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<document id="DCFA7B0FDBE8881AF7E5E0FBB18917C4" ID-CLB-Dataset="21486" ID-DOI="10.3161/150811013X678955" ID-GBIF-Dataset="69c434f8-762e-4e43-bc29-d0da90994625" ID-ISSN="1733-5329" ID-Zenodo-Dep="3943563" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594705346288" checkinUser="felipe" docAuthor="Bates, Paul J. J., Cameron, Kenneth, Pearch, Malcolm J. &amp; Hayes, Benjamin" docDate="2013" docId="837C87DBFF9BC075FE84FBDEFC38FAF2" docLanguage="en" docName="ActaChiropterologica.15.2.313-340.pdf.imf" docOrigin="Acta Chiropterologica 15 (2)" docStyle="DocumentStyle:A870862413AAF9DEC7D3C2B4FC7FB2AC.1:ActaChiropterol.2008-2015.journal_article" docStyleId="A870862413AAF9DEC7D3C2B4FC7FB2AC" docStyleName="ActaChiropterol.2008-2015.journal_article" docStyleVersion="1" docTitle="Neoromicia nana" docType="treatment" docVersion="11" lastPageNumber="327" masterDocId="7F45FFA3FF94C07AFF9CFFFFFFC9FFC4" masterDocTitle="A review of the bats (Chiroptera) of the Republic of Congo, including eight species new to the country" masterLastPageNumber="340" masterPageNumber="313" pageNumber="327" updateTime="1698843371229" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9F672B903F5035F6E9B9D90018656DD3">A review of the bats (Chiroptera) of the Republic of Congo, including eight species new to the country</mods:title>
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<mods:namePart id="7448150492600BE88DA60F08AFE56118">Bates, Paul J. J.</mods:namePart>
<mods:affiliation id="B04A558D87723BBDC3E658F1E133E174">Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St Botolphs Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom &amp; Corresponding author: E-mail: pjjbates 2 @ hotmail. com</mods:affiliation>
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<mods:namePart id="BD36E96325C00EF2DC9344878ED84160">Cameron, Kenneth</mods:namePart>
<mods:affiliation id="68880EEEEAAE67A8674D3BE727752B5D">Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, New York 10460, USA</mods:affiliation>
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<mods:namePart id="587C7E15D01D3EEBDFE29F6D78209510">Pearch, Malