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<mods:title id="F63A141877282FF38D5AC5067D202F26">Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species</mods:title>
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<heading id="990081C3FFB09A14FF19B56169EEB57B" bold="true" box="[151,484,570,596]" fontSize="11" level="1" pageId="1" pageNumber="60" reason="1">
<taxonomicName id="05F74D2CFFB09A14FF19B56169EEB57B" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,484,570,596]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FF19B56169EEB57B" bold="true" box="[151,484,570,596]" pageId="1" pageNumber="60">
<emphasis id="F083EABDFFB09A14FF19B561690EB57B" bold="true" box="[151,260,570,596]" italics="true" pageId="1" pageNumber="60">Wahydra</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FE85B56169EEB57B" author="Steinhauser" box="[267,484,570,596]" pageId="1" pageNumber="60" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="8AED6524FFB09A14FF19B5DA6D46B43D" pageId="1" pageNumber="60" type="reference_group">
<paragraph id="C24836AFFFB09A14FF19B5DA6D7EB5DC" blockId="1.[151,1436,641,786]" pageId="1" pageNumber="60">
<taxonomicName id="05F74D2CFFB09A14FF19B5DA69B8B5B8" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,434,641,663]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FF19B5DA68F8B5B8" box="[151,242,641,663]" italics="true" pageId="1" pageNumber="60">Wahydra</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FF77B5DA69B8B5B8" author="Steinhauser" box="[249,434,641,663]" pageId="1" pageNumber="60" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</taxonomicName>
.
<emphasis id="F083EABDFFB09A14FE33B5DA6AABB5B8" bold="true" box="[445,673,641,663]" pageId="1" pageNumber="60">Bull. Allyn Mus. 132</emphasis>
: 30; type-species:
<taxonomicName id="05F74D2CFFB09A14FCEAB5DA6C94B5B8" authority="Butler, 1870" authorityName="Butler" authorityYear="1870" box="[868,1182,641,663]" class="Insecta" family="Hesperiidae" genus="Pamphila" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="kenava">
<emphasis id="F083EABDFFB09A14FCEAB5DA6C1CB5B8" box="[868,1046,641,663]" italics="true" pageId="1" pageNumber="60">Pamphila kenava</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FB91B5DA6C94B5B8" author="Butler" box="[1055,1182,641,663]" pageId="1" pageNumber="60" refString="Butler, A. (1870) Descriptions of some new diurnal Lepidoptera, chiefly Hesperiidae. Transactions of the entomological Society of London, (4), 485 - 520. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01881. x" type="journal article" year="1870">Butler, 1870</bibRefCitation>
</taxonomicName>
.—
<bibRefCitation id="A6664B5EFFB09A14FB33B5DA6D4CB5B8" author="Lamas" box="[1213,1350,641,663]" pageId="1" pageNumber="60" refString="Lamas, G. (2003) Las Mariposas de Machu Picchu. Guia ilustrada de las mariposas del Santuario Historico Machu Picchu, Cuzco, Peru. Lima, Profonanpe, 221 pp." type="book" year="2003">Lamas, 2003</bibRefCitation>
.
<emphasis id="F083EABDFFB09A14FADFB5DA6960B599" bold="true" pageId="1" pageNumber="60">Marip. Machu Picchu</emphasis>
, p. 30.—
<bibRefCitation id="A6664B5EFFB09A14FE58B5FB6A63B599" author="Mielke" box="[470,617,672,694]" pageId="1" pageNumber="60" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, pp. 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 84,
<emphasis id="F083EABDFFB09A14FCECB5FB6B7DB59A" bold="true" box="[866,887,672,693]" italics="true" pageId="1" pageNumber="60">in</emphasis>
Lamas (ed.).
<emphasis id="F083EABDFFB09A14FB97B5FB6951B5FB" bold="true" pageId="1" pageNumber="60">Checklist: Part 4A, Hesperioidea- Papilionoidea</emphasis>
,
<emphasis id="F083EABDFFB09A14FEE2B5E4698BB5FB" bold="true" box="[364,385,703,724]" italics="true" pageId="1" pageNumber="60">in</emphasis>
Heppner (ed.).
