId="13.[151,1437,1137,2026]" pageId="13" pageNumber="542">
Van
<bibRefCitation id="66267BA88972FF9AC79B64C6FEE597B1" author="Tol" box="[255,380,1713,1738]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
also questioned whether
<emphasis id="30C3DA4B8972FF9AC5D464C5FCBB97B2" box="[688,802,1714,1737]" italics="true" pageId="13" pageNumber="542">P. c u r i o s a</emphasis>
and
<taxonomicName id="C5B77DDA8972FF9AC40C64C5FBDB97B1" box="[872,1090,1713,1738]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC40C64C5FBDB97B1" box="[872,1090,1713,1738]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
are truly distinct species, as structural differences seemed to be small. In fact there are some clear differences. Whereas
<taxonomicName id="C5B77DDA8972FF9AC3FF64A1FAF69795" box="[1179,1391,1749,1774]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC3FF64A1FAF69795" box="[1179,1391,1749,1774]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
has a large cup-like structure with a large spine on its distal margin at the dorsal apex of the paraprocts (
<figureCitation id="9A8C1ADC8972FF9AC27C648EFAC59669" box="[1304,1372,1785,1810]" captionStart="FIGURE 5" captionStartId="7.[151,250,1584,1606]" captionTargetBox="[150,1433,668,1559]" captionTargetId="figure@7.[150,1436,663,1562]" captionTargetPageId="7" captionText="FIGURE 5. Protosticta satoi (dark form), Cao Bang. [A – G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] male— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view; male, Tam Dao— (K) prothorax in dorsal view." httpUri="https://zenodo.org/record/257728/files/figure.png" pageId="13" pageNumber="542">Fig. 5</figureCitation>
B),
<taxonomicName id="C5B77DDA8972FF9AC2E3648DFF72964E" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="30C3DA4B8972FF9AC2E3648DFF72964E" italics="true" pageId="13" pageNumber="542">P. curiosa</emphasis>
</taxonomicName>
has only a very small cup-like structure lacking a spine (
<figureCitation id="9A8C1ADC8972FF9AC403656AFC33964D" box="[871,938,1821,1846]" captionStart="FIGURE 9" captionStartId="12.[151,250,1246,1268]" captionTargetBox="[155,1429,273,1221]" captionTargetId="figure@12.[151,1436,265,1225]" captionTargetPageId="12" captionText="FIGURE 9. Protosticta spp. [A – M] Protosticta pseudocuriosa spec. nov. [A – I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N – Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257732/files/figure.png" pageId="13" pageNumber="542">Fig. 9</figureCitation>
N). The female of
<emphasis id="30C3DA4B8972FF9AC3136569FB7D964E" box="[1143,1252,1822,1845]" italics="true" pageId="13" pageNumber="542">P. c ur i os a</emphasis>
differs from the female of
<taxonomicName id="C5B77DDA8972FF9AC66F6535FE789621" box="[267,481,1857,1882]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC66F6535FE789621" box="[267,481,1857,1882]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
by a large oval yellow spot on the dorsum of S9 (
<figureCitation id="9A8C1ADC8972FF9AC37B6536FBFA9621" box="[1055,1123,1857,1882]" captionStart="FIGURE 9" captionStartId="12.[151,250,1246,1268]" captionTargetBox="[155,1429,273,1221]" captionTargetId="figure@12.[151,1436,265,1225]" captionTargetPageId="12" captionText="FIGURE 9. Protosticta spp. [A – M] Protosticta pseudocuriosa spec. nov. [A – I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N – Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257732/files/figure.png" pageId="13" pageNumber="542">Fig. 9</figureCitation>
Q), whereas this segment is generally, but not always, entirely black in
<taxonomicName id="C5B77DDA8972FF9AC51D6511FCD79605" box="[633,846,1893,1918]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC51D6511FCD79605" box="[633,846,1893,1918]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
(
<figureCitation id="9A8C1ADC8972FF9AC4396512FC379605" box="[861,942,1893,1918]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="13" pageNumber="542">Fig. 10</figureCitation>
E).
