treatments-xml/data/22/14/6B/22146B0FBDE45A458C99DB1C29B01FFE.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.941.49100" ID-GBIF-Dataset="0a42a21d-4646-481d-a3e0-6adb2693eb9b" ID-PMC="PMC7311483" ID-Pensoft-Pub="1313-2970-941-25" ID-Pensoft-UUID="1997660E3DE658719B9E6AA8DBE6E2D0" ID-PubMed="32595406" ID-ZooBank="9095718C8FB046188F9A466CD3B0D3BF" ModsDocID="1313-2970-941-25" checkinTime="1592452103238" checkinUser="pensoft" docAuthor="Chiu, Ming-Chung, Huang, Chin-Gi, Wu, Wen-Jer, Lin, Zhao-Hui, Chen, Hsuan-Wien &amp; Shiao, Shiuh-Feng" docDate="2020" docId="22146B0FBDE45A458C99DB1C29B01FFE" docLanguage="en" docName="ZooKeys 941: 25-48" docOrigin="ZooKeys 941" docSource="http://dx.doi.org/10.3897/zookeys.941.49100" docTitle="Gordius chiashanus Chiu 2020, sp. nov." docType="treatment" docUuid="E904851F-6F48-423D-9AC2-5A7BB595FA7B" docUuidSource="ZooBank" docVersion="4" id="1997660E3DE658719B9E6AA8DBE6E2D0" lastPageNumber="25" masterDocId="1997660E3DE658719B9E6AA8DBE6E2D0" masterDocTitle="A new millipede-parasitizing horsehair worm, Gordius chiashanus sp. nov., at medium altitudes in Taiwan (Nematomorpha, Gordiida)" masterLastPageNumber="48" masterPageNumber="25" pageNumber="25" updateTime="1668168965315" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new millipede-parasitizing horsehair worm, Gordius chiashanus sp. nov., at medium altitudes in Taiwan (Nematomorpha, Gordiida)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chiu, Ming-Chung</mods:namePart>
<mods:affiliation>Department of Biology, Kobe University, Kobe 657 - 8501, Japan &amp; Current address: Department of Biology, National Changhua University of Education, Changhua City 55007, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Chin-Gi</mods:namePart>
<mods:affiliation>National Mosquito-Borne Diseases Control Research Center, National Health Research Institutes, Tainan City 704, Taiwan &amp; Department of Earth and Life Science, University of Taipei, Taipei 100, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wu, Wen-Jer</mods:namePart>
<mods:affiliation>Department of Entomology, National Taiwan University, Taipei 106, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Zhao-Hui</mods:namePart>
<mods:affiliation>Department of Biological Resources, National Chiayi University, Chiayi 300, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Hsuan-Wien</mods:namePart>
<mods:affiliation>Department of Biological Resources, National Chiayi University, Chiayi 300, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shiao, Shiuh-Feng</mods:namePart>
<mods:affiliation>Department of Entomology, National Taiwan University, Taipei 106, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">sfshiao@ntu.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>941</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>25</mods:start>
<mods:end>48</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.941.49100</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.941.49100</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-941-25</mods:identifier>
<mods:identifier type="ZooBank">9095718C8FB046188F9A466CD3B0D3BF</mods:identifier>
<mods:identifier type="Pensoft-UUID">1997660E3DE658719B9E6AA8DBE6E2D0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="164613286" LSID="urn:lsid:zoobank.org:act:E904851F-6F48-423D-9AC2-5A7BB595FA7B" httpUri="http://treatment.plazi.org/id/22146B0FBDE45A458C99DB1C29B01FFE" lastPageNumber="25" pageId="0" pageNumber="25">
<subSubSection pageId="0" pageNumber="25" type="nomenclature">
<paragraph pageId="0" pageNumber="25">
<taxonomicName LSID="22146B0F-BDE4-5A45-8C99-DB1C29B01FFE" authority="Chiu" authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus" status="sp. nov.">Gordius chiashanus Chiu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="25">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type locality">
<paragraph pageId="0" pageNumber="25">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="25">
Dinghu (
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="29.10" value="23.491417">23°29'29.10&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="43" orientation="longitude" precision="1" seconds="19.00" value="120.72195">120°43'19.00&quot;E</geoCoordinate>
), Alishan township, Chiayi county, Taiwan (holotype). Paratypes were collected from Dasyueshan (Heping district, Taichung city), Xitou (Lugu township, Nantou county), Shihjhuo, Fenqihu (Zhuqi township, Chiayi county), Dinghu (Alishan township, Chiayi county), and Hongshi forest road (Haituan township, Taitung county). Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Gordius chiashanus sp. nov. specimen information." httpUri="http://table.plazi.org/id/7BB6E77FF7BC881F889427151EE368B7" pageId="0" pageNumber="25" tableUuid="7BB6E77FF7BC881F889427151EE368B7">1</tableCitation>
presents detailed information of the locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type material">
<paragraph pageId="0" pageNumber="25">Type material.</paragraph>
<paragraph pageId="0" pageNumber="25">
Partial bodies of the holotype and allotype were deposited at the National Museum of Natural Science, Taichung, Taiwan. Paratypes were deposited at the National Museum of Natural Science, Taichung, Taiwan and Lake Biwa Museum, Shiga, Japan (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Gordius chiashanus sp. nov. specimen information." httpUri="http://table.plazi.org/id/7BB6E77FF7BC881F889427151EE368B7" pageId="0" pageNumber="25" tableUuid="7BB6E77FF7BC881F889427151EE368B7">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="7BB6E77FF7BC881F889427151EE368B7" httpUri="http://table.plazi.org/id/7BB6E77FF7BC881F889427151EE368B7" pageId="0" pageNumber="25" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Table 1.</emphasis>
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. specimen information.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
<table pageId="0" pageNumber="25">
<tr pageId="0" pageNumber="25">
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Collection date</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">GenBank no.</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Locality</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Longitude and latitude</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Collector</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Depository</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Sex</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Status</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Length (mm)</th>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">20-XI-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN7848311</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Dasyueshan (Heping, Taichung, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="24" direction="north" minutes="14" orientation="latitude" precision="1" seconds="47.90" value="24.24664">24°14'47.90&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="56" orientation="longitude" precision="1" seconds="06.80" value="120.93523">120°56'06.80&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ta-Chih Chen</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">430</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">26-XI-2008</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784832</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hongshi trail (Haituan, Taitung, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="04" orientation="latitude" precision="1" seconds="14.50" value="23.070696">23°04'14.50&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="07" orientation="longitude" precision="1" seconds="58.30" value="121.132866">121°07'58.30&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Po-Yen Chen</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">744</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">22-I-2008</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784841</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="01.70" value="23.483807">23°29'01.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="42" orientation="longitude" precision="1" seconds="05.90" value="120.70164">120°42'05.90&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Hsuan Tsai</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">860</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">9-II-2007</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784833</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="01.70" value="23.483807">23°29'01.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="42" orientation="longitude" precision="1" seconds="05.90" value="120.70164">120°42'05.90&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Hsuan Tsai</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">707</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">8-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784819</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Dinghu (Alishan, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="29.10" value="23.491417">23°29'29.10&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="43" orientation="longitude" precision="1" seconds="19.00" value="120.72195">120°43'19.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">LBM</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">771</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">8-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784820</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Dinghu (Alishan, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="29.10" value="23.491417">23°29'29.10&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="43" orientation="longitude" precision="1" seconds="19.00" value="120.72195">120°43'19.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">734</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">8-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784821</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Dinghu (Alishan, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="29.10" value="23.491417">23°29'29.10&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="43" orientation="longitude" precision="1" seconds="19.00" value="120.72195">120°43'19.