treatments-xml/data/CA/10/36/CA103649E06BDC46E71595960D0B34BC.xml

<document id="02F2FADC952B7B174CEE296E217E54C7" ID-DOI="10.11646/zootaxa.1095.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5051358" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625109620475" checkinUser="felipe" docAuthor="Macpherson, Enrique &amp; Segonzac, Michel" docDate="2005" docId="CA103649E06BDC46E71595960D0B34BC" docLanguage="en" docName="zt01095p060.pdf" docOrigin="Zootaxa 1095 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1095.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Munidopsis exuta Macpherson &amp; Segonzac 2005, n. sp." docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="36294E31E07EDC5EE61D97420F42323A" masterDocTitle="Species of the genus Munidopsis (Crustacea, Decapoda, Galatheidae) from the deep Atlantic Ocean, including cold-seep and hydrothermal vent areas" masterLastPageNumber="60" masterPageNumber="1" pageNumber="22" updateTime="1698981493787" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9F9DBDF4D7C673076578A7DA9C1EBC12">Species of the genus Munidopsis (Crustacea, Decapoda, Galatheidae) from the deep Atlantic Ocean, including cold-seep and hydrothermal vent areas</mods:title>
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<mods:namePart id="2FF58FF7F89F67C9A6362518E87F5A69">Macpherson, Enrique</mods:namePart>
<mods:affiliation id="F9076F26234F96D8E087ECB30B53A1A9">Centro de Estudios Avanzados de Blanes (CSIC), C. acc. Cala San Francesc 14, 17300 Blanes, Spain (email: macpherson @ ceab. csic. es). Ifremer, Centre de Brest, DEEP / Laboratoire Environnement Profond, BP 70, 29280 Plouzané, France (email: segonzac @ ifremer. fr).</mods:affiliation>
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<mods:date id="67FCFAC549830C028BC9334E320E838F">2005</mods:date>
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<taxonomicName id="85B9FCDCE06BDC4BE71595960E9D30D4" authority="Macpherson &amp; Segonzac, 2005" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[264,479,724,750]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="exuta" status="sp. nov.">
<emphasis id="70CD5B4DE06BDC4BE71595960E9D30D4" bold="true" box="[264,479,724,750]" italics="true" pageId="21" pageNumber="22">Munidopsis exuta</emphasis>
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<taxonomicNameLabel id="6BFEE636E06BDC4BE7FA95960D6B30D4" box="[487,553,724,750]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="DA829BDAE06BDC4BE42595960DC230D4" box="[568,640,724,750]" captionStart="FIGURE 5" captionStartId="22.[264,357,1719,1740]" captionTargetBox="[295,1315,288,1699]" captionTargetId="figure-99@22.[264,1323,276,1723]" captionTargetPageId="22" captionText="FIGURE 5. Munidopsis exuta n. sp., holotype, ovigerous female (34.7 mm), Mid­Atlantic Ridge, HYDROSNAKE, Stn HS08, MNHN­Ga 4621. A, carapace, dorsal. B, same, lateral. C, posterior part of sixth abdominal segment and telson. D, anterior part of sternal plastron. E, left antennule, antenna and ocular peduncle, ventral. F, endopod of right third maxilliped, lateral. G, right cheliped, lateral. H, right second pereiopod, lateral. I, dactylus of right second pereiopod, lateral. Scale: A–B, D–E, G. I = 5 mm, C, F, H = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5051372" httpUri="https://zenodo.org/record/5051372/files/figure.png" pageId="21" pageNumber="22">Fig. 5</figureCitation>
)
</emphasis>
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<taxonomicName id="85B9FCDCE06BDC4BE715945D0D59310C" authorityName="Henderson" authorityYear="1885" box="[264,539,799,822]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="70CD5B4DE06BDC4BE715945D0D59310C" box="[264,539,799,822]" italics="true" pageId="21" pageNumber="22">Munidopsis subsquamosa</emphasis>
</taxonomicName>
.—de Saint­Laurent, 1985: 475 (not 
<taxonomicName id="85B9FCDCE06BDC4BE582945D0A52310F" authority="Henderson, 1885" authorityName="Henderson" authorityYear="1885" box="[927,1296,798,822]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subsquamosa">
<emphasis id="70CD5B4DE06BDC4BE582945D0B11310C" box="[927,1107,799,822]" italics="true" pageId="21" pageNumber="22">M. subsquamosa</emphasis>
