treatments-xml/data/DE/34/C9/DE34C90D9A425CC69E158B25667916DA.xml

<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="DE34C90D9A425CC69E158B25667916DA" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) brunneus subsp. brunneus brunneus Karsch 1881" docType="treatment" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Machard, Patrice</mods:namePart>
<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2022</mods:date>
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15.1. 
<taxonomicName LSID="DE34C90D-9A42-5CC6-9E15-8B25667916DA" authority="brunneus Karsch, 1881" authorityName="brunneus Karsch" authorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="Eremoderus" subSpecies="brunneus">Platyderus (Eremoderus) brunneus brunneus Karsch, 1881</taxonomicName>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3D, E</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche &amp; Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Urite, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, E: topotype male specimen; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) vrabeci, sp. nov., holotype; E. P. (E.) lassallei, sp. nov., holotype; F. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; G. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; H. P. (E.) afghanistanicus, sp. nov., holotype; I. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; J. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; K. P. (E.) jordanensis, sp. nov., holotype; L. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel; N. P. (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; O. P. (P.) reticulatus (Chaudoir), male specimen, Tashehzeh, Nowshahr County, Mazandaran Province, Iran. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure7" httpUri="https://binary.pensoft.net/fig/695644" pageId="0" pageNumber="71">, 7I</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9C</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12J</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15A</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="reference_group">
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Karsch, 1881: 43 (type locality: &quot;Bir Milrha&quot;, Jabal 
<normalizedToken originalValue="Tarhūnah">Tarhūnah</normalizedToken>
[Tarhuna plateau], Libya, based on lectotype designation), stat. rev.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
= 
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
Bedel, 1900: 170 (type locality: &quot;Sud de la Tunisie&quot;), syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
This taxon was described from &quot;Bir Milrha&quot; (Murqub District, Libya; locality cannot be localized) and &quot;Uadi Mbellem&quot; (Misrata District, Libya; locality with approximate GPS coordinates: 31.167, 15.050) both localities in the 
<normalizedToken originalValue="Gebèl">Gebel</normalizedToken>
<normalizedToken originalValue="Tarhúna">Tarhuna</normalizedToken>
Plateaux. As a result of current lectotype designation, the former place becomes the type locality of 
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
</taxonomicName>
was first discovered from &quot;Sud de la Tunisie&quot; (Bedel, 1900), and for the second time - from 
<normalizedToken originalValue="“Aïn">&quot;Ain</normalizedToken>
Segoufta 
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken originalValue="l’O">l'O</normalizedToken>
. du Dj. Bon-Hedma&quot; (Bedel, 1902); the second location refers to present Jebel Bou-Hedma National Park, situated on Gafsa Governorate and Sidi Bouzid Governorate, Tunisia.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 14.</emphasis>
Spermathecal complex and gonocoxites, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="Eremoderus">Platyderus (Eremoderus) lassallei</taxonomicName>
, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; 
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) ledouxi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="ledouxi" subGenus="E.">P. (E.) ledouxi</taxonomicName>
Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; 
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; 
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (Platyderus) umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus" subGenus="Platyderus">P. (Platyderus) umbratus</taxonomicName>
(
<normalizedToken originalValue="Ménétriés">Menetries</normalizedToken>
), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; 
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" subGenus="E.">P. (E.) arabicus</taxonomicName>
, sp. nov., holotype; 
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" subGenus="E.">P. (E.) brunki</taxonomicName>
, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13</figureCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="references">
<paragraph pageId="0" pageNumber="71">References.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus elegans</emphasis>
</taxonomicName>
: 
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 211, 214-215; 
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 768; 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B33" refString="Lorenz, W, 1998. Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." title="Nomina Carabidarum - a directory of the scientific names of ground beetles (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). First edition. Wolfgang Lorenz, Tutzing, [ii +] 502 pp." year="1998">Lorenz 1998</bibRefCitation>
: 375; 
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 522; 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 395; 
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 757. 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
(data for 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus brunneus</emphasis>
</taxonomicName>
Karsch): 
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel 1902</bibRefCitation>
: 211, 214-215; 
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II. W. Junk, Berlin" pageId="0" pageNumber="71" pagination="739 - 1022" refId="B16" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II. W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
: 769; 