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">726</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784822</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">LBM</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">803</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784823</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">LBM</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">756</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784824</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">594</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784825</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">383</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784826</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">676</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-XII-2013</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784827</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Hua-Te Fang</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">474</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784828</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">749</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784829</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">666</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2017</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784830</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Fenqihu (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="30" orientation="latitude" precision="1" seconds="12.70" value="23.503529">23°30'12.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="36.00" value="120.69334">120°41'36.00&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">717</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2016</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784816</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">LBM</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">498</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2016</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784817</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">403</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18-XII-2016</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784818</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">LBM</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">549</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">9-II-2008</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784842</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">572</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">10-XII-2011</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784840</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Ming-Chung Chiu</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Free-living adult</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">502</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17-III-2019</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784839</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Xitou (Lugu, Nantou, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="40" orientation="latitude" precision="1" seconds="21.30" value="23.672583">23°40'21.30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="47" orientation="longitude" precision="1" seconds="27.50" value="120.79097">120°47'27.50&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Zhao-Hui Lin</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Dead worm in host</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">23-VII-2018</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784834</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="29" orientation="latitude" precision="1" seconds="01.70" value="23.483807">23°29'01.70&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="42" orientation="longitude" precision="1" seconds="05.90" value="120.70164">120°42'05.90&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Wei Li</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Immature worm</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">660</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">28-VII-2018</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784835</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="28" orientation="latitude" precision="1" seconds="22.60" value="23.472944">23°28'22.60&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="42.80" value="120.69522">120°41'42.80&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Wei Li</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Immature worm</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">894</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">28-VII-2018</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784836</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="28" orientation="latitude" precision="1" seconds="22.60" value="23.472944">23°28'22.60&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="42.80" value="120.69522">120°41'42.80&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Wei Li</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Immature worm</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">420</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">28-VII-2018</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784837</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="28" orientation="latitude" precision="1" seconds="22.60" value="23.472944">23°28'22.60&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="42.80" value="120.69522">120°41'42.80&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Wei Li</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Immature worm</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">442</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">28-VII-2018</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">MN784838</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Shihjhuo (Zhuqi, Chiayi, Taiwan)</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<geoCoordinate degrees="23" direction="north" minutes="28" orientation="latitude" precision="1" seconds="22.60" value="23.472944">23°28'22.60&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="41" orientation="longitude" precision="1" seconds="42.80" value="120.69522">120°41'42.80&quot;E</geoCoordinate>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Yu-Wei Li</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">NMNS</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">Immature worm</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">426</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="25">
<tableNote pageId="0" pageNumber="25">LBM: Lake Biwa Museum; NMNS: National Museum of Natural Science. 1 Holotype.</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type hosts">
<paragraph pageId="0" pageNumber="25">Type hosts.</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spirobolus" order="Spirobolida" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Spirobolus</emphasis>
</taxonomicName>
sp. nov. (Hsu and Chang, unpublished) (
<taxonomicName authorityName="Brandt" authorityYear="1833" class="Diplopoda" lsidName="" pageId="0" pageNumber="25" rank="class">Diplopoda</taxonomicName>
:
<taxonomicName authorityName="Bollman" authorityYear="1893" family="Spirobolidae" lsidName="" pageId="0" pageNumber="25" rank="family">Spirobolidae</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="etymology">
<paragraph pageId="0" pageNumber="25">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="25">
The specific name is the combination of
<emphasis italics="true" pageId="0" pageNumber="25">chia</emphasis>
, referring to the place (Chiayi county) where the first sample was found, and
<emphasis italics="true" pageId="0" pageNumber="25">shan</emphasis>
, referring to the Chinese word for
<normalizedToken originalValue="“mountains.”">&quot;mountains.&quot;</normalizedToken>
The word
<emphasis italics="true" pageId="0" pageNumber="25">chia</emphasis>
is also in memory of our friend, Chia-Chih Lin, who died in an accident in a field experiment.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="description">
<paragraph pageId="0" pageNumber="25">Description.</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Male adults (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 11) (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side B, C SEM images of the anterior end surface that is (B) smooth with scattered short bristles and (C) wrinkled D close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (A), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25">1</figureCitation>
-
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mid-body of male Gordius chiashanus sp. nov. A, B SEM images of (A) cuticle in the mid-body with scattered short bristles (arrows) and (B) close-up view of a short bristle C, D white spots and dorsal and ventral dark pigmented line examined using (C) a compound microscope and (D) a stereomicroscope. Scale bars: 1 mm (A, C, D), 5 μm (B)." figureDoi="10.3897/zookeys.941.49100.figure3" httpUri="https://binary.pensoft.net/fig/421156" pageId="0" pageNumber="25">3</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5</figureCitation>
).
</emphasis>
Body length 627.94
<normalizedToken originalValue="±">+/-</normalizedToken>
154.75 (383-860) mm, width (widest, after dehydration) 1.30
<normalizedToken originalValue="±">+/-</normalizedToken>
0.31 (0.81-2.06) mm, light to dark brown, smooth, and covered with mucus-like structure (viscous liquid on live worms with rainbow-like reflection (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5C</figureCitation>
, Suppl. material 1: Video S1), and created haze that surrounded the body surface in hot water (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 1.</emphasis>
Anterior end of male
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side
<emphasis bold="true" pageId="0" pageNumber="25">B, C</emphasis>
SEM images of the anterior end surface that is (
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
) smooth with scattered short bristles and (
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
) wrinkled
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
), 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">B-D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
Anterior end columnar and spherical; anterior tip white (white cap) with a dark -brown collar and a vertical white stripe on the ventral side (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side B, C SEM images of the anterior end surface that is (B) smooth with scattered short bristles and (C) wrinkled D close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (A), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25">1A</figureCitation>
). Under SEM, surface of anterior end appeared smooth (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side B, C SEM images of the anterior end surface that is (B) smooth with scattered short bristles and (C) wrinkled D close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (A), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25">1B</figureCitation>