<bibRefCitation id="2628FAAEE06BDC4BE244945C0A52310F" author="Henderson, J. R." box="[1113,1296,798,821]" pageId="21" pageNumber="22" pagination="407 - 421" refId="ref29177" refString="Henderson, J. R. (1885) Diagnoses of the new species of Galatheidea collected during the &quot; Challenger &quot; Expedition. Annals and Magazine of natural History, (5) 16, 407 - 421." type="journal article" year="1885">Henderson, 1885</bibRefCitation>
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) 
<taxonomicName id="85B9FCDCE06BDC4BE715947D0E93316C" authorityName="Segonzac" authorityYear="1992" box="[264,465,831,854]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="70CD5B4DE06BDC4BE715947D0E93316C" box="[264,465,831,854]" italics="true" pageId="21" pageNumber="22">Munidopsis crassa</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="C318A14EE06BDC4BE7EC947C0D93316F" author="Segonzac, M." box="[497,721,830,853]" page="596" pageId="21" pageNumber="22" year="1992">
<bibRefCitation id="2628FAAEE06BDC4BE7EC947C0D93316F" author="Segonzac, M." box="[497,721,830,853]" pageId="21" pageNumber="22" pagination="593 - 600" refId="ref31246" refString="Segonzac, M. (1992) Les peuplements associes a l'hydrothermalisme oceanique du Snake Pit (dorsale medio-atlantique; 23 ºN, 3480 m): composition et microdistribution de la megafaune. Comptes Rendus de l'Academie des Sciences de Paris, Serie III, 314, 593 - 600." type="journal article" year="1992">Segonzac, 1992: 596</bibRefCitation>
</treatmentCitation>
(not 
<taxonomicName id="85B9FCDCE06BDC4BE514947D0CBD316F" authority="Smith, 1885" authorityName="Smith" authorityYear="1885" box="[777,1023,830,854]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="70CD5B4DE06BDC4BE514947D0C31316C" box="[777,883,831,854]" italics="true" pageId="21" pageNumber="22">M. crassa</emphasis>
<bibRefCitation id="2628FAAEE06BDC4BE564947C0CBD316F" author="Smith, S. I." box="[889,1023,830,853]" pageId="21" pageNumber="22" pagination="493 - 511" refId="ref31591" refString="Smith, S. I. (1885) On some new or little known decapod Crustacea, from recent Fish Commission dredging off the east coast of the United States. Proceedings of the United States National Museum, 7, 493 - 511." type="journal article" year="1885">Smith, 1885</bibRefCitation>
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).
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</paragraph>
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<emphasis id="70CD5B4DE06BDC4BE71594C40EB4319A" bold="true" box="[264,502,902,928]" pageId="21" pageNumber="22">Material examined</emphasis>
. 
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<collectingRegion id="807D49BDE06BDC4BE41594C50D75319B" box="[520,567,903,929]" country="Somalia" name="Bay" pageId="21" pageNumber="22">Bay</collectingRegion>
of 
<collectingMunicipality id="A2621D25E06BDC4BE47E94C50DF6319B" box="[611,692,903,929]" pageId="21" pageNumber="22">Biscay</collectingMunicipality>
, BIOGAS 5, 
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<specimenCode id="121F2F24E06BDC4BE59A94C50C92319B" box="[903,976,903,929]" collectionCode="CV" country="China" name="Municipal Museum of Chungking" pageId="21" pageNumber="22">CV40</specimenCode>
, 
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, 
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, 
<quantity id="85412ABAE06BDC4BE2D594C50A67319B" box="[1224,1317,903,929]" metricMagnitude="3" metricUnit="m" metricValue="2.86" pageId="21" pageNumber="22" unit="m" value="2860.0">
<elevation id="C994606CE06BDC4BE2D594C50A67319B" box="[1224,1317,903,929]" metricMagnitude="3" metricUnit="m" metricValue="2.86" pageId="21" pageNumber="22" unit="m" value="2860.0">2860 m</elevation>
</quantity>
, 
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<collectingDate id="26435877E06BDC4BE71594ED0ED231F3" box="[264,400,943,969]" pageId="21" pageNumber="22" value="1974-06-15">15.06.1974</collectingDate>
</date>
: 1 ovig. female 
<quantity id="85412ABAE06BDC4BE44F94ED0DFC31F3" box="[594,702,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="21" pageNumber="22" unit="mm" value="29.2">29.2 mm</quantity>
, 
<specimenCount id="54BF4CD6E06BDC4BE4D194ED0C7631F3" box="[716,820,943,969]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
<quantity id="85412ABAE06BDC4BE52094ED0CE931F3" box="[829,939,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="21" pageNumber="22" unit="mm" value="18.2">18.2 mm</quantity>
</materialsCitation>
. 