<bibRefCitation author="Hovorka, O" editor="Loebl, L" journalOrPublisher="Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" pageId="0" pageNumber="71" pagination="521 - 524" refId="B22" refString="Hovorka, O, Sciaky, R, 2003. Atranopsina Baehr, 1982. In: Loebl, L, Smetana, A, Eds., Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup: 521 - 524" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2003">Hovorka and Sciaky 2003</bibRefCitation>
: 521; 
<bibRefCitation author="Lorenz, W" journalOrPublisher="Acta Societatis Zoologicae Bohemicae" pageId="0" pageNumber="71" refId="B34" refString="Lorenz, W, 2005. Systematic list of extant ground beetles of the world (InsectaColeoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." title="Systematic list of extant ground beetles of the world (InsectaColeoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Wolfgang Lorenz, Tutzing, [ii +] 530 pp." year="2005">Lorenz 2005</bibRefCitation>
: 396; 
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka 2017</bibRefCitation>
: 756.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
</taxonomicName>
Karsch, 1881. The original type series of Karsch consisted of six syntypes, 4♂♂, 2♀♀, all stored in MFNB. Their examination revealed the presence of two diagnosable taxa. One male and one female specimen were found to represent the subspecies 
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
(see section &quot;Type material&quot; under 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
). The other four specimens belong to 
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
, the first of them designated lectotype and other three paralectotypes: ♂, '60906 [p, w] // Bir Milrha / Exp. Rohlfs. [h, tq] // Hist.-Coll. (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 / 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha / Zool. Mus. Berlin [p, w] // SYNTYPE / 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♂, 'Uadi 
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
/ Exp. Rohlfs. [h, tq] // Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 / 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Uadi 
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
, Exped. / Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE / 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♂, 'gn 2469. [h, w] // Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 / 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha &amp; Uadi 
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
, / Exped. Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE / 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'; ♀, 'Hist.-Coll. (
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
) / Nr. 60906 / 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">Platyderus brunneus</taxonomicName>
Klug, / Karsch* / Bir Milrha &amp; Uadi 
<normalizedToken originalValue="M’bellem">M'bellem</normalizedToken>
, / Exped. Rohlfs / Zool. Mus. Berlin [p, w] // SYNTYPE / 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus" subGenus="Eremoderus">Platyderus Eremoderus brunneus</taxonomicName>
Karsch, 1881 / labelled by MFNB 2017 [p, r]'.
</paragraph>
<paragraph pageId="0" pageNumber="71">
The synonymy of 
<taxonomicName authorityName="Karsch" authorityYear="1881" baseAuthorityName="brunneus Karsch" baseAuthorityYear="1881" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus</emphasis>
</taxonomicName>
with 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
was proposed by 
<bibRefCitation author="Bedel, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="71" refId="B11" refString="Bedel, L, 1902. Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209-220." title="Catalogue raisonne des Coleopteres du nord de l'Afrique (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 209 - 220." year="1902">Bedel (1902</bibRefCitation>
: 211). Revision of the type material and study of the original descriptions of the both aforementioned forms and that of 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
</taxonomicName>
, as well the study of other 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-specimens from North Africa and the Levant, revealed that 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
</taxonomicName>
represent a single species in contrast with 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
which is a separate species not occurring in North Africa.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Tunisia</emphasis>
: 
<materialsCitation collectingDate="1998-04-22" collectorName="R. Borovec, Bedel" country="TUNISIA" location="Ben Gardane" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Medenine Governorate">
<collectingRegion country="Tunisia" name="Medenine">Medenine Governorate</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
, ' 
<collectingCountry name="Tunisia">TUNISIA</collectingCountry>
c. or. / 
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
W of 
<location LSID="urn:lsid:plazi:treatment:DE34C90D9A425CC69E158B25667916DA:E650DDF8528D0FB64660AEC58F557611" country="TUNISIA" name="Ben Gardane" stateProvince="Medenine Governorate">Ben Gardane</location>
/ 
<collectingDate value="1998-04-22">22.4.1998</collectingDate>
/ 
<collectorName>R. Borovec</collectorName>
lgt. // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName authorityName="Bedel" authorityYear="1900" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">Platyderus elegans</taxonomicName>
<collectorName>Bedel</collectorName>
/ JEANNE det. 2003 // COLL. WRASE / 
<normalizedToken originalValue="BERLIN’">BERLIN'</normalizedToken>
(cWR)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="2006-02-04" collectorName="Weill'" country="Libya" location="Nuqat al Khams District" specimenCount="1" specimenCount-female="1" stateProvince="Nuqat al Khams District">
<emphasis bold="true" pageId="0" pageNumber="71">
<collectingCountry name="Libya">Libya</collectingCountry>
</emphasis>
: 
<collectingRegion country="Libya" name="An Nuqat al Khams">Nuqat al Khams District</collectingRegion>
: 
<specimenCount type="female">1♀</specimenCount>
, 'BUKAMASH [near Zuwarah City] / 
<collectingDate value="2006-02-04">04-II-2006</collectingDate>
/ LIBYA 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<normalizedToken originalValue="Weill’">
<collectorName>Weill'</collectorName>
</normalizedToken>
(cWEI)
</materialsCitation>
. 