) or wrinkled (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side B, C SEM images of the anterior end surface that is (B) smooth with scattered short bristles and (C) wrinkled D close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (A), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25">1C</figureCitation>
) on the tip of one sample; scattered short bristles (11.24
<normalizedToken originalValue="±">+/-</normalizedToken>
6.57 (4.92-22.24)
<normalizedToken originalValue="µm">µm</normalizedToken>
in length) scattered except on tip in most samples (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the ventral side of the anterior end showing a white cap, dark-brown collar, and vertical white stripe on the ventral side B, C SEM images of the anterior end surface that is (B) smooth with scattered short bristles and (C) wrinkled D close-up view of the dotted square in C showing the short bristles (arrows) covered by a wrinkled structure. Scale bars: 2 mm (A), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure1" httpUri="https://binary.pensoft.net/fig/421154" pageId="0" pageNumber="25">1B, D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
Cuticle in mid-body ornamented with a dorsal and a ventral dark pigment line; white spots scattered across entire body surface (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mid-body of male Gordius chiashanus sp. nov. A, B SEM images of (A) cuticle in the mid-body with scattered short bristles (arrows) and (B) close-up view of a short bristle C, D white spots and dorsal and ventral dark pigmented line examined using (C) a compound microscope and (D) a stereomicroscope. Scale bars: 1 mm (A, C, D), 5 μm (B)." figureDoi="10.3897/zookeys.941.49100.figure3" httpUri="https://binary.pensoft.net/fig/421156" pageId="0" pageNumber="25">3C, D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5A</figureCitation>
). Under SEM, cuticle surface appeared smooth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mid-body of male Gordius chiashanus sp. nov. A, B SEM images of (A) cuticle in the mid-body with scattered short bristles (arrows) and (B) close-up view of a short bristle C, D white spots and dorsal and ventral dark pigmented line examined using (C) a compound microscope and (D) a stereomicroscope. Scale bars: 1 mm (A, C, D), 5 μm (B)." figureDoi="10.3897/zookeys.941.49100.figure3" httpUri="https://binary.pensoft.net/fig/421156" pageId="0" pageNumber="25">3A</figureCitation>
) with a few scattered short or cone-like bristles (6.75
<normalizedToken originalValue="±">+/-</normalizedToken>
2.37 (2.31-10.34)
<normalizedToken originalValue="µm">µm</normalizedToken>
in length) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mid-body of male Gordius chiashanus sp. nov. A, B SEM images of (A) cuticle in the mid-body with scattered short bristles (arrows) and (B) close-up view of a short bristle C, D white spots and dorsal and ventral dark pigmented line examined using (C) a compound microscope and (D) a stereomicroscope. Scale bars: 1 mm (A, C, D), 5 μm (B)." figureDoi="10.3897/zookeys.941.49100.figure3" httpUri="https://binary.pensoft.net/fig/421156" pageId="0" pageNumber="25">3A, B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
Posterior end divided into two tail lobes (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2A, B</figureCitation>
), each lobe 855.24
<normalizedToken originalValue="±">+/-</normalizedToken>
100.89 (658.39-994.88)
<normalizedToken originalValue="µm">µm</normalizedToken>
long and 458.55
<normalizedToken originalValue="±">+/-</normalizedToken>
76.52 (365.95-643.00)
<normalizedToken originalValue="µm">µm</normalizedToken>
wide with length-to-width ratio of 1.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.26 (1.49-2.42). Inner side of lobe tips white (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2A</figureCitation>
). Under SEM, inner side of tail lobes concave in some samples; cuticle surface smooth, but one sample exhibited flat areoles on inner side of lobe tips; short bristles scattered across the surface and concentrated in most samples on lobe tips (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2C</figureCitation>
) and on inner side of lobe tips forming a bristle field (322.67
<normalizedToken originalValue="±">+/-</normalizedToken>
99.34 (187.60-412.75)
<normalizedToken originalValue="µm">µm</normalizedToken>
long and 71.82
<normalizedToken originalValue="±">+/-</normalizedToken>
35.49 (44.81-114.54)
<normalizedToken originalValue="µm">µm</normalizedToken>
wide) on each of tail lobe posterior to tips of postcloacal crescent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2D</figureCitation>
). Postcloacal crescent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2A, B</figureCitation>
) 718.61
<normalizedToken originalValue="±">+/-</normalizedToken>
118.77 (536.14-984.34)
<normalizedToken originalValue="µm">µm</normalizedToken>
long and 86.7
<normalizedToken originalValue="±">+/-</normalizedToken>
15.62 (54.73-118.65)
<normalizedToken originalValue="µm">µm</normalizedToken>
wide and located on ventral side near base of tail lobes. Crescent generally semicircular or slightly angled, but a few samples exhibited a straightened form of crescent. Branches of postcloacal crescent usually ended at tail lobes. Cloacal opening circular (40.5
<normalizedToken originalValue="±">+/-</normalizedToken>
21.87 (27.41-56.14)
<normalizedToken originalValue="µm">µm</normalizedToken>
) and anterior to postcloacal crescent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2A, B</figureCitation>
). Wall inside cloacal opening exhibited areoles (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Posterior end of male Gordius chiashanus sp. nov. A stereomicroscopic image of the posterior end B-D SEM images of (B) overview of the posterior end with bristles concentrated on the (C) lobe tips (arrow), and (D) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows) E cloacal opening with areoles on the inside wall. Scale bars: 1 mm (A), 500 μm (B), 200 μm (C-D), 50 μm (E)." figureDoi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25">2E</figureCitation>
); no circumcloacal spine or bristles observed in region next to cloacal opening.
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure2" httpUri="https://binary.pensoft.net/fig/421155" pageId="0" pageNumber="25" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 2.</emphasis>
Posterior end of male
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
stereomicroscopic image of the posterior end
<emphasis bold="true" pageId="0" pageNumber="25">B-D</emphasis>
SEM images of (
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
) overview of the posterior end with bristles concentrated on the (
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
) lobe tips (arrow), and (
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
) inner side of the lobe tips and the formation of a bristle field on each tail lobe posterior to the tips of the postcloacal crescent (arrows)
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
cloacal opening with areoles on the inside wall. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
), 500
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
), 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">C-D</emphasis>
), 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.941.49100.figure3" httpUri="https://binary.pensoft.net/fig/421156" pageId="0" pageNumber="25" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 3.</emphasis>
Mid-body of male
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A, B</emphasis>
SEM images of (
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
) cuticle in the mid-body with scattered short bristles (arrows) and (
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
) close-up view of a short bristle
<emphasis bold="true" pageId="0" pageNumber="25">C, D</emphasis>
white spots and dorsal and ventral dark pigmented line examined using (
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
) a compound microscope and (
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
) a stereomicroscope. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">A, C, D</emphasis>
), 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Female adults (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 4) (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5</figureCitation>
).
</emphasis>
Body length 659.75
<normalizedToken originalValue="±">+/-</normalizedToken>
77.06 (549-717) mm, width (widest, after dehydration) 1.54
<normalizedToken originalValue="±">+/-</normalizedToken>
0.54 (1.00-2.03) mm, light to dark brown, smooth, and covered with mucus-like structure. White spots scattered on surface but relatively less obvious than those of male adults (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4F, G</figureCitation>
). Anterior end columnar and spherical. Anterior tip white (white cap) with a dark-brown collar and exhibited a vertical white stripe on the ventral side (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4A</figureCitation>
). Under SEM, surface of anterior end smooth and exhibited scattered short bristles (16.75
<normalizedToken originalValue="±">+/-</normalizedToken>
4.60 (13.39-23.56)
<normalizedToken originalValue="µm">µm</normalizedToken>
in length) except at tip (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4B</figureCitation>
). Cuticle in mid-body ornamented with a dorsal and a ventral dark pigment line (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4G</figureCitation>
). Under SEM, cuticle surface smooth with a few short or cone-like bristles (7.24
<normalizedToken originalValue="±">+/-</normalizedToken>
2.01 (4.94-9.99)
<normalizedToken originalValue="µm">µm</normalizedToken>
in length) scattered. Posterior end columnar and rounded at tip (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4E</figureCitation>
) and did not exhibit scattered bristles (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4D</figureCitation>
). Cloacal opening on terminal end (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female Gordius chiashanus sp. nov. A, B anterior end examined using a (A) stereomicroscope and (B) SEM C-E posterior end with the terminal view examined using a (C) stereomicroscope and (D) SEM, and the (E) lateral view examined using a stereomicroscope F, G mid-body examined using a (F) stereomicroscope and (G) compound microscope. Co, cloacal opening. Scale bars: 1 mm (A, F, G), 200 μm (B-D)." figureDoi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25">4C, D</figureCitation>
) circular and 36.56
<normalizedToken originalValue="±">+/-</normalizedToken>
23.23 (24.68-48.45)
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter.