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<collectingRegion id="807D49BDE06BDC4BE5AA94ED0CA431F3" box="[951,998,943,969]" country="Somalia" name="Bay" pageId="21" pageNumber="22">Bay</collectingRegion>
of 
<location id="4766D184E06BDC4BE20C94ED0B2031F3" LSID="urn:lsid:plazi:treatment:CA103649E06BDC46E71595960D0B34BC:4766D184E06BDC4BE20C94ED0B2031F3" box="[1041,1122,943,969]" latitude="26.133333" longLatPrecision="1243" longitude="-44.816666" name="Biscay" pageId="21" pageNumber="22" stateProvince="Bay">Biscay</location>
, BIOGAS 6, 
<collectorName id="EF4CE289E06BDC4BE31994ED0E6D31CB" pageId="21" pageNumber="22">Stn CP</collectorName>
10, 
<geoCoordinate id="278DE198E06BDC4BE74094950E8031CB" box="[349,450,983,1009]" degrees="47" direction="north" minutes="30" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="47.5">47º30’N</geoCoordinate>
, 
<geoCoordinate id="278DE198E06BDC4BE7CE94950D7E31CB" box="[467,572,983,1009]" degrees="09" direction="west" minutes="04" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-9.066667">09º04’W</geoCoordinate>
, 
<quantity id="85412ABAE06BDC4BE45394950DEC31CB" box="[590,686,983,1009]" metricMagnitude="3" metricUnit="m" metricValue="2.878" pageId="21" pageNumber="22" unit="m" value="2878.0">
<elevation id="C994606CE06BDC4BE45394950DEC31CB" box="[590,686,983,1009]" metricMagnitude="3" metricUnit="m" metricValue="2.878" pageId="21" pageNumber="22" unit="m" value="2878.0">2878 m</elevation>
</quantity>
, 
<date id="3607A19FE06BDC4BE4DD94950C0B31CB" box="[704,841,983,1009]" pageId="21" pageNumber="22" value="1974-10-21">
<collectingDate id="26435877E06BDC4BE4DD94950C0B31CB" box="[704,841,983,1009]" pageId="21" pageNumber="22" value="1974-10-21">21.10.1974</collectingDate>
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: 
<specimenCount id="54BF4CD6E06BDC4BE54594950CEF31CB" box="[856,941,983,1009]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
<quantity id="85412ABAE06BDC4BE5A494950B6831CB" box="[953,1066,983,1009]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="21" pageNumber="22" unit="mm" value="17.1">17.1 mm</quantity>
. Mid­Atlantic­Ridge, 
<collectionCode id="24A81F9AE06BDC4BE71594BD0E093623" box="[264,331,1023,1049]" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="21" pageNumber="22">MAR</collectionCode>
vent site Snake Pit­Les Ruches, HYDROSNAKE, 
<collectorName id="EF4CE289E06BDC4BE5D194BD0B653623" box="[972,1063,1023,1049]" pageId="21" pageNumber="22">Stn HS</collectorName>
08, 
<collectingMethod id="9BF8FF48E06BDC4BE24A94BD0B9A3623" box="[1111,1240,1023,1049]" pageId="21" pageNumber="22">baited trap</collectingMethod>
PT01, 
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, 
<geoCoordinate id="278DE198E06BDC4BE76193650EA7367B" box="[380,485,1063,1089]" degrees="45" direction="west" minutes="57" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-45.95">45º57’W</geoCoordinate>
, 
<quantity id="85412ABAE06BDC4BE7E993650D11367B" box="[500,595,1063,1089]" metricMagnitude="3" metricUnit="m" metricValue="3.502" pageId="21" pageNumber="22" unit="m" value="3502.0">
<elevation id="C994606CE06BDC4BE7E993650D11367B" box="[500,595,1063,1089]" metricMagnitude="3" metricUnit="m" metricValue="3.502" pageId="21" pageNumber="22" unit="m" value="3502.0">3502 m</elevation>
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, 
<date id="3607A19FE06BDC4BE47F93650DA9367B" box="[610,747,1063,1089]" pageId="21" pageNumber="22" value="1988-06-26">
<collectingDate id="26435877E06BDC4BE47F93650DA9367B" box="[610,747,1063,1089]" pageId="21" pageNumber="22" value="1988-06-26">26.06.1988</collectingDate>
</date>
: 1 ovig. female 
<quantity id="85412ABAE06BDC4BE5A093650B6E367B" box="[957,1068,1063,1089]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="21" pageNumber="22" unit="mm" value="34.7">34.7 mm</quantity>
. Mid­Atlantic­Ridge. 
<collectionCode id="24A81F9AE06BDC4BE715930D0E093653" box="[264,331,1103,1129]" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="21" pageNumber="22">MAR</collectionCode>
vent site Snake Pit­Elan. 
<specimenCode id="121F2F24E06BDC4BE4BC930D0C413653" box="[673,771,1103,1129]" collectionCode="MAR" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="21" pageNumber="22">MAR93</specimenCode>
, Dive 2617, 
<geoCoordinate id="278DE198E06BDC4BE5AC930D0B663653" box="[945,1060,1103,1129]" degrees="23" direction="north" minutes="23" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="23.383333">23°23' N</geoCoordinate>
, 
<geoCoordinate id="278DE198E06BDC4BE225930D0BED3653" box="[1080,1199,1103,1129]" degrees="44" direction="west" minutes="56" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-44.933334">44°56' W</geoCoordinate>
, 
<quantity id="85412ABAE06BDC4BE2DF930D0A673653" box="[1218,1317,1103,1129]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="m" value="3500.0">
<elevation id="C994606CE06BDC4BE2DF930D0A673653" box="[1218,1317,1103,1129]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="m" value="3500.0">3500 m</elevation>
</quantity>
, 
<date id="3607A19FE06BDC4BE71593350ED236AB" box="[264,400,1143,1169]" pageId="21" pageNumber="22" value="1993-06-18">
<collectingDate id="26435877E06BDC4BE71593350ED236AB" box="[264,400,1143,1169]" pageId="21" pageNumber="22" value="1993-06-18">18.06.1993</collectingDate>
</date>
: 
<specimenCount id="54BF4CD6E06BDC4BE78393350D4536AB" box="[414,519,1143,1169]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
32.0 mm. Mid­Atlantic­Ridge. 