<emphasis id="F083EABDFFB09A14FDA3B5E56B3EB5FB" bold="true" box="[557,820,702,724]" pageId="1" pageNumber="60">
<taxonomicName id="05F74D2CFFB09A14FDA3B5E56B01B5FB" authority="Neotrop. Lep." authorityName="Neotrop. Lep." box="[557,779,702,724]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—
<bibRefCitation id="A6664B5EFFB09A14FCDCB5E56BE8B5FB" author="Mielke" box="[850,994,702,724]" pageId="1" pageNumber="60" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 5: Hesperiinae 2. In: Megaleas - Zenis. Sociedade Brasileira de Zoologia, Curitiba, pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="F083EABDFFB09A14FC7EB5E56D10B5FB" bold="true" box="[1008,1306,702,724]" pageId="1" pageNumber="60">Cat. Amer. Hesperioidea 5</emphasis>
, p. 1362.—
<bibRefCitation id="A6664B5EFFB09A14FF49B5856A38B5DC" author="Warren" box="[199,562,733,755]" pageId="1" pageNumber="60" refString="Warren, A. D., Ogawa, J. R. &amp; Brower, A. V. Z. (2008) Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics, 24, 642 - 676. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2008.00218. x" type="journal article" year="2008">Warren; Ogawa &amp; A. Brower, 2008</bibRefCitation>
.
<emphasis id="F083EABDFFB09A14FDB2B5866ACFB5DC" bold="true" box="[572,709,733,755]" pageId="1" pageNumber="60">Cladistics 24</emphasis>
: 27.—
<bibRefCitation id="A6664B5EFFB09A14FC87B5866C7DB5DC" author="Warren" box="[777,1143,733,755]" pageId="1" pageNumber="60" refString="Warren, A. D., Ogawa, J. R. &amp; Brower, A. V. Z. (2009) Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic Entomology, 34, 467 - 523. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2008.00463. x" type="journal article" year="2009">Warren; Ogawa &amp; A. Brower, 2009</bibRefCitation>
.
<emphasis id="F083EABDFFB09A14FB0EB5866D0DB5DC" bold="true" box="[1152,1287,733,755]" pageId="1" pageNumber="60">Syst. Ent. 34</emphasis>
: 498, 516.
</paragraph>
<paragraph id="C24836AFFFB09A14FF19B5A76D46B43D" blockId="1.[151,1436,641,786]" box="[151,1356,764,786]" pageId="1" pageNumber="60">
<emphasis id="F083EABDFFB09A14FF19B5A76904B43E" box="[151,270,764,786]" italics="true" pageId="1" pageNumber="60">
<taxonomicName id="05F74D2CFFB09A14FF19B5A768F8B43D" box="[151,242,764,786]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">Wahydra</taxonomicName>
<emphasis id="F083EABDFFB09A14FF76B5A76904B43E" bold="true" box="[248,270,764,785]" italics="true" pageId="1" pageNumber="60">sp</emphasis>
</emphasis>
. Orellana, 2003,
<emphasis id="F083EABDFFB09A14FE4AB5A769D3B43E" bold="true" box="[452,473,764,785]" italics="true" pageId="1" pageNumber="60">in</emphasis>
Andressen &amp; Monasterio.
<emphasis id="F083EABDFFB09A14FD65B5A76C91B43D" bold="true" box="[747,1179,764,786]" pageId="1" pageNumber="60">Mem. IV Simp. Intern. Des. Sust. Andes</emphasis>
, Mérida, p. 5-61.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB09A14FF19B46268E0B361" pageId="1" pageNumber="60" type="diagnosis">
<paragraph id="C24836AFFFB09A14FF19B46268E0B361" blockId="1.[151,1437,825,1935]" pageId="1" pageNumber="60">
<emphasis id="F083EABDFFB09A14FF19B4626957B47D" bold="true" box="[151,349,825,850]" pageId="1" pageNumber="60">Diagnosis. Male:</emphasis>
Genus traditionally recognized by the presence of orange bands on both DFW and DHW, though some species have those bands reduced.
<taxonomicName id="05F74D2CFFB09A14FDFEB4066B6DB459" authority="Bell, 1942" authorityName="Bell" authorityYear="1942" box="[624,871,861,886]" class="Insecta" family="Hesperiidae" genus="Anthoptus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FDFEB4066AEBB45A" box="[624,737,861,885]" italics="true" pageId="1" pageNumber="60">Anthoptus</emphasis>
Bell, 1942
</taxonomicName>
,
<taxonomicName id="05F74D2CFFB09A14FCFAB4056C7BB459" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[884,1137,861,886]" class="Insecta" family="Hesperiidae" genus="Corticea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FCFAB4056BDCB45A" box="[884,982,862,885]" italics="true" pageId="1" pageNumber="60">Corticea</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FC6FB4056C7BB459" author="Evans" box="[993,1137,861,886]" pageId="1" pageNumber="60" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="05F74D2CFFB09A14FBF1B4066D63B459" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[1151,1385,861,886]" class="Insecta" family="Hesperiidae" genus="Cantha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FBF1B4066CD8B45A" box="[1151,1234,861,885]" italics="true" pageId="1" pageNumber="60">Cantha</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FB52B4056D63B459" author="Evans" box="[1244,1385,861,886]" pageId="1" pageNumber="60" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="05F74D2CFFB09A14FF19B4DA6971B4B5" authority="Bell, 1947" authorityName="Bell" authorityYear="1947" box="[151,379,897,922]" class="Insecta" family="Hesperiidae" genus="Zalomes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FF19B4DA68FCB4B6" box="[151,246,897,921]" italics="true" pageId="1" pageNumber="60">Zalomes</emphasis>
Bell, 1947
</taxonomicName>
(genera of
<taxonomicName id="05F74D2CFFB09A14FD87B4DA6A82B4B5" box="[521,648,897,922]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="tribe" tribe="Anthoptini">Anthoptini</taxonomicName>
);
<taxonomicName id="05F74D2CFFB09A14FD11B4D96B76B4B5" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[671,892,897,922]" class="Insecta" family="Hesperiidae" genus="Lento" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FD11B4D96AD5B4B6" box="[671,735,898,921]" italics="true" pageId="1" pageNumber="60">Lento</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FD64B4D96B76B4B5" author="Evans" box="[746,892,897,922]" pageId="1" pageNumber="60" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="05F74D2CFFB09A14FC02B4D96C81B4B5" authority="Godman, 1900" authorityName="Godman" authorityYear="1900" box="[908,1163,897,922]" class="Insecta" family="Hesperiidae" genus="Vinius" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FC02B4D96BDBB4B6" box="[908,977,898,921]" italics="true" pageId="1" pageNumber="60">Vinius</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FC53B4DA6C81B4B5" author="Godman" box="[989,1163,897,922]" pageId="1" pageNumber="60" refString="Godman F. D. (1900) Notes on the American species of Hesperiidae described by Plotz. Annals and Magazine of natural History, (7) 20 (116), 132 - 155." type="journal article" year="1900">Godman, 1900</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="05F74D2CFFB09A14FB15B4D96D96B4B5" authority="Austin, 1997" authorityName="Austin" authorityYear="1997" box="[1179,1436,897,922]" class="Insecta" family="Hesperiidae" genus="Vinpeius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FB15B4D96CF3B4B6" box="[1179,1273,898,921]" italics="true" pageId="1" pageNumber="60">Vinpeius</emphasis>
Austin, 1997
</taxonomicName>
(genera of
<taxonomicName id="05F74D2CFFB09A14FE9FB4FE697EB491" box="[273,372,933,958]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="tribe" tribe="Moncini">Moncini</taxonomicName>
),
<taxonomicName id="05F74D2CFFB09A14FE06B4FE6A7EB491" authority="Godman, 1900" authorityName="Godman" authorityYear="1900" box="[392,628,933,958]" class="Insecta" family="Hesperiidae" genus="Molo" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FE06B4FE69CEB492" box="[392,452,933,957]" italics="true" pageId="1" pageNumber="60">Molo</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FE45B4FE6A7EB491" author="Godman" box="[459,628,933,958]" pageId="1" pageNumber="60" refString="Godman F. D. (1900) Notes on the American species of Hesperiidae described by Plotz. Annals and Magazine of natural History, (7) 20 (116), 132 - 155." type="journal article" year="1900">Godman, 1900</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="05F74D2CFFB09A14FD0FB4FD6B6BB491" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[641,865,933,958]" class="Insecta" family="Hesperiidae" genus="Misius" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FD0FB4FD6AC6B492" box="[641,716,934,957]" italics="true" pageId="1" pageNumber="60">Misius</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FD5DB4FD6B6BB491" author="Evans" box="[723,865,933,958]" pageId="1" pageNumber="60" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="05F74D2CFFB09A14FC13B4FD6C7BB491" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[925,1137,933,958]" class="Insecta" family="Hesperiidae" genus="Racta" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="F083EABDFFB09A14FC13B4FD6BD5B492" box="[925,991,934,957]" italics="true" pageId="1" pageNumber="60">Racta</emphasis>