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure4" httpUri="https://binary.pensoft.net/fig/421157" pageId="0" pageNumber="25" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 4.</emphasis>
Female
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A, B</emphasis>
anterior end examined using a (
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
) stereomicroscope and (
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
) SEM
<emphasis bold="true" pageId="0" pageNumber="25">C-E</emphasis>
posterior end with the terminal view examined using a (
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
) stereomicroscope and (
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
) SEM, and the (
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
) lateral view examined using a stereomicroscope
<emphasis bold="true" pageId="0" pageNumber="25">F, G</emphasis>
mid-body examined using a (
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
) stereomicroscope and (
<emphasis bold="true" pageId="0" pageNumber="25">G</emphasis>
) compound microscope. Co, cloacal opening. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">A, F, G</emphasis>
), 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">B-D</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 5.</emphasis>
Field observation of
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
hazy appearance (arrows) surrounding the body surface in hot water
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
spermatophore (arrow) on a female collected on the surface of the soil
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
rainbow-like reflection on the body surface
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
free-living adult collected in wet soil
<emphasis bold="true" pageId="0" pageNumber="25">E, F</emphasis>
infected host,
<taxonomicName authorityName="Brandt" authorityYear="1833" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spirobolus" order="Spirobolida" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Spirobolus</emphasis>
</taxonomicName>
sp. nov. (Hsu and Chang, unpublished), harboring (
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
) three immature worms (arrow) and (
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
) an adult worm. Photographs courtesy of (
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
) Fang, Hua-Te and (
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
) Hung, Ming-Chin. Scale bars: 1 cm (
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Eggs (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 12) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6C-E</figureCitation>
).
</emphasis>
Egg strings (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6E</figureCitation>
) 7.41
<normalizedToken originalValue="±">+/-</normalizedToken>
3.46 (3.78-13.70) mm in length and 1.13
<normalizedToken originalValue="±">+/-</normalizedToken>
0.12 (0.86-1.25) mm in width; white or light yellow in color, deposited in water as short pieces not adhering to substrate. Eggs round, 54.16
<normalizedToken originalValue="±">+/-</normalizedToken>
242 2.89 (49.88-58.61)
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter. Developing embryo surrounded by an inner membrane (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6C, D</figureCitation>
) separated by a distinct space from outer egg shell 14.35
<normalizedToken originalValue="±">+/-</normalizedToken>
1.41 (12.43-17.33)
<normalizedToken originalValue="µm">µm</normalizedToken>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Living larvae (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 10) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6B</figureCitation>
).
</emphasis>
Eggs developed for approximately 49 days. Hatched larvae remained near egg strings or moved inside eggshells. Under light microscopy, living larvae appeared cylindrical with a single posterior spine. Preseptum length 32.33
<normalizedToken originalValue="±">+/-</normalizedToken>
4.53 (27.06-40.04)
<normalizedToken originalValue="µm">µm</normalizedToken>
, and the width 18.04
<normalizedToken originalValue="±">+/-</normalizedToken>
0.86 (16.70-19.12)
<normalizedToken originalValue="µm">µm</normalizedToken>
. Postseptum length 83.05
<normalizedToken originalValue="±">+/-</normalizedToken>
8.31 (66.50-92.66)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 15.05
<normalizedToken originalValue="±">+/-</normalizedToken>
0.73 (14.21-16.10)
<normalizedToken originalValue="µm">µm</normalizedToken>
; proboscis length 14.94
<normalizedToken originalValue="±">+/-</normalizedToken>
1.99 (12.35-18.48)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 4.11
<normalizedToken originalValue="±">+/-</normalizedToken>
0.85 (2.77-5.34)
<normalizedToken originalValue="µm">µm</normalizedToken>
; pseudointestine length 60.60
<normalizedToken originalValue="±">+/-</normalizedToken>
5.40 (54.99-70.12)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 11.66
<normalizedToken originalValue="±">+/-</normalizedToken>
1.42 (8.84-13.56)
<normalizedToken originalValue="µm">µm</normalizedToken>
, unequally subdivided, elongated oval with a depression in anterior end (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Larvae treated with hot water (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 2) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6A</figureCitation>
).
</emphasis>
Larvae treated with hot water similar in morphology but larger than living larvae. Preseptum length 44.57
<normalizedToken originalValue="±">+/-</normalizedToken>
0.13 (44.48-44.66)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 17.96
<normalizedToken originalValue="±">+/-</normalizedToken>
0.16 (17.85-18.08)
<normalizedToken originalValue="µm">µm</normalizedToken>
. Postseptum length 118.23
<normalizedToken originalValue="±">+/-</normalizedToken>
1.91 (116.88-119.58)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 15.36
<normalizedToken originalValue="±">+/-</normalizedToken>
0.68 (14.88-15.84)
<normalizedToken originalValue="µm">µm</normalizedToken>
. Proboscis length 12.63
<normalizedToken originalValue="±">+/-</normalizedToken>
1.18 (11.80-13.47)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 3.26
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05 (3.23-3.30)
<normalizedToken originalValue="µm">µm</normalizedToken>
; pseudointestine length 77.99
<normalizedToken originalValue="±">+/-</normalizedToken>
5.22 (74.30-81.68)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 13.99
<normalizedToken originalValue="±">+/-</normalizedToken>
0.81 (13.41-14.56)
<normalizedToken originalValue="µm">µm</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">
Field-collected cysts (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 5) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6F-H</figureCitation>
).
</emphasis>
Larvae in cysts unfolded (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 4) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6F</figureCitation>
) or exhibited a postseptum folded twice (
<emphasis italics="true" pageId="0" pageNumber="25">N</emphasis>
= 1) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6G, H</figureCitation>
). Unfolded larvae morphologically similar to larvae but larger in size; preseptum length was 60.18
<normalizedToken originalValue="±">+/-</normalizedToken>
6.72 (50.40-65.18)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 20.87
<normalizedToken originalValue="±">+/-</normalizedToken>
0.52 (20.28-21.33)
<normalizedToken originalValue="µm">µm</normalizedToken>
; postseptum length 127.33
<normalizedToken originalValue="±">+/-</normalizedToken>
20.05 (105.10-146.05)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 19.82
<normalizedToken originalValue="±">+/-</normalizedToken>
2.27 (17.61-22.91)
<normalizedToken originalValue="µm">µm</normalizedToken>
; proboscis length 15.46
<normalizedToken originalValue="±">+/-</normalizedToken>
1.67 (13.84-17.56)
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 4.10
<normalizedToken originalValue="±">+/-</normalizedToken>
0.68 (3.09-4.52)
<normalizedToken originalValue="µm">µm</normalizedToken>
; pseudointestine not visible (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6F</figureCitation>
). Folded larva (length 34.97
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 30.47
<normalizedToken originalValue="µm">µm</normalizedToken>
) fold twice and surrounded by a clear cyst wall, 47.86
<normalizedToken originalValue="µm">µm</normalizedToken>
in total length and 42.40
<normalizedToken originalValue="µm">µm</normalizedToken>
in total width; proboscis length 15.57
<normalizedToken originalValue="µm">µm</normalizedToken>
, width 5.09
<normalizedToken originalValue="µm">µm</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6G</figureCitation>
); a single posterior spine visible after treatment with a solution of 5% KOH (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Immature stages of Gordius chiashanus sp. nov. A, B free-living larva (A) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow) C, D eggs with the inner membrane examined using an (C) SEM and (D) compound microscope E egg strings F-H cysts in the paratenic host with (F) a unfolded larva and (G) a folded larva, showing (H) a single posterior spine (arrow) after treatment with a 5 % KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50 µm (A-D, F-H), 1 mm (E)." figureDoi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25">6H</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure6" httpUri="https://binary.pensoft.net/fig/421159" pageId="0" pageNumber="25" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 6.</emphasis>
Immature stages of
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="25">A, B</emphasis>
free-living larva (
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
) treated with hot water and a living larva showing the depression in the anterior end of the pseudointestine (arrow)
<emphasis bold="true" pageId="0" pageNumber="25">C, D</emphasis>
eggs with the inner membrane examined using an (
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
) SEM and (
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
) compound microscope
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
egg strings
<emphasis bold="true" pageId="0" pageNumber="25">F-H</emphasis>
cysts in the paratenic host with (
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
) a unfolded larva and (
<emphasis bold="true" pageId="0" pageNumber="25">G</emphasis>
) a folded larva, showing (
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
) a single posterior spine (arrow) after treatment with a 5% KOH solution. Abbreviations: Ho, hooklet; PostS, postseptum; PreS, preseptum; Pro, proboscis; PsI, pseudointestine. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="25">A-D, F-H</emphasis>
), 1 mm (
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
).