<collectionCode id="24A81F9AE06BDC4BE59793350C8F36AB" box="[906,973,1143,1169]" country="Zimbabwe" httpUri="http://grbio.org/cool/p96j-ip08" name="Grasslands Rhizobium Collection" pageId="21" pageNumber="22">MAR</collectionCode>
vent site TAG. AMK47, Stn 4343, slurp gun 4, 
<geoCoordinate id="278DE198E06BDC4BE7EC93DD0D143683" box="[497,598,1183,1209]" degrees="26" direction="north" minutes="08" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="26.133333">26º08’N</geoCoordinate>
, 
<geoCoordinate id="278DE198E06BDC4BE47B93DD0D8D3683" box="[614,719,1183,1209]" degrees="44" direction="west" minutes="49" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="-44.816666">44º49’W</geoCoordinate>
, 
<quantity id="85412ABAE06BDC4BE4C293DD0C7C3683" box="[735,830,1183,1209]" metricMagnitude="3" metricUnit="m" metricValue="3.65" pageId="21" pageNumber="22" unit="m" value="3650.0">
<elevation id="C994606CE06BDC4BE4C293DD0C7C3683" box="[735,830,1183,1209]" metricMagnitude="3" metricUnit="m" metricValue="3.65" pageId="21" pageNumber="22" unit="m" value="3650.0">3650 m</elevation>
</quantity>
, 
<date id="3607A19FE06BDC4BE55093DD0C943683" box="[845,982,1183,1209]" pageId="21" pageNumber="22" value="2002-06-26">
<collectingDate id="26435877E06BDC4BE55093DD0C943683" box="[845,982,1183,1209]" pageId="21" pageNumber="22" value="2002-06-26">26.06.2002</collectingDate>
</date>
: 1 ovig. female 
<quantity id="85412ABAE06BDC4BE28393DD0A4E3683" box="[1182,1292,1183,1209]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" pageId="21" pageNumber="22" unit="mm" value="39.5">39.5 mm</quantity>
, 
<specimenCount id="54BF4CD6E06BDC4BE30193DD0E1A36DB" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
<quantity id="85412ABAE06BDC4BE74293850E8936DB" box="[351,459,1223,1249]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="21" pageNumber="22" unit="mm" value="31.6">31.6 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="4206875FE06BDC4BE72593AC0B3A370B" blockId="21.[264,1323,902,1889]" pageId="21" pageNumber="22">
<emphasis id="70CD5B4DE06BDC4BE72593AC0EC23732" bold="true" box="[312,384,1262,1288]" pageId="21" pageNumber="22">Types</emphasis>
. The ovigerous female of 
<quantity id="85412ABAE06BDC4BE4A493AD0C633733" box="[697,801,1263,1289]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="21" pageNumber="22" unit="mm" value="34.7">34.7 mm</quantity>
from HYDROSNAKE, Stn HS08 (MNHN­ Ga 4621) has been selected as 
<typeStatus id="9D0239FDE06BDC4BE47292550D9B370B" box="[623,729,1303,1329]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, the other specimens are 
<typeStatus id="9D0239FDE06BDC4BE21D92550B36370B" box="[1024,1140,1303,1329]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA3D4D4E06BDC4BE725927C0CD937BB" pageId="21" pageNumber="22" type="etymology">
<paragraph id="4206875FE06BDC4BE725927C0CD937BB" blockId="21.[264,1323,902,1889]" pageId="21" pageNumber="22">
<emphasis id="70CD5B4DE06BDC4BE725927C0EFF3762" bold="true" box="[312,445,1342,1368]" pageId="21" pageNumber="22">Etymology</emphasis>
. From the Latin 
<emphasis id="70CD5B4DE06BDC4BE499927D0D8D3762" box="[644,719,1343,1368]" italics="true" pageId="21" pageNumber="22">exutus</emphasis>
, bared, naked, in reference to the absence of spines (except epigastric spines) on the carapace and abdomen.
</paragraph>
</subSubSection>
<subSubSection id="0AA3D4D4E06BDC49E72592CC0D1334BC" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
<paragraph id="4206875FE06BDC4BE72592CC0BE7355B" blockId="21.[264,1323,902,1889]" pageId="21" pageNumber="22">
<emphasis id="70CD5B4DE06BDC4BE72592CC0E8B3792" bold="true" box="[312,457,1422,1448]" pageId="21" pageNumber="22">Description</emphasis>
. Carapace, exclusive of rostrum, longer than broad, moderately convex from side to side. Cervical groove distinct. Gastric region distinct, posterior part delimited by shallow groove; surface with pair of epigastric spines (2 or 3 additional small spines in 
<specimenCount id="54BF4CD6E06BDC4BE71591450EEC341B" box="[264,430,1543,1569]" pageId="21" pageNumber="22" type="generic">one specimen</specimenCount>
), covered by scale­like rugosities much larger on posterior triangle than on remainder. Anterior branchial region also with scale­like rugosities, as illustrated. Rostrum broad at base, distally narrowed and upturned, weakly carinated dorsally, with fine rugosities, sometimes with 1–4 small lateral spines on each side; antennal spine absent. Oblique frontal margin leading to short anterolateral spine (first spine) directed straight forward, followed by 2 spines on anterior part of anterior branchial region, anterior (second spine) of these strong, directed anterolaterally, situated lateral to level of anterolateral spine, posterior (third spine) much smaller, followed by a few knobs and another small spine (fourth) bordering anterior and posterior branchial regions.
