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<document ID-DOI="http://doi.org/10.5281/zenodo.4762287" ID-GBIF-Dataset="1e9005b3-eebe-4cde-8807-181213fc7c60" ID-GBIF-Taxon="182071331" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762287" checkinTime="1620934442152" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2008" docId="19428793FFDD79140710FE8A6E65AC0B" docLanguage="en" docName="Illiesia.4.1.1-10.pdf" docOrigin="Illiesia 4 (1)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Neoperla peterzwicki Stark &amp; Sivec 2008, sp. n." docType="treatment" docVersion="5" lastPageNumber="3" masterDocId="E57BFFEBFFDF7916062FFF956B15AA52" masterDocTitle="New Stoneflies (Plecoptera) From Asia" masterLastPageNumber="10" masterPageNumber="1" pageId="2" pageNumber="3" updateTime="1632016262587" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Stoneflies (Plecoptera) From Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-01-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>10</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758743</mods:identifier>
<mods:identifier type="GBIF-Dataset">1e9005b3-eebe-4cde-8807-181213fc7c60</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758743</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762287" ID-GBIF-Taxon="182071331" ID-Zenodo-Dep="4762287" LSID="urn:lsid:plazi:treatment:19428793FFDD79140710FE8A6E65AC0B" httpUri="http://treatment.plazi.org/id/19428793FFDD79140710FE8A6E65AC0B" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection box="[319,640,287,312]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph blockId="2.[319,640,287,312]" box="[319,640,287,312]" pageId="2" pageNumber="3">
<heading bold="true" box="[319,640,287,312]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis bold="true" box="[319,640,287,312]" pageId="2" pageNumber="3">
<taxonomicName authority="Stark &amp; Sivec, 2008" authorityName="Stark &amp; Sivec" authorityYear="2008" box="[319,565,287,311]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="peterzwicki" status="sp. n.">
<emphasis bold="true" box="[319,565,287,311]" italics="true" pageId="2" pageNumber="3">Neoperla peterzwicki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[578,640,288,312]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[417,541,322,346]" pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[417,541,322,346]" box="[417,541,322,346]" pageId="2" pageNumber="3">
(
<figureCitation box="[425,533,322,346]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Figs. 5-11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="materials_examined">
<paragraph blockId="2.[184,776,391,1972]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[184,418,392,416]" pageId="2" pageNumber="3">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111463308" collectingDate="1994-04-05" collectingDateMax="1994-04-09" collectingDateMin="1994-04-05" collectionCode="MZB" collectorName="B. Hubley &amp; D. C. Darling" country="Indonesia" elevation="440" latitude="3.1666667" location="Bahan River" longLatPrecision="1307" longitude="115.78333" municipality="Long Tua" pageId="2" pageNumber="3" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="East Kalimantan" typeStatus="holotype">
<typeStatus box="[430,538,392,416]" pageId="2" pageNumber="3">Holotype</typeStatus>
<specimenCount box="[550,569,391,415]" pageId="2" pageNumber="3" type="male"></specimenCount>
and 1 pinned
<specimenCount box="[756,775,391,415]" pageId="2" pageNumber="3" type="female"></specimenCount>
paratype from
<collectingCountry box="[369,484,427,451]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
,
<collectingRegion box="[501,698,427,451]" country="Indonesia" name="Kalimantan Timur" pageId="2" pageNumber="3">East Kalimantan</collectingRegion>
,
<collectingMunicipality pageId="2" pageNumber="3">Long Tua</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:19428793FFDD79140710FE8A6E65AC0B:9434605EFFDD791406DAFE586A92ABB7" box="[245,391,461,485]" country="Indonesia" latitude="3.1666667" longLatPrecision="1307" longitude="115.78333" municipality="Long Tua" name="Bahan River" pageId="2" pageNumber="3" stateProvince="East Kalimantan">Bahan River</location>
,
<geoCoordinate box="[405,504,461,485]" degrees="3" direction="north" minutes="10" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="3.1666667">3° 10 N</geoCoordinate>
,
<geoCoordinate box="[519,639,461,485]" degrees="115" direction="east" minutes="47" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="115.78333">115° 47 E</geoCoordinate>
,
<quantity box="[653,727,461,485]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="3" unit="m" value="440.0">
<elevation box="[653,727,461,485]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="3" unit="m" value="440.0">440 m</elevation>
</quantity>
,
<date pageId="2" pageNumber="3" value="1994-04-05" valueMax="1994-04-09" valueMin="1994-04-05">
<collectingDate pageId="2" pageNumber="3" value="1994-04-05" valueMax="1994-04-09" valueMin="1994-04-05">5-9 April 1994</collectingDate>
</date>
, IIS 940526,
<collectorName box="[478,603,496,520]" pageId="2" pageNumber="3">B. Hubley</collectorName>
,
<collectorName box="[622,776,496,520]" pageId="2" pageNumber="3">D.C. Darling</collectorName>