</paragraph>
</caption>
<caption ID-Table-UUID="B55C18A583798285A8D76C16525946D4" httpUri="http://table.plazi.org/id/B55C18A583798285A8D76C16525946D4" pageId="0" pageNumber="25" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Table 2.</emphasis>
List of COI sequences obtained from GenBank for phylogenetic analyses in this study.
</paragraph>
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<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Accession number</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">Species/clade</th>
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<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Heinze" authorityYear="1952" class="Gordioida" family="Gordiidae" genus="Acutogordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acutogordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Acutogordius</emphasis>
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<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade 8)
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<td colspan="1" pageId="0" pageNumber="25" rowspan="1">KY172803</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<normalizedToken originalValue="">''</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<bibRefCitation DOI="https://doi.org/10.1017/S0031182017000233" author="Tobias, ZJC" journalOrPublisher="Parasitology" pageId="0" pageNumber="25" pagination="1026 - 1040" refId="B39" refString="Tobias, ZJC, Yadav, AK, Schmidt-Rhaesa, A, Poulin, R, 2017. Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha). Parasitology 144: 1026 - 1040, DOI: https://doi.org/10.1017/S0031182017000233" title="Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha)." url="https://doi.org/10.1017/S0031182017000233" volume="144" year="2017">Tobias et al. 2017</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">KY172747</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName baseAuthorityName="Örley" baseAuthorityYear="1881" class="Gordioida" family="Chordodidae" genus="Parachordodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parachordodes diblastus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="diblastus">
<emphasis italics="true" pageId="0" pageNumber="25">Parachordodes diblastus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<bibRefCitation DOI="https://doi.org/10.1017/S0031182017000233" author="Tobias, ZJC" journalOrPublisher="Parasitology" pageId="0" pageNumber="25" pagination="1026 - 1040" refId="B39" refString="Tobias, ZJC, Yadav, AK, Schmidt-Rhaesa, A, Poulin, R, 2017. Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha). Parasitology 144: 1026 - 1040, DOI: https://doi.org/10.1017/S0031182017000233" title="Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha)." url="https://doi.org/10.1017/S0031182017000233" volume="144" year="2017">Tobias et al. 2017</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">KY172778</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<normalizedToken originalValue="">''</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<bibRefCitation DOI="https://doi.org/10.1017/S0031182017000233" author="Tobias, ZJC" journalOrPublisher="Parasitology" pageId="0" pageNumber="25" pagination="1026 - 1040" refId="B39" refString="Tobias, ZJC, Yadav, AK, Schmidt-Rhaesa, A, Poulin, R, 2017. Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha). Parasitology 144: 1026 - 1040, DOI: https://doi.org/10.1017/S0031182017000233" title="Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha)." url="https://doi.org/10.1017/S0031182017000233" volume="144" year="2017">Tobias et al. 2017</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="25">
<tableNote pageId="0" pageNumber="25">
* KY172770 was excluded from the analysis since its high difference from the member of
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
and the high similarity with
<taxonomicName authorityName="Poinar" authorityYear="1991" class="Gordioida" family="Chordodidae" genus="Euchordodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euchordodes nigromaculatus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="nigromaculatus">
<emphasis italics="true" pageId="0" pageNumber="25">Euchordodes nigromaculatus</emphasis>
</taxonomicName>
.
</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="phylogeny">
<paragraph pageId="0" pageNumber="25">Phylogeny.</paragraph>
<paragraph pageId="0" pageNumber="25">
The partial COI sequences of the 18 free-living adults contained 15 haplotypes with 392 invariable sites, nine singletons, and 21 parsimoniously informative sites. The genetic distance among them was 0.0024 within the range of 0.0000-0.0510. The three living adults and six worms inside the hosts were considered conspecific with the 18 free-living adults because of their small genetic distances (0.0000-0.0719). The mean interspecific genetic distances between
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. and other
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
species or clades were in the range of 0.2320-0.4242, and that between
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName authorityName="Chiu, Huang, Wu &amp; Shiao" authorityYear="2017" class="Gordioida" family="Gordiidae" genus="Acutogordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acutogordius taiwanensis" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="taiwanensis">
<emphasis italics="true" pageId="0" pageNumber="25">Acutogordius taiwanensis</emphasis>
</taxonomicName>
was 0.3648 (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Intra- and interspecific mean COI genetic distances of Gordius / Acutogordius species or clades under K 2 P model." httpUri="http://table.plazi.org/id/94A3D0B4E166C3BAB8040575EA7BE3A8" pageId="0" pageNumber="25" tableUuid="94A3D0B4E166C3BAB8040575EA7BE3A8">3</tableCitation>
). In addition to short genetic distances, the conspecific status of the 18 free-living adults was also supported because all the samples were located in a single clade, as indicated by a high bootstrap value. No subgroup was detected because the polytomic topology exhibited low bootstrap values and short genetic distances. The
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
species/clades in the present result were consistent with the results of
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2014.09.010" author="Hanelt, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="211 - 218" refId="B16" refString="Hanelt, B, Schmidt-Rhaesa, A, Bolek, MG, 2015. Cryptic species of hairworm parasites revealed by molecular data and crowdsourcing of specimen collections. Molecular Phylogenetics and Evolution 82: 211 - 218, DOI: https://doi.org/10.1016/j.ympev.2014.09.010" title="Cryptic species of hairworm parasites revealed by molecular data and crowdsourcing of specimen collections." url="https://doi.org/10.1016/j.ympev.2014.09.010" volume="82" year="2015">Hanelt et al. (2015)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.1017/S0031182017000233" author="Tobias, ZJC" journalOrPublisher="Parasitology" pageId="0" pageNumber="25" pagination="1026 - 1040" refId="B39" refString="Tobias, ZJC, Yadav, AK, Schmidt-Rhaesa, A, Poulin, R, 2017. Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha). Parasitology 144: 1026 - 1040, DOI: https://doi.org/10.1017/S0031182017000233" title="Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha)." url="https://doi.org/10.1017/S0031182017000233" volume="144" year="2017">Tobias et al. (2017)</bibRefCitation>
, despite slight differences in the relative relationships among species, which might be attributable to the differences in models used or the shorter sequence adopted in previous studies. The clade of
<taxonomicName lsidName="A. taiwanensis" pageId="0" pageNumber="25" rank="species" species="taiwanensis">
<emphasis italics="true" pageId="0" pageNumber="25">A. taiwanensis</emphasis>
</taxonomicName>
was located within that of the