</paragraph>
<caption id="16C6D7D7E068DC48E71591F50E763556" ID-DOI="http://doi.org/10.5281/zenodo.5051372" ID-Zenodo-Dep="5051372" httpUri="https://zenodo.org/record/5051372/files/figure.png" pageId="22" pageNumber="23" startId="22.[264,357,1719,1740]" targetBox="[295,1315,288,1699]" targetPageId="22">
<paragraph id="4206875FE068DC48E71591F50E763556" blockId="22.[264,1323,1719,1900]" pageId="22" pageNumber="23">
<emphasis id="70CD5B4DE068DC48E71591F50EC534F6" bold="true" box="[264,391,1719,1740]" pageId="22" pageNumber="23">FIGURE 5.</emphasis>
<taxonomicName id="85B9FCDCE068DC48E78A91FA0D0934F7" authority="Macpherson &amp; Segonzac, 2005" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[407,587,1720,1741]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="exuta" status="sp. nov.">
<emphasis id="70CD5B4DE068DC48E78A91FA0D0934F7" box="[407,587,1720,1741]" italics="true" pageId="22" pageNumber="23">Munidopsis exuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BFEE636E068DC48E44191F50DD934F6" box="[604,667,1719,1740]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
, holotype, ovigerous female (34.7 mm), Mid­Atlantic Ridge, HYDROSNAKE, Stn HS08, MNHN­Ga 4621. A, carapace, dorsal. B, same, lateral. C, posterior part of sixth abdominal segment and telson. D, anterior part of sternal plastron. E, left antennule, antenna and ocular peduncle, ventral. F, endopod of right third maxilliped, lateral. G, right cheliped, lateral. H, right second pereiopod, lateral. I, dactylus of right second pereiopod, lateral. Scale: A–B, D–E, G. I = 5 mm, C, F, H = 2 mm.
</paragraph>
</caption>
<paragraph id="4206875FE069DC49E725965E0E313394" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Thoracic sternites with a few arcuate setiferous ridges; third thoracic sternite posteriorly narrowed, anterior margin with acute submedian granules and small anterolateral spine on each side; fourth thoracic sternite subtriangular, anteriorly narrow elongate.</paragraph>
<paragraph id="4206875FE069DC49E72596FE0C6E3074" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Abdomen smooth, with few granules on pleura; segments 2–4 each with 2 elevated transverse ridges; segment 6 bearing posterolateral lobes somewhat exceeding nearly straight transverse median margin. Telson composed of 8 plates; length­width ratio 0.78; midlateral plate convex on distolateral margin.</paragraph>
<paragraph id="4206875FE069DC49E725951E0D4630D4" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Eyes not movable; smoothly ovate cornea cupped within broad­based ocular peduncle; peduncle extended into strong but relatively short mesiodorsal spine directed upward at low angle; cornea relatively large, clearly wider than eyespine, and as wide as third article of antennal peduncle.</paragraph>
<paragraph id="4206875FE069DC49E72595BE0D6531B4" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Basal article of antennule having dorsolateral spine much smaller than ventrolateral. Article 1 of antennal peduncle distomesially bearing flattish process rather straight laterally, somewhat convex mesially, distolateral spine well­developed; article 2 with strong distolateral spine.</paragraph>
<paragraph id="4206875FE069DC49E72594DE0CB6363C" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Third maxillipeds with weak spination; ischium with distodorsal and distoventral spines, both small, latter in particular; mesial crest with 21–23 denticles. Flexor margin of merus with 5 small spines, extensor distal marginal spine small.</paragraph>
<paragraph id="4206875FE069DC49E72593560B4D36CC" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">Chelipeds longer than carapace including rostrum. Merus nearly reaching end of rostrum, with 4 terminal spines (2 dorsal, 1 mesial, 1 lateral), a few small spines on dorsal sides, covered with scale­like elevated rugosities. Carpus with a few distal and mesial spines. Palm spineless, slightly longer than broad, with small granules along mesial margin. Fingers slightly longer than palm, distally spooned, prehensile edges crenulated; fixed finger moderately ridged along distal third of lateral margin.</paragraph>
<paragraph id="4206875FE069DC49E72592460A4A340C" blockId="23.[264,1325,284,1870]" pageId="23" pageNumber="24">First walking leg slightly exceeding cheliped by half length of dactylus; dorsal crest of merus with row of small spines, lateral side with granules; carpus with row of spines along dorsal border, two granulate crests along lateral side and continued on to propodus, ventrolateral crest less conspicuous than dorsolateral; propodus nearly twice as long as dactylus, sometimes with spines on proximal half of dorsal margin; dactylus smoothly narrowed distally, ending in curved sharp spine, flexor margin slightly curving, bearing 13 or14 proximally diminishing low spines, ultimate spine rather remote from end of dactylus and much closer to penultimate. Meri of walking legs progressively shorter posteriorly.</paragraph>
<paragraph id="4206875FE069DC49E72591060CC03464" blockId="23.[264,1325,284,1870]" box="[312,898,1604,1630]" pageId="23" pageNumber="24">Epipods on chelipeds, but absent on walking legs.</paragraph>
<paragraph id="4206875FE069DC49E725912E0D1334BC" blockId="23.[264,1325,284,1870]" box="[312,593,1644,1670]" pageId="23" pageNumber="24">
Eggs diameter: 
<quantity id="85412ABAE069DC49E7ED912E0D0F34BC" box="[496,589,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="23" pageNumber="24" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0AA3D4D4E069DC46E72591D60B10363C" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="discussion">
<paragraph id="4206875FE069DC46E72591D60DC03394" blockId="23.[264,1325,284,1870]" lastBlockId="24.[264,1325,284,1670]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="70CD5B4DE069DC49E72591D60EE83494" bold="true" box="[312,426,1684,1710]" pageId="23" pageNumber="24">Remarks</emphasis>
. The new species belongs to the group of species having (usually) 2 epigastric spines, epipods on the chelipeds, the cornea distinctly broader than the eyespine, the fixed finger of the cheliped without a denticulate carina on the distolateral margin, the first walking leg overreaching end of the cheliped, the dactyli of the walking legs slightly curving on flexor margin, and the abdominal segment 6 with the posteromedian margin weakly convex, not produced. 