(
<collectionCode box="[192,254,530,554]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="3" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111463306" collectingDate="1993-02-20" collectingDateMax="1993-03-04" collectingDateMin="1993-02-20" collectionCode="ROM" collectorName="D. C. Darling" country="Indonesia" county="Balungan" elevation="355" latitude="2.85" location="Lalut Birai Research Station" longLatPrecision="1307" longitude="115.8" municipality="Kayan Mentarang Nature Reserve" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="East Kalimantan" typeStatus="paratype">
<typeStatus box="[286,402,531,555]" pageId="2" pageNumber="3">Paratypes</typeStatus>
:
<collectingCountry box="[427,543,530,554]" name="Indonesia" pageId="2" pageNumber="3">Indonesia</collectingCountry>
:
<collectingRegion box="[566,771,530,554]" country="Indonesia" name="Kalimantan Timur" pageId="2" pageNumber="3">East Kalimantan</collectingRegion>
,
<collectingCounty box="[184,295,565,589]" pageId="2" pageNumber="3">Balungan</collectingCounty>
,
<collectingMunicipality box="[307,704,565,589]" pageId="2" pageNumber="3">Kayan Mentarang Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:19428793FFDD79140710FE8A6E65AC0B:9434605EFFDD791404E4FDA06AA4A83D" country="Indonesia" county="Balungan" latitude="2.85" longLatPrecision="1307" longitude="115.8" municipality="Kayan Mentarang Nature Reserve" name="Lalut Birai Research Station" pageId="2" pageNumber="3" stateProvince="East Kalimantan">Lalut Birai Research Station</location>
,
<geoCoordinate box="[442,552,599,624]" degrees="115" direction="east" minutes="48" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="115.8">115° 48 E</geoCoordinate>
,
<geoCoordinate box="[561,663,599,624]" degrees="02" direction="north" minutes="51" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="2.85">02° 51 N</geoCoordinate>
,
<quantity box="[672,741,599,623]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="2" pageNumber="3" unit="m" value="355.0">
<elevation box="[672,741,599,623]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="2" pageNumber="3" unit="m" value="355.0">355 m</elevation>
</quantity>
,
<date pageId="2" pageNumber="3" value="1993-02-20" valueMax="1993-03-04" valueMin="1993-02-20">
<collectingDate pageId="2" pageNumber="3" value="1993-02-20" valueMax="1993-03-04" valueMin="1993-02-20">20 February-4 March 1993</collectingDate>
</date>
, IIS 930000,
<collectorName box="[582,729,634,658]" pageId="2" pageNumber="3">D.C. Darling</collectorName>
,
<specimenCount box="[738,775,633,658]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
pinned (
<collectionCode box="[279,346,669,693]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="2" pageNumber="3" type="Museum">ROM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[184,776,391,1972]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[184,354,703,727]" pageId="2" pageNumber="3">Adult habitus.</emphasis>
General color pale brown. Head pale brown but with obscure darker spot between ocelli; lappets and triangular area forward of M-line brown; dusky area and small pale spot define obscure M-line (
<figureCitation box="[192,261,841,865]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Basal two antennal segments pale brown, flagellum dark brown; basal palpal segments pale, apical segments brown. Pronotum brown with slightly darker rugosities. Wing membrane pale amber, veins brown. Femora pale brown, darker on dorsum and at narrow knee band; tibiae and tarsi dark brown.
</paragraph>
<paragraph blockId="2.[184,776,391,1972]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[184,251,1084,1108]" pageId="2" pageNumber="3">Male.</emphasis>
Forewing length
<quantity box="[455,537,1084,1108]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="3" unit="mm" value="13.0">13 mm</quantity>
. Process of tergum 7 wide and notched; apex lined with a few large sensilla basiconica. Median sclerite of tergum 8 slightly raised, weakly sclerotized and bearing a pair of sensilla basiconica patches. Median field of tergum 9 bearing a boomerang shaped mound, slightly sclerotized at each end, and bearing a few sensilla basiconica. Hemitergal tips turned abruptly inwards, posterolateral margins bearing dense row of short stout bristles and a lateral notch; finger lobes slender, relatively long, directed strongly mesad and curved slightly (
<figureCitation box="[288,356,1463,1488]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
). Aedeagal tube poorly sclerotized, plump and sparsely armed with small scale-like spines below and forward of bulb and in a narrow mid-dorsal band; apex of tube armed with a pair of dorsolateral spiny lobes. Everted aedeagal sac with large basoventral spiny lobe, large cushion-like basodorsal spiny lobe and a pair of small lateral spiny lobes at midlength; apex of sac bearing a subapical ring of large spines and, near bases of lateral lobes on dorsal margin, a pair of small patches of ca. 5-6 large spines; much of sac surface covered with fine spinules grading into small spines (
<figureCitation captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Figs. 7- 8</figureCitation>
).