</paragraph>
<paragraph id="020806598972FF9AC7A365FEFA049692" blockId="13.[151,1437,1137,2026]" pageId="13" pageNumber="542">
The Vietnamese material identified by van
<bibRefCitation id="66267BA88972FF9AC5DB65FEFCA096D9" author="Tol" box="[703,825,1929,1954]" pageId="13" pageNumber="542" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
as
<emphasis id="30C3DA4B8972FF9AC40065FDFBCD96D9" box="[868,1108,1929,1954]" italics="true" pageId="13" pageNumber="542">
&quot;
<taxonomicName id="C5B77DDA8972FF9AC40965FDFBD296D9" box="[877,1099,1929,1954]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">P. khaosoidaoensis</taxonomicName>
&quot;
</emphasis>
is clearly much closer to
<taxonomicName id="C5B77DDA8972FF9AC2E365FDFF7296BE" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="30C3DA4B8972FF9AC2E365FDFF7296BE" italics="true" pageId="13" pageNumber="542">P. curiosa</emphasis>
</taxonomicName>
than to
<taxonomicName id="C5B77DDA8972FF9AC62265D9FD8296BD" box="[326,539,1965,1990]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC62265D9FD8296BD" box="[326,539,1965,1990]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
. Therefore we reject his specimen as
<taxonomicName id="C5B77DDA8972FF9AC4A665D9FB0F96BD" box="[962,1174,1965,1990]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8972FF9AC4A665D9FB0F96BD" box="[962,1174,1965,1990]" italics="true" pageId="13" pageNumber="542">P. khaosoidaoensis</emphasis>
</taxonomicName>
and cannot support the occurrence of that species in
<collectingCountry id="7AA046C98972FF9AC68665A6FDDC9691" box="[482,581,2001,2026]" name="Vietnam" pageId="13" pageNumber="542">Vietnam</collectingCountry>
. In fact his material matches our description of
<taxonomicName id="C5B77DDA8972FF9AC30065A5FABD9691" box="[1124,1316,2001,2026]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="542" phylum="Arthropoda" rank="species" species="pseudocuriosa" status="sp. nov.">
<emphasis id="30C3DA4B8972FF9AC30065A5FABD9691" box="[1124,1316,2001,2026]" italics="true" pageId="13" pageNumber="542">P. pseudocuriosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2BF067308972FF9AC24865A5FA049692" box="[1324,1437,2002,2025]" pageId="13" pageNumber="542" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="020806598971FF99C7F362E0FDF79183" blockId="14.[151,1437,151,320]" pageId="14" pageNumber="543">
from the same locality, especially by having the same two small oval yellow spots at the posterior end of the mesepisternum and mesepimeron (van
<bibRefCitation id="66267BA88971FF99C53D62CBFD5D91AF" author="Tol" box="[601,708,188,213]" pageId="14" pageNumber="543" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol 2008</bibRefCitation>
: p. 221,
<figureCitation id="9A8C1ADC8971FF99C44862CBFCEB91AF" box="[812,882,188,213]" captionStart="FIGURE 8" captionStartId="11.[151,250,1179,1201]" captionTargetBox="[157,1428,303,1152]" captionTargetId="figure@11.[151,1436,303,1156]" captionTargetPageId="11" captionText="FIGURE 8. Protosticta spinosa spec. nov., [A – G] holotype male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) paraprocts in ventral view; (F) penile organ in dorsal view; (G) wings; [H – J] holotype female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257731/files/figure.png" pageId="14" pageNumber="543">Fig. 8</figureCitation>
). This character is absent in
<taxonomicName id="C5B77DDA8971FF99C3A062CAFA0591AE" box="[1220,1436,188,213]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="543" phylum="Arthropoda" rank="species" species="khaosoidaoensis">
<emphasis id="30C3DA4B8971FF99C3A062CAFA0591AE" box="[1220,1436,188,213]" italics="true" pageId="14" pageNumber="543">P. khaosoidaoensis</emphasis>
</taxonomicName>
(
<figureCitation id="9A8C1ADC8971FF99C7FB62A8FF639183" box="[159,250,223,248]" captionStart="FIGURE 10" captionStartId="14.[151,250,1195,1217]" captionTargetBox="[150,1436,378,1170]" captionTargetId="figure@14.[150,1437,375,1173]" captionTargetPageId="14" captionText="FIGURE 10. Protosticta khaosoidaoensis, Chantaburi, [A – C] male— (A) head &amp; thorax in lateral view; (B) anal appendages in oblique-lateral view; (C) tip of cerci in posteroventral view; [D – E] female— (D) head &amp; thorax in lateral view; (E) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257733/files/figure.png" pageId="14" pageNumber="543">Figs. 10</figureCitation>
A, 10D) and
<emphasis id="30C3DA4B8971FF99C6E86296FE609183" box="[396,505,225,248]" italics="true" pageId="14" pageNumber="543">P. c u r i o s a</emphasis>
(
<figureCitation id="9A8C1ADC8971FF99C56F62A8FDD69183" box="[523,591,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,1246,1268]" captionTargetBox="[155,1429,273,1221]" captionTargetId="figure@12.[151,1436,265,1225]" captionTargetPageId="12" captionText="FIGURE 9. Protosticta spp. [A – M] Protosticta pseudocuriosa spec. nov. [A – I] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view and (E) oblique-dorsal view; (F) tip of cerci in posteroventral view; (G) paraprocts in ventral view; (H) penile organ in dorsal view; (I) wings; [J – M] female— (J) head &amp; thorax in lateral view; (K) prothorax in dorsal view; (M) abdomen tip in lateral view; (L) male, Son Kim, prothorax in dorsal view; [N – Q] Protosticta curiosa, Thailand, (later Asahina's col.) male— (N) anal appendages in oblique-dorsal view; (O) tip of cerci in posteroventral view; (P) female, head &amp; thorax in lateral view; (Q) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257732/files/figure.png" pageId="14" pageNumber="543">Fig. 9</figureCitation>
P).
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D28971FF99C7A36373FE95903B" pageId="14" pageNumber="543" type="distribution">
<paragraph id="020806598971FF99C7A36373FE95903B" blockId="14.[151,1437,151,320]" pageId="14" pageNumber="543">
<emphasis id="30C3DA4B8971FF99C7A36373FEC09066" bold="true" box="[199,345,260,285]" pageId="14" pageNumber="543">Distribution</emphasis>
.
<collectingCountry id="7AA046C98971FF99C6086373FE499066" box="[364,464,260,285]" name="Thailand" pageId="14" pageNumber="543">Thailand</collectingCountry>
(Chantaburi and Chiang Mai Provinces), Lao PDR (Louang Phabang, Oudomxay Province).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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