<bibRefCitation id="A6664B5EFFB09A14FC6BB4FD6C7BB491" author="Evans" box="[997,1137,933,958]" pageId="1" pageNumber="60" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
(genera of
<taxonomicName id="05F74D2CFFB09A14FB7DB4FE6D66B491" box="[1267,1388,933,958]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="tribe" tribe="Hesperiini">Hesperiini</taxonomicName>
), all present such orange bands, but lack the elongated tripartite or bipartite brand on male DFW. Other genera of
<taxonomicName id="05F74D2CFFB09A14FF19B4B66906B329" box="[151,268,1005,1030]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="tribe" tribe="Hesperiini">Hesperiini</taxonomicName>
might also present such bands, but have shorter antennae and/or flatten palp. Additionally, the presence and development of lateral projections in the arms of tegumen besides the gnathos have not been seen in these genera.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB09A14FF49B3026D43B0A0" pageId="1" pageNumber="60" type="description">
<paragraph id="C24836AFFFB09A14FF49B3026D43B0A0" blockId="1.[151,1437,825,1935]" pageId="1" pageNumber="60">
<emphasis id="F083EABDFFB09A14FF49B30269A2B35D" bold="true" box="[199,424,1113,1138]" pageId="1" pageNumber="60">Description. Male:</emphasis>
Forewing length
<quantity id="050F9B4AFFB09A14FDF9B3026B08B35D" box="[631,770,1113,1138]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.4" metricValueMin="1.05" pageId="1" pageNumber="60" unit="mm" value="12.25" valueMax="14.0" valueMin="10.5">10.514mm</quantity>
. Eyes red. Vertex black scattered with yellowish to range scales. Antennae longer than 1/2 costa; antennal club short (1/4 shaft), shaft ventrally yellowish in basal portion or on whole ventral surface; nudum 1214, on apiculus extended to the club. Palpus quadrate (inner edge equal to transverse width), second segment ventrally yellowish, third segment of medium length (around half of the second segment length), cylindrical. Thorax and legs black covered with long yellowish to orange scales; midtibiae spined; hindtibiae with two pairs of spurs. DFW marked with orange spots or bands; brand thin or large, perpendicular to CuA, extended from CuA1 to 2A. VFW costal area and apical spots marked from ground color, either yellowish or orange; colored scales compact, present in entire costal area; apical spot contiguous with marginal band, sometimes also with costa; distal cellular spot faint or absent, opaque, yellowish; cell area different by variable yellowish or orange spot. DHW marked with yellowish to orange discal bands, or completely dark brown. VHW yellowish to dark orange; reddish discal spot, yellowish bands and a marked yellowish streak in CuA2-2A are variably present according to the species.
<emphasis id="F083EABDFFB09A14FE3FB2B06A15B12B" bold="true" box="[433,543,1515,1540]" pageId="1" pageNumber="60">Genitalia</emphasis>
. Median posterior apophyses of tegumen present, developed (almost the size or longer than the fenestra), thin or large (lobular), simple or bilobed; fenestra present, triangular or semicircular; lateral apophyses of tegumen present, symmetrical or asymetrical, pointed, squared or rounded. Saccus reduced (shorter than tegumen lenght), lobed. Uncus symmetric or asymmetric, bifid (arms projected); arms medially separated with a pair of small spines on each tip. Gnathos developed, hooked with median membranous patch. Valvae symmetrical or asymmetrical, with dorso-posterior cleft dividing ampulla from harpe; sacculus triangular; harpe projected, hook-like, filled with spines on apex; ampulla thin (less than half of valva width), margin smooth. Aedeagus cylindrical, slightly shorter or longer than the saccs + valva length; coecum of aedeagus globular, short, curved or dorsally straight; dorso-posterior end of aedeagus hollowed, hollow long, extending anteriorly on aedeagus, until the of bulbus ejaculatorius; ventro-posterior end of aedeagus projected as a lobe, with lateral dorsal projections folded over the vesica opening; vesica without cornuti. Fultura inferior directed dorsally and posteriorly; projections broad, extending only laterally of aedeagus; antero-ventral lobes reduced or absent.
</paragraph>
</subSubSection>
</treatment>
</document>