</paragraph>
<paragraph blockId="2.[184,776,391,1972]" lastBlockId="2.[812,1405,288,1625]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[184,276,1913,1937]" pageId="2" pageNumber="3">Female.</emphasis>
Forewing length
<quantity box="[488,573,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="3" unit="mm" value="17.0">17 mm</quantity>
. Subgenital plate scarcely produced, posterior margin of sternum 8 almost straight but with two widely separated small projecting points on either side of a slightly scalloped area (
<figureCitation box="[873,935,356,381]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
). Vagina approximately circular with tiers of internalwrinkles; mostofvaginal areawithbrown lining; spermatheca J-shaped without apical hook (
<figureCitation box="[820,899,460,484]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
).
</paragraph>
<paragraph blockId="2.[812,1405,288,1625]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[812,863,495,519]" pageId="2" pageNumber="3">Egg.</emphasis>
Outline oval, collar short but distinctly stalked; rim of collar flanged and emarginate (
<figureCitation box="[1303,1391,530,554]" captionStart="Figs" captionStartId="3.[184,234,1566,1590]" captionTargetBox="[379,1222,288,1469]" captionTargetId="figure-157@3.[352,1235,279,1462]" captionTargetPageId="3" captionText="Figs. 5-11. Neoperla peterzwicki. 5. Head and pronotum, 6. Male abdominal terga 7-9, 7. Aedeagus lateral, 8. Aedeagus dorsal, 9. Female abdominal sterna 8-9, 10. Vagina, 11. Egg." figureDoi="http://doi.org/10.5281/zenodo.4758747" httpUri="https://zenodo.org/record/4758747/files/figure.png" pageId="2" pageNumber="3">Fig. 11</figureCitation>
). Chorion finely punctate throughout; equatorial region with obscure follicle cell impressions enclosing punctations.
</paragraph>
<paragraph blockId="2.[812,1405,288,1625]" box="[812,1014,668,692]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[812,885,668,692]" pageId="2" pageNumber="3">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="etymology">
<paragraph blockId="2.[812,1405,288,1625]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[812,946,703,727]" pageId="2" pageNumber="3">Etymology.</emphasis>
The patronym honors our friend and colleague Professor Dr. Peter Zwick in recognition of his major contributions to the systematics of
<taxonomicName authorityName="Needham" authorityYear="1905" box="[1311,1403,772,796]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[1311,1403,772,796]" italics="true" pageId="2" pageNumber="3">Neoperla</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation author="Zwick, P." box="[866,996,807,831]" pageId="2" pageNumber="3" pagination="263 - 269" refId="ref4452" refString="Zwick, P. 1980. The genus Neoperla (Plecoptera: Perlidae) from Sri Lanka. Oriental Insects, 14: 263 - 269." type="journal article" year="1980">Zwick 1980</bibRefCitation>
,
<bibRefCitation author="Zwick, P." box="[1005,1059,807,831]" pageId="2" pageNumber="3" pagination="113 - 126" refId="ref4479" refString="Zwick, P. 1981. The South Indian species of Neoperla (Plecoptera: Perlidae). Oriental Insects, 15: 113 - 126." type="journal article" year="1981">1981</bibRefCitation>
,
<bibRefCitation author="Zwick, P." box="[1068,1122,807,831]" pageId="2" pageNumber="3" pagination="167 - 204" refId="ref4506" refString="Zwick, P. 1983. The Neoperla of Sumatra and Java (Indonesia) (Plecoptera: Perlidae). Spixiana, 6: 167 - 204." type="journal article" year="1983">1983</bibRefCitation>
,
<bibRefCitation author="Zwick, P." box="[1131,1184,807,831]" pageId="2" pageNumber="3" pagination="393 - 407" refId="ref4594" refString="Zwick, P. 1988. Species of Neoperla from the South- East Asian mainland (Plecoptera: Perlidae). Entomologica Scandinavica, 18: 393 - 407." type="journal article" year="1988">1988</bibRefCitation>
) and in the study of Bornean stoneflies (
<bibRefCitation author="Zwick, P." box="[1036,1172,841,865]" pageId="2" pageNumber="3" pagination="1 - 53" refId="ref4565" refString="Zwick, P. 1986. The Bornean species of the stonefly genus Neoperla (Plecoptera: Perlidae). Aquatic Insects, 8: 1 - 53." type="journal article" year="1986">Zwick 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="diagnosis">