<taxonomicName id="85B9FCDCE066DC46E469965E0BFE330C" authority="Pequegnat &amp; Pequegnat, 1970" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[628,1212,284,310]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E469965E0C0C330F" box="[628,846,284,309]" italics="true" pageId="24" pageNumber="25">Munidopsis geyeri</emphasis>
<bibRefCitation id="2628FAAEE066DC46E54A965E0BFE330C" author="Pequegnat, L. H. &amp; Pequegnat, W. E." box="[855,1212,284,310]" pageId="24" pageNumber="25" pagination="125 - 170" refId="ref30885" refString="Pequegnat, L. H. &amp; Pequegnat, W. E. (1970) Deep-sea anomurans of superfamily Galatheoidea with description of three new species. Part 5. Contributions on the Biology of the Gulf of Mexico. Texas A &amp; M University oceanographic Studies, 1, 125 - 170." type="journal article" year="1970">Pequegnat &amp; Pequegnat, 1970</bibRefCitation>
</taxonomicName>
from the Caribbean Sea and South and Central Atlantic (see below) is the most similar species geographically and taxonomically, but it is readily differentiated by the following characters (see also 
<bibRefCitation id="2628FAAEE066DC46E7EF96D60D343394" author="Baba, K." box="[498,630,404,430]" pageId="24" pageNumber="25" refId="ref27771" refString="Baba, K. (2005) Deep-sea chirostylid and galatheid crustaceans (Decapoda: Anomura) from the Indo-Pacific, with a list of species. Galathea Report, 20, 317 pp." type="book" year="2005">Baba 2005</bibRefCitation>
):
</paragraph>
<paragraph id="4206875FE066DC46E71596FE0A6833EC" blockId="24.[264,1325,284,1670]" box="[264,1322,444,470]" pageId="24" pageNumber="25">
– The rostrum is triangular with a marked dorsal carina in 
<taxonomicName id="85B9FCDCE066DC46E21096FE0BC633EF" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[1037,1156,444,469]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E21096FE0BC633EF" box="[1037,1156,444,469]" italics="true" pageId="24" pageNumber="25">M. geyeri</emphasis>
</taxonomicName>
, whereas the
</paragraph>
<paragraph id="4206875FE066DC46E72596A60B8433C4" blockId="24.[264,1325,284,1670]" box="[312,1222,484,510]" pageId="24" pageNumber="25">rostrum is clearly spiniform, without marked dorsal carina in the new species.</paragraph>
<paragraph id="4206875FE066DC46E715954E0BA6301C" blockId="24.[264,1325,284,1670]" box="[264,1252,524,550]" pageId="24" pageNumber="25">
– The antennal spine is always present in 
<taxonomicName id="85B9FCDCE066DC46E517954E0C38301F" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[778,890,524,549]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E517954E0C38301F" box="[778,890,524,549]" italics="true" pageId="24" pageNumber="25">M. geyeri</emphasis>
</taxonomicName>
, but absent in the new species.
</paragraph>
<paragraph id="4206875FE066DC46E71595760B123074" blockId="24.[264,1325,284,1670]" box="[264,1104,564,590]" pageId="24" pageNumber="25">
– The eyespine is usually longer in 
<taxonomicName id="85B9FCDCE066DC46E4DC95760C733077" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[705,817,564,589]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E4DC95760C733077" box="[705,817,564,589]" italics="true" pageId="24" pageNumber="25">M. geyeri</emphasis>
</taxonomicName>
than in the new species.