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
species, and it did not behave as a sister group (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationship of Gordius / Acutogordius spp. restructured using COI partial sequences compared with C. formosanus, E. nigromaculatus, and P. diblastus as out groups. Numbers at the nodes represent the percentage of 1000 bootstrap replicates." figureDoi="10.3897/zookeys.941.49100.figure7" httpUri="https://binary.pensoft.net/fig/421160" pageId="0" pageNumber="25">7</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure7" httpUri="https://binary.pensoft.net/fig/421160" pageId="0" pageNumber="25" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 7.</emphasis>
Phylogenetic relationship of
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Heinze" authorityYear="1952" class="Gordioida" family="Gordiidae" genus="Acutogordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acutogordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Acutogordius</emphasis>
</taxonomicName>
spp. restructured using COI partial sequences compared with
<taxonomicName lsidName="C. formosanus" pageId="0" pageNumber="25" rank="species" species="formosanus">
<emphasis italics="true" pageId="0" pageNumber="25">C. formosanus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="E. nigromaculatus" pageId="0" pageNumber="25" rank="species" species="nigromaculatus">
<emphasis italics="true" pageId="0" pageNumber="25">E. nigromaculatus</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. diblastus" pageId="0" pageNumber="25" rank="species" species="diblastus">
<emphasis italics="true" pageId="0" pageNumber="25">P. diblastus</emphasis>
</taxonomicName>
as out groups. Numbers at the nodes represent the percentage of 1000 bootstrap replicates.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="reproductive season">
<paragraph pageId="0" pageNumber="25">Reproductive season.</paragraph>
<paragraph pageId="0" pageNumber="25">
Free-living adult worms frequently aggregate and mate on wet ground (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5B, C</figureCitation>
) after rain or fog, and they are sometimes found in water or soil (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Field observation of Gordius chiashanus sp. nov. A hazy appearance (arrows) surrounding the body surface in hot water B spermatophore (arrow) on a female collected on the surface of the soil C rainbow-like reflection on the body surface D free-living adult collected in wet soil E, F infected host, Spirobolus sp. nov. (Hsu and Chang, unpublished), harboring (E) three immature worms (arrow) and (F) an adult worm. Photographs courtesy of (D) Fang, Hua-Te and (F) Hung, Ming-Chin. Scale bars: 1 cm (E)." figureDoi="10.3897/zookeys.941.49100.figure5" httpUri="https://binary.pensoft.net/fig/421158" pageId="0" pageNumber="25">5D</figureCitation>
). They suddenly emerge in early December, and their number decreases within 1-2 months (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Seasonal occurrence of free-living adults of Gordius chiashanus sp. nov. Numbers at the bottom indicate the actual number of each bar." figureDoi="10.3897/zookeys.941.49100.figure8" httpUri="https://binary.pensoft.net/fig/421161" pageId="0" pageNumber="25">8</figureCitation>
). During the reproductive season, no infected host was found. The seasonality and pattern of
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. differed from the graph constructed using data from
<taxonomicName lsidName="C. formosanus" pageId="0" pageNumber="25" rank="species" species="formosanus">
<emphasis italics="true" pageId="0" pageNumber="25">C. formosanus</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1645/15-907" author="Chiu, MC" journalOrPublisher="Journal of Parasitology" pageId="0" pageNumber="25" pagination="319 - 326" refId="B9" refString="Chiu, MC, Huang, CG, Wu, WJ, Shiao, SF, 2016. Annual survey of horsehair worm cysts in northern Taiwan, with notes on a single seasonal infection peak in chironomid larvae (Diptera: Chironomidae). Journal of Parasitology 102: 319 - 326, DOI: https://doi.org/10.1645/15-907" title="Annual survey of horsehair worm cysts in northern Taiwan, with notes on a single seasonal infection peak in chironomid larvae (Diptera: Chironomidae)." url="https://doi.org/10.1645/15-907" volume="102" year="2016">Chiu et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="diagnosis">
<paragraph pageId="0" pageNumber="25">Diagnosis and comments.</paragraph>
<paragraph pageId="0" pageNumber="25">
The 21 free-living
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
adults and six juvenile worms from round-backed millipedes were judged as belonging to the same species in accordance with the results that they all were located in the same clade in the phylogenetic tree and had low genetic distances (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationship of Gordius / Acutogordius spp. restructured using COI partial sequences compared with C. formosanus, E. nigromaculatus, and P. diblastus as out groups. Numbers at the nodes represent the percentage of 1000 bootstrap replicates." figureDoi="10.3897/zookeys.941.49100.figure7" httpUri="https://binary.pensoft.net/fig/421160" pageId="0" pageNumber="25">7</figureCitation>
, Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Intra- and interspecific mean COI genetic distances of Gordius / Acutogordius species or clades under K 2 P model." httpUri="http://table.plazi.org/id/94A3D0B4E166C3BAB8040575EA7BE3A8" pageId="0" pageNumber="25" tableUuid="94A3D0B4E166C3BAB8040575EA7BE3A8">3</tableCitation>
). These samples were regarded as a new species,
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov., on the basis of their distribution patterns of bristles on the male tail and presence of a vertical white stripe on the anterior ventral side and areoles on the inside wall of the cloacal opening.
</paragraph>
<caption ID-Table-UUID="94A3D0B4E166C3BAB8040575EA7BE3A8" httpUri="http://table.plazi.org/id/94A3D0B4E166C3BAB8040575EA7BE3A8" pageId="0" pageNumber="25" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Table 3.</emphasis>
Intra- and interspecific mean COI genetic distances of
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Heinze" authorityYear="1952" class="Gordioida" family="Gordiidae" genus="Acutogordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acutogordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Acutogordius</emphasis>
</taxonomicName>
species or clades under K2P model.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
<table pageId="0" pageNumber="25">
<tr pageId="0" pageNumber="25">
<th colspan="2" pageId="0" pageNumber="25" rowspan="1">Species/Clade</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">3</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">4</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">5</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">6</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">7</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">8</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">9</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">10</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">11</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">12</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">13</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">14</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">15</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">16</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">17</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">18</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">19</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">20</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">21</th>
<th colspan="1" pageId="0" pageNumber="25" rowspan="1">22</th>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov.