<paragraph blockId="2.[812,1405,288,1625]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[812,1129,876,900]" pageId="2" pageNumber="3">Diagnosis and Discussion.</emphasis>
<taxonomicName authorityName="Stark &amp; Sivec" authorityYear="2008" box="[1136,1356,876,900]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="peterzwicki">
<emphasis box="[1136,1356,876,900]" italics="true" pageId="2" pageNumber="3">Neoperla peterzwicki</emphasis>
</taxonomicName>
is a member of the
<emphasis box="[1029,1156,911,934]" italics="true" pageId="2" pageNumber="3">“Borneensis</emphasis>
Subgroup” of the
<emphasis box="[812,927,946,969]" italics="true" pageId="2" pageNumber="3">Montivaga</emphasis>
Group of
<emphasis box="[1051,1152,945,969]" italics="true" pageId="2" pageNumber="3">
<taxonomicName authorityName="Needham" authorityYear="1905" box="[1051,1148,945,969]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Neoperla</taxonomicName>
.
</emphasis>
As defined by
<bibRefCitation author="Zwick, P." pageId="2" pageNumber="3" pagination="1 - 53" refId="ref4565" refString="Zwick, P. 1986. The Bornean species of the stonefly genus Neoperla (Plecoptera: Perlidae). Aquatic Insects, 8: 1 - 53." type="journal article" year="1986">Zwick (1986)</bibRefCitation>
, males of the subgroup have a notched or bifurcate process of tergum 7 and the hemiterga have relatively wide bases. The subgroup is presently known only from Borneo and includes seven species or subspecies recognized and described by
<bibRefCitation author="Zwick, P." pageId="2" pageNumber="3" pagination="1 - 53" refId="ref4565" refString="Zwick, P. 1986. The Bornean species of the stonefly genus Neoperla (Plecoptera: Perlidae). Aquatic Insects, 8: 1 - 53." type="journal article" year="1986">Zwick (1986)</bibRefCitation>
. Externally the new species is quite similar to
<taxonomicName authority="Navas" authorityName="Navas" authorityYear="1929" box="[812,1006,1187,1211]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="harina">
<emphasis box="[812,919,1187,1211]" italics="true" pageId="2" pageNumber="3">N. harina</emphasis>
Navas
</taxonomicName>
,
<taxonomicName authority="Klapalek" authorityName="Klapalek" authorityYear="1909" box="[1023,1266,1187,1211]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="furcifera">
<emphasis box="[1023,1150,1187,1211]" italics="true" pageId="2" pageNumber="3">N. furcifera</emphasis>
Klapálek
</taxonomicName>
,
<taxonomicName authority="Zwick" authorityName="Zwick" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bilobata">
<emphasis box="[1283,1403,1187,1211]" italics="true" pageId="2" pageNumber="3">N. bilobata</emphasis>
Zwick
</taxonomicName>
and
<taxonomicName authority="Zwick" authorityName="Zwick" box="[940,1125,1221,1246]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="furcata">
<emphasis box="[940,1047,1222,1246]" italics="true" pageId="2" pageNumber="3">N. furcata</emphasis>
Zwick
</taxonomicName>
and less similar to other members of the complex. The aedeagal lobing and armature are suggestive of
<taxonomicName box="[1124,1239,1290,1314]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bilobata">
<emphasis box="[1124,1239,1290,1314]" italics="true" pageId="2" pageNumber="3">N. bilobata</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Navas" authorityYear="1929" box="[1296,1396,1290,1314]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="harina">
<emphasis box="[1296,1396,1290,1314]" italics="true" pageId="2" pageNumber="3">N. harina</emphasis>
</taxonomicName>
; from the latter, the new species differs in having a much more prominent dorsobasal spiny cushion and in having small spines scattered over most of the sac surface; from the former species it differs in having a larger ventrobasal spiny lobe and in having the largest spines in the apical region arranged in a circumlinear subapical patch rather than along the dorsal margin in a more linear arrangement. The following key is offered for males of the subgroup.
</paragraph>
</subSubSection>
</treatment>
</document>