</paragraph>
<paragraph id="4206875FE066DC46E715951E0A6A304C" blockId="24.[264,1325,284,1670]" box="[264,1320,604,630]" pageId="24" pageNumber="25">– The posterolateral lobes of the abdominal segment 6 do not reach the posterior median</paragraph>
<paragraph id="4206875FE066DC46E72595C60B8C30A4" blockId="24.[264,1325,284,1670]" box="[312,1230,644,670]" pageId="24" pageNumber="25">
margin in 
<taxonomicName id="85B9FCDCE066DC46E7B295C60D6230A7" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[431,544,644,669]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E7B295C60D6230A7" box="[431,544,644,669]" italics="true" pageId="24" pageNumber="25">M. geyeri</emphasis>
</taxonomicName>
, whereas in 
<taxonomicName id="85B9FCDCE066DC46E4A995C60C5830A7" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[692,794,644,669]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="exuta">
<emphasis id="70CD5B4DE066DC46E4A995C60C5830A7" box="[692,794,644,669]" italics="true" pageId="24" pageNumber="25">M. exuta</emphasis>
</taxonomicName>
it distinctly overreaches that margin.
</paragraph>
<paragraph id="4206875FE066DC46E71595EE0A6930FC" blockId="24.[264,1325,284,1670]" box="[264,1323,684,710]" pageId="24" pageNumber="25">
– The propodus of the walking legs is nearly as long as the dactylus in 
<taxonomicName id="85B9FCDCE066DC46E24A95EE0BFE30FF" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[1111,1212,684,709]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="exuta">
<emphasis id="70CD5B4DE066DC46E24A95EE0BFE30FF" box="[1111,1212,684,709]" italics="true" pageId="24" pageNumber="25">M. exuta</emphasis>
</taxonomicName>
, whereas
</paragraph>
<paragraph id="4206875FE066DC46E72595960B7030D4" blockId="24.[264,1325,284,1670]" box="[312,1074,724,750]" pageId="24" pageNumber="25">
in 
<taxonomicName id="85B9FCDCE066DC46E74B95960E8430D7" authorityName="Pequegnat &amp; Pequegnat" authorityYear="1970" box="[342,454,724,749]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="geyeri">
<emphasis id="70CD5B4DE066DC46E74B95960E8430D7" box="[342,454,724,749]" italics="true" pageId="24" pageNumber="25">M. geyeri</emphasis>
</taxonomicName>
the propodus is 1.3–1.4 times the length of dactylus.
</paragraph>
<paragraph id="4206875FE066DC46E72595BE0B10363C" blockId="24.[264,1325,284,1670]" pageId="24" pageNumber="25">
<taxonomicName id="85B9FCDCE066DC46E72595BE0EDD312F" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[312,415,764,789]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="exuta">
<emphasis id="70CD5B4DE066DC46E72595BE0EDD312F" box="[312,415,764,789]" italics="true" pageId="24" pageNumber="25">M. exuta</emphasis>
</taxonomicName>
is also close to 
<taxonomicName id="85B9FCDCE066DC46E47E95BE0D9B312F" authorityName="Smith" authorityYear="1885" box="[611,729,764,789]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="70CD5B4DE066DC46E47E95BE0D9B312F" box="[611,729,764,789]" italics="true" pageId="24" pageNumber="25">M. crassa</emphasis>
</taxonomicName>
(see above), but they can be differentiated by the shape of the distal margin of abdominal segment 6: weakly convex in the new species and strongly produced in 
<taxonomicName id="85B9FCDCE066DC46E41E940E0D34315F" authorityName="Smith" authorityYear="1885" box="[515,630,844,869]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="70CD5B4DE066DC46E41E940E0D34315F" box="[515,630,844,869]" italics="true" pageId="24" pageNumber="25">M. crassa</emphasis>
</taxonomicName>
. Furthermore, 
<taxonomicName id="85B9FCDCE066DC46E53A940E0CCF315F" authorityName="Macpherson &amp; Segonzac" authorityYear="2005" box="[807,909,844,869]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="exuta">
<emphasis id="70CD5B4DE066DC46E53A940E0CCF315F" box="[807,909,844,869]" italics="true" pageId="24" pageNumber="25">M. exuta</emphasis>
</taxonomicName>
is characterized by the presence of two epigastric spines, the absence of antennal spine, the rostrum not triangular, and the dactyli of the walking legs slightly curving on the flexor margin, whereas 
<taxonomicName id="85B9FCDCE066DC46E29994DE0BB9318F" authorityName="Smith" authorityYear="1885" box="[1156,1275,924,949]" class="Malacostraca" family="Galatheidae" genus="Munidopsis" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="70CD5B4DE066DC46E29994DE0BB9318F" box="[1156,1275,924,949]" italics="true" pageId="24" pageNumber="25">M. crassa</emphasis>
</taxonomicName>
has some gastric spines, the antennal spine present, the rostrum moderately broadly triangular, and the dactyli of the walking legs clearly curving on the flexor margin.