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.024</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade1)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.285</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.009</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade2)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.312</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.217</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.015</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade3)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.293</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.297</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.275</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.007</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade4)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.308</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.208</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.249</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.157</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.012</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade5)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.272</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.165</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.211</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.227</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.222</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.003</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade6)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.293</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.257</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.251</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.255</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.228</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.259</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.006</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. terrestris" pageId="0" pageNumber="25" rank="species" species="terrestris">
<emphasis italics="true" pageId="0" pageNumber="25">G. terrestris</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.232</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.209</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.265</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.250</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.230</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.222</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.238</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.020</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. robustus" pageId="0" pageNumber="25" rank="species" species="robustus">G. cf. robustus</taxonomicName>
(Clade8)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.265</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.203</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.307</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.338</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.253</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.244</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.251</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.122</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.026</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. attoni" pageId="0" pageNumber="25" rank="species" species="attoni">
<emphasis italics="true" pageId="0" pageNumber="25">G. attoni</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.277</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.229</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.288</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.337</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.274</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.238</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.289</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.231</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.249</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.010</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. balticus" pageId="0" pageNumber="25" rank="species" species="balticus">
<emphasis italics="true" pageId="0" pageNumber="25">G. balticus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.316</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.260</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.298</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.288</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.269</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.274</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.304</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.264</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.323</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.337</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. N178
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.352</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.260</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.313</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.370</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.289</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.340</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.330</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.256</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.290</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.271</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.323</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">13</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. N183
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.329</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.302</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.290</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.373</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.317</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.344</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.365</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.294</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.336</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.277</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.301</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.246</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. N297B
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.424</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.416</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.462</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.547</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.441</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.443</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.478</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.375</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.412</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.348</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.455</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.343</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.414</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. N357
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.332</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.366</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.387</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.420</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.376</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.396</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.302</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.357</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.359</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.379</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.439</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.375</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.434</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.441</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">16</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. KW-2011-A
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.384</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.325</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.327</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.453</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.371</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.336</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.345</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.347</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.348</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.331</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.376</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.332</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.376</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.372</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.424</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. KW-2011-B
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.334</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.375</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.365</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.370</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.334</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.407</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.364</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.302</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.363</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.333</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.380</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.323</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.358</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.333</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.308</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.290</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp. KW-2011-D
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.375</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.300</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.344</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.393</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.373</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.294</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.388</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.388</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.367</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.369</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.405</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.384</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.390</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.374</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.403</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.312</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.301</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">19</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. paranensis" pageId="0" pageNumber="25" rank="species" species="paranensis">
<emphasis italics="true" pageId="0" pageNumber="25">G. paranensis</emphasis>
</taxonomicName>
(Clade1)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.369</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.405</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.381</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.450</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.381</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.410</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.359</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.373</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.395</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.409</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.398</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.408</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.466</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.426</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.453</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.415</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.386</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.440</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.049</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName lsidName="G. paranensis" pageId="0" pageNumber="25" rank="species" species="paranensis">
<emphasis italics="true" pageId="0" pageNumber="25">G. paranensis</emphasis>
</taxonomicName>
(Clade2)
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.337</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.348</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.391</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.436</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.384</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.372</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.368</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.333</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.368</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.345</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.339</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.334</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.385</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.324</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.404</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.357</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.327</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.344</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.377</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.010</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">21</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
sp.