<materialsCitation collectingDate="2010-01-30" collectorName="Det., Weill" country="Libya" location="Jafara District" specimenCount="1" specimenCount-female="1" stateProvince="Jafara District">
<collectingRegion inferredBySuffix="true">Jafara District</collectingRegion>
: 
<specimenCount type="female">1♀</specimenCount>
, 'AZIZIYAH 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
Weill / 
<collectingDate value="2010-01-30">30-I-2010</collectingDate>
LIBYE // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
. R &amp; S / 
<collectorName>Det.</collectorName>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P." order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">P.</taxonomicName>
<collectorName>Weill</collectorName>
2010' (cWEI)
</materialsCitation>
. 
<materialsCitation collectingDate="2002-03-02" collectorName="JC Ringenbach Leg." country="Libya" location="Tripoli District" specimenCount="1" specimenCount-male="1" stateProvince="Tripoli District">
<collectingRegion country="Libya" name="Tarabulus">Tripoli District</collectingRegion>
: 
<specimenCount type="male">1♂</specimenCount>
, ' 
<collectingDate value="2002-03-02">02-III-2002</collectingDate>
/ JANZUR / LIBYA - TRIPOLITANIA // 
<collectorName>JC Ringenbach Leg.</collectorName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus languidus</emphasis>
</taxonomicName>
' (cWEI)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
Three additional records from 
<collectingCountry name="Libya">Libya</collectingCountry>
that concern 
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
are pointed out for 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
in the webpage http://jcringenbach.free.fr/website/beetles/carabidae/Platyderus_languidus.htm, namely: 'East of 
<normalizedToken originalValue="Tajura’">Tajura'</normalizedToken>
, 
<date value="2002-01-04">04/01/2002</date>
, JCR Leg.'; 'Road Yafran-Az Zintan, 
<date value="2001-12-25">25/12/2001</date>
, JCR Leg.'; 'Sidi As 
<normalizedToken originalValue="Sa’ih">Sa'ih</normalizedToken>
<date value="2006-02-03">03/02/2006</date>
, PW Leg.'. The webpage is part of the database http://jcringenbach.free.fr/website/beetles/carabidae/carablibya.htm created by Jean-Claude Ringenbach (
<normalizedToken originalValue="Pardies-Piétat">Pardies-Pietat</normalizedToken>
, 
<collectingCountry name="France">France</collectingCountry>
) and Patrick Weill (Pau, 
<collectingCountry name="France">France</collectingCountry>
) and representing online checklist of Libyan ground-beetles.
</paragraph>
<paragraph pageId="0" pageNumber="71">
TME: 
<specimenCount type="generic">9 specimens</specimenCount>
. TGE: 
<specimenCount type="male">2♂♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
Nominotypical subspecies differs from ssp. 
<taxonomicName authorityName="Reitter" authorityYear="1909" baseAuthorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" baseAuthorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunneus ferrantei</emphasis>
</taxonomicName>
in parascutellar striole and elytral striae 1-7 very shallow, almost indistinct, with bases not reaching basal bead (vs. parascutellar striole and elytral striae 1-7 distinct, slightly impressed, with bases reaching basal bead). In addition, the former has the pronotal anterior margin proportionally broader, compared to the posterior margin (vs. anterior margin of pronotum proportionally narrower, compared to posterior margin, see PA/PB, Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">Two of the three Libyan specimens were collected under stones or plant debris. Bukamash and Aziziyah are two biotopes of Tripolitania located in a semi-desert zone (Jaffarah Plain), with a few sparse trees and abundant ground vegetation, mainly in early spring (Patrick Weill, pers. comm.).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
Tunisia, Libya (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of species from subgenus Platyderus Eremoderus in continental North Africa and the southern Levant: Platyderus brunneus brunneus (blue circles), P. brunneus ferrantei (yellow circles), P. insignitus insignitus (red circles), and P. insignitus presaharensis (brown circle)." figureDoi="10.3897/dez.69.83840.figure18" httpUri="https://binary.pensoft.net/fig/695655" pageId="0" pageNumber="71">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
<bibRefCitation author="Hovorka, O" editor="Loebl, I" journalOrPublisher="Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston" pageId="0" pageNumber="71" pagination="755 - 760" refId="B21" refString="Hovorka, O, 2017. Atranopsina Baehr, 1982. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera. Volume 1 Revised and Updated Edition. Archostemata - Myxophaga - Adephaga. Brill, Leiden - Boston: 755 - 760" title="Atranopsina Baehr, 1982." volumeTitle="Catalogue of Palaearctic Coleoptera." year="2017">Hovorka (2017</bibRefCitation>
: 757) placed 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. elegans" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="71">P. elegans</emphasis>
</taxonomicName>
in 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
(s. str.). Present study proved however that the species currently validly known as 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus P. brunneus</emphasis>
</taxonomicName>
is indeed a member of subgenus 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="(Eremoderus)" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subGenus" subGenus="Eremoderus">Eremoderus</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>