</paragraph>
</subSubSection>
<subSubSection id="0AA3D4D4E066DC46E72593560D0B34BC" pageId="24" pageNumber="25" type="distribution">
<paragraph id="4206875FE066DC46E72593560D0B34BC" blockId="24.[264,1325,284,1670]" pageId="24" pageNumber="25">
<emphasis id="70CD5B4DE066DC46E72593560D2F3614" bold="true" box="[312,621,1044,1070]" pageId="24" pageNumber="25">Distribution and habitat</emphasis>
. The species is only known from the northeast Atlantic at 
<quantity id="85412ABAE066DC46E715937E0EF2366C" box="[264,432,1084,1110]" metricMagnitude="3" metricUnit="m" metricValue="3.181" metricValueMax="3.502" metricValueMin="2.86" pageId="24" pageNumber="25" unit="m" value="3181.0" valueMax="3502.0" valueMin="2860.0">2860–3502 m</quantity>
. This species occurs in both abyssal and hydrothermal vent environments. The BIOGAS specimens were collected in the Bay of Biscay, among actiniarians, polychaetes, brachiopods, molluscans and echinoderms. Two specimens were collected in the vent area of Snake Pit (HYDROSNAKE and MAR93 cruises). The HYDROSNAKE specimen was caught by a baited trap 
<quantity id="85412ABAE066DC46E4DC939E0C4936CC" box="[705,779,1244,1270]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="100.0">100 m</quantity>
north of the site Les Ruches, together with the nephropid lobster 
<taxonomicName id="85B9FCDCE066DC46E7FC92460B5C3724" authority="Segonzac &amp; Macpherson" authorityName="Segonzac &amp; Macpherson" authorityYear="2003" box="[481,1054,1284,1310]" class="Malacostraca" family="Nephropidae" genus="Thymopides" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="laurentae">
<emphasis id="70CD5B4DE066DC46E7FC92460DA53727" box="[481,743,1284,1309]" italics="true" pageId="24" pageNumber="25">Thymopides laurentae</emphasis>
Segonzac &amp; Macpherson
</taxonomicName>
, and the macrurid fish 
<taxonomicName id="85B9FCDCE066DC46E715926E0B33377C" authority="(Hector) (Segonzac &amp; Macpherson 2003)" authorityName="Segonzac &amp; Macpherson" authorityYear="2003" baseAuthorityName="Hector" baseAuthorityYear="1875" box="[264,1137,1324,1350]" class="Actinopterygii" family="Macrouridae" genus="Coryphaenoides" kingdom="Animalia" order="Gadiformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="armatus">
<emphasis id="70CD5B4DE066DC46E715926E0D7E377F" box="[264,572,1324,1349]" italics="true" pageId="24" pageNumber="25">Coryphaenoides armatus</emphasis>
(Hector) (
<bibRefCitation id="2628FAAEE066DC46E4CC926E0B2A377C" author="Segonzac, M. &amp; Macpherson, E." box="[721,1128,1324,1350]" pageId="24" pageNumber="25" pagination="361 - 367" refId="ref31298" refString="Segonzac, M. &amp; Macpherson, E. (2003) A new deep-sea lobster of the genus Thymopides (Crustacea: Decapoda: Nephropidae) collected near the hydrothermal vent Snake-Pit, Mid-Atlantic Ridge. Cahiers de Biologie Marine, 44, 361 - 367." type="journal article" year="2003">Segonzac &amp; Macpherson 2003</bibRefCitation>
)
</taxonomicName>
. The MAR 
<specimenCount id="54BF4CD6E066DC46E30D926E0E343754" pageId="24" pageNumber="25" type="generic">93 specimen</specimenCount>
was collected from the site Elan, about 
<quantity id="85412ABAE066DC46E55192160CD53754" box="[844,919,1364,1390]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="m" value="100.0">100 m</quantity>
west of Les Ruches, at the base of the active edifice, on sulfide rocks, among scattered sea anemones, chaetopterid tubeworms and alvinocaridid shrimps. The 
<specimenCount id="54BF4CD6E066DC46E50D92E60CFD3784" box="[784,959,1444,1470]" pageId="24" pageNumber="25" type="generic">two specimens</specimenCount>
from the AMK47 cruise were collected at the base of an inactive chimney complex, among dead mytilid bivalves. An additional specimen (unfortunately lost) was collected during the cruise BRIDGE 1993, at the hydrothermal vent site Broken Spur site (dive Alvin 2625, 
<date id="3607A19FE066DC46E23B915E0BEC340C" box="[1062,1198,1564,1590]" pageId="24" pageNumber="25" value="1993-06-27">27.06.1993</date>
, 
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, 
<geoCoordinate id="278DE198E066DC46E71591060E363464" box="[264,372,1604,1630]" degrees="43" direction="west" minutes="10" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-43.166668">43°10’W</geoCoordinate>
, 
<quantity id="85412ABAE066DC46E79F91060EA23464" box="[386,480,1604,1630]" metricMagnitude="3" metricUnit="m" metricValue="3.056" pageId="24" pageNumber="25" unit="m" value="3056.0">3056 m</quantity>
; E. Southward, Marine Biological Association of the 
<collectingCountry id="3AAEC7CFE066DC46E27491060BE63464" box="[1129,1188,1604,1630]" name="United Kingdom" pageId="24" pageNumber="25">U.K.</collectingCountry>
, 
<collectingRegion id="807D49BDE066DC46E2AE91060A6A3464" box="[1203,1320,1604,1630]" country="United Kingdom" name="Plymouth" pageId="24" pageNumber="25">Plymouth</collectingRegion>
, personnal communication).
</paragraph>
</subSubSection>
</treatment>
</document>