<bibRefCitation DOI="https://doi.org/10.1017/S0031182017000233" author="Tobias, ZJC" journalOrPublisher="Parasitology" pageId="0" pageNumber="25" pagination="1026 - 1040" refId="B39" refString="Tobias, ZJC, Yadav, AK, Schmidt-Rhaesa, A, Poulin, R, 2017. Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha). Parasitology 144: 1026 - 1040, DOI: https://doi.org/10.1017/S0031182017000233" title="Intra- and interspecific genetic diversity of New Zealand hairworms (Nematomorpha)." url="https://doi.org/10.1017/S0031182017000233" volume="144" year="2017">Tobias et al. 2017</bibRefCitation>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.335</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.283</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.293</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.436</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.355</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.311</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.366</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.287</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.337</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.347</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.358</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.254</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.308</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.343</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.353</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.304</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.335</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.321</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.354</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.337</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.012</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="25" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="25">
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">22</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<taxonomicName authorityName="Chiu, Huang, Wu &amp; Shiao" authorityYear="2017" class="Gordioida" family="Gordiidae" genus="Acutogordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acutogordius taiwanensis" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="taiwanensis">
<emphasis italics="true" pageId="0" pageNumber="25">Acutogordius taiwanensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.365</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.343</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.327</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.401</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.386</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.368</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.345</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.322</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.375</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.304</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.336</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.270</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.210</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.462</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.469</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.432</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.376</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.366</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.435</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.389</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">0.311</td>
<td colspan="1" pageId="0" pageNumber="25" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="25">0.002</emphasis>
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="25">
<tableNote pageId="0" pageNumber="25">-Indicates a single haplotype whose intraspecific distance could not be calculated.</tableNote>
</paragraph>
<paragraph pageId="0" pageNumber="25">
The concentration of bristles and spines on the male tail lobes has been previously described in species from the Palaearctic (
<bibRefCitation author="Spiridonov, SE" journalOrPublisher="Proceedings of the Zoological Institute USSR Academy of Sciences" pageId="0" pageNumber="25" pagination="97 - 101" refId="B34" refString="Spiridonov, SE, 1984. Two new Nematomorpha species of the family Gordiidae. Proceedings of the Zoological Institute USSR Academy of Sciences 126: 97 - 101" title="Two new Nematomorpha species of the family Gordiidae." volume="126" year="1984">Spiridonov 1984</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2533.1.1" author="Schmidt-Rhaesa, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="25" pagination="1 - 35" refId="B26" refString="Schmidt-Rhaesa, A, 2010. Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species. Zootaxa 2533: 1 - 35, DOI: https://doi.org/10.11646/zootaxa.2533.1.1" title="Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species." url="https://doi.org/10.11646/zootaxa.2533.1.1" volume="2533" year="2010">Schmidt-Rhaesa 2010</bibRefCitation>
) and Nearctic realms (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.892.38868" author="Anaya, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="59 - 75" refId="B2" refString="Anaya, C, Schmidt-Rhaesa, A, Hanelt, B, Bolek, MG, 2019. A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America. ZooKeys 892: 59 - 75, DOI: https://doi.org/10.3897/zookeys.892.38868" title="A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America." url="https://doi.org/10.3897/zookeys.892.38868" volume="892" year="2019">Anaya et al. 2019</bibRefCitation>
). In
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov., this dense patch of bristles is a stable characteristic that was detected in all samples. The distribution pattern was similar to that of
<taxonomicName lsidName="G. helveticus" pageId="0" pageNumber="25" rank="species" species="helveticus">
<emphasis italics="true" pageId="0" pageNumber="25">G. helveticus</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2533.1.1" author="Schmidt-Rhaesa, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="25" pagination="1 - 35" refId="B26" refString="Schmidt-Rhaesa, A, 2010. Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species. Zootaxa 2533: 1 - 35, DOI: https://doi.org/10.11646/zootaxa.2533.1.1" title="Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species." url="https://doi.org/10.11646/zootaxa.2533.1.1" volume="2533" year="2010">Schmidt-Rhaesa 2010</bibRefCitation>
) because the bristles exhibited a progressively broader distribution instead of being concentrated along the row of the ventral border, such as in
<taxonomicName lsidName="G. karwendeli" pageId="0" pageNumber="25" rank="species" species="karwendeli">
<emphasis italics="true" pageId="0" pageNumber="25">G. karwendeli</emphasis>
</taxonomicName>
Schmidt-Rhaesa, 2010 (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2533.1.1" author="Schmidt-Rhaesa, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="25" pagination="1 - 35" refId="B26" refString="Schmidt-Rhaesa, A, 2010. Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species. Zootaxa 2533: 1 - 35, DOI: https://doi.org/10.11646/zootaxa.2533.1.1" title="Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species." url="https://doi.org/10.11646/zootaxa.2533.1.1" volume="2533" year="2010">Schmidt-Rhaesa 2010</bibRefCitation>
) and
<taxonomicName lsidName="G. terrestris" pageId="0" pageNumber="25" rank="species" species="terrestris">
<emphasis italics="true" pageId="0" pageNumber="25">G. terrestris</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.892.38868" author="Anaya, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="59 - 75" refId="B2" refString="Anaya, C, Schmidt-Rhaesa, A, Hanelt, B, Bolek, MG, 2019. A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America. ZooKeys 892: 59 - 75, DOI: https://doi.org/10.3897/zookeys.892.38868" title="A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America." url="https://doi.org/10.3897/zookeys.892.38868" volume="892" year="2019">Anaya et al. 2019</bibRefCitation>
), or in a circular patch of concentrated spines, such as in
<taxonomicName lsidName="G. spiridonovi" pageId="0" pageNumber="25" rank="species" species="spiridonovi">
<emphasis italics="true" pageId="0" pageNumber="25">G. spiridonovi</emphasis>
</taxonomicName>
Schmidt-Rhaesa, 2010 (
<bibRefCitation author="Spiridonov, SE" journalOrPublisher="Proceedings of the Zoological Institute USSR Academy of Sciences" pageId="0" pageNumber="25" pagination="97 - 101" refId="B34" refString="Spiridonov, SE, 1984. Two new Nematomorpha species of the family Gordiidae. Proceedings of the Zoological Institute USSR Academy of Sciences 126: 97 - 101" title="Two new Nematomorpha species of the family Gordiidae." volume="126" year="1984">Spiridonov 1984</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
Although the distribution pattern of the bristles is similar to that of
<taxonomicName lsidName="G. helveticus" pageId="0" pageNumber="25" rank="species" species="helveticus">
<emphasis italics="true" pageId="0" pageNumber="25">G. helveticus</emphasis>
</taxonomicName>
,
<emphasis italics="true" pageId="0" pageNumber="25">
G
<taxonomicName lsidName="chiashanus" pageId="0" pageNumber="25" rank="species" species="chiashanus">chiashanus</taxonomicName>
</emphasis>
sp. nov. is morphologically distinct because of the presence of stout bristles on the mid-body, a vertical white stripe on the anterior ventral side, and areoles on the inside wall of the cloacal opening. The vertical white stripe on the anterior ventral side can be easily observed by the naked eye, but it has rarely been mentioned thus far. The presence of a white stripe was previously reported in the terrestrial hairworm,
<taxonomicName lsidName="G. terrestris" pageId="0" pageNumber="25" rank="species" species="terrestris">
<emphasis italics="true" pageId="0" pageNumber="25">G. terrestris</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.892.38868" author="Anaya, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="59 - 75" refId="B2" refString="Anaya, C, Schmidt-Rhaesa, A, Hanelt, B, Bolek, MG, 2019. A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America. ZooKeys 892: 59 - 75, DOI: https://doi.org/10.3897/zookeys.892.38868" title="A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America." url="https://doi.org/10.3897/zookeys.892.38868" volume="892" year="2019">Anaya et al. 2019</bibRefCitation>
), which exhibits a broad white patch; however, the patch is likely to be the intensive aggregation of white spots in
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. The presence of areoles on the inside wall of the cloacal opening has only been reported in an unknown
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1515/9783110272536" author="Schmidt-Rhaesa, A" editor="Schmidt-Rhaesa, A" journalOrPublisher="Gastrotricha, Cycloneuralia and Gnathifera (Vol. 1). Nematomorpha, Priapulida, Kinorhyncha and Loricifera. De Gruyter, Berlin" pageId="0" pageNumber="25" pagination="29 - 145" refId="B27" refString="Schmidt-Rhaesa, A, 2012. Nematomorpha. In: Schmidt-Rhaesa, A, Ed., Handbook of Zoology. Gastrotricha, Cycloneuralia and Gnathifera (Vol. 1). Nematomorpha, Priapulida, Kinorhyncha and Loricifera. De Gruyter, Berlin: 29 - 145, DOI: https://doi.org/10.1515/9783110272536" title="Nematomorpha." url="https://doi.org/10.1515/9783110272536" volumeTitle="Handbook of Zoology." year="2012">Schmidt-Rhaesa 2012</bibRefCitation>
, fig. 3.2.2). Although cloacal openings are usually covered by contamination in many
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
species, as was the case in most of our samples, the areole on the inside wall of cloacal opening might not be a general characteristic of the genus
<taxonomicName class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius</emphasis>
</taxonomicName>
because it is absent in at least some species (e.g.,
<taxonomicName lsidName="G. serratus" pageId="0" pageNumber="25" rank="species" species="serratus">
<emphasis italics="true" pageId="0" pageNumber="25">G. serratus</emphasis>
</taxonomicName>
Schmidt-Rhaesa, 2010,
<taxonomicName lsidName="G. terrestris" pageId="0" pageNumber="25" rank="species" species="terrestris">
<emphasis italics="true" pageId="0" pageNumber="25">G. terrestris</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="G. spiridonovi" pageId="0" pageNumber="25" rank="species" species="spiridonovi">
<emphasis italics="true" pageId="0" pageNumber="25">G. spiridonovi</emphasis>
</taxonomicName>
) (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2533.1.1" author="Schmidt-Rhaesa, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="25" pagination="1 - 35" refId="B26" refString="Schmidt-Rhaesa, A, 2010. Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species. Zootaxa 2533: 1 - 35, DOI: https://doi.org/10.11646/zootaxa.2533.1.1" title="Considerations on the genus Gordius (Nematomorpha, horsehair worms), with the description of seven new species." url="https://doi.org/10.11646/zootaxa.2533.1.1" volume="2533" year="2010">Schmidt-Rhaesa 2010</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3176/eco.2010.1.03" author="Schmidt-Rhaesa, A" journalOrPublisher="Estonian Journal of Ecology" pageId="0" pageNumber="25" pagination="39 - 51" refId="B31" refString="Schmidt-Rhaesa, A, Prous, M, 2010. Records of horsehair worms (Nematomorpha) in Estonia, with description of three new species from the genus Gordius L. Estonian Journal of Ecology 59: 39 - 51, DOI: https://doi.org/10.3176/eco.2010.1.03" title="Records of horsehair worms (Nematomorpha) in Estonia, with description of three new species from the genus Gordius L." url="https://doi.org/10.3176/eco.2010.1.03" volume="59" year="2010">Schmidt-Rhaesa and Prous 2010</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.892.38868" author="Anaya, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="59 - 75" refId="B2" refString="Anaya, C, Schmidt-Rhaesa, A, Hanelt, B, Bolek, MG, 2019. A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America. ZooKeys 892: 59 - 75, DOI: https://doi.org/10.3897/zookeys.892.38868" title="A new species of Gordius (Phylum Nematomorpha) from terrestrial habitats in North America." url="https://doi.org/10.3897/zookeys.892.38868" volume="892" year="2019">Anaya et al. 2019</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.941.49100.figure8" httpUri="https://binary.pensoft.net/fig/421161" pageId="0" pageNumber="25" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 8.</emphasis>
Seasonal occurrence of free-living adults of
<taxonomicName authorityName="Chiu" authorityYear="2020" class="Gordioida" family="Gordiidae" genus="Gordius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gordius chiashanus" order="Gordioidea" pageId="0" pageNumber="25" phylum="Nematomorpha" rank="species" species="chiashanus">
<emphasis italics="true" pageId="0" pageNumber="25">Gordius chiashanus</emphasis>
</taxonomicName>
sp. nov. Numbers at the bottom indicate the actual number of each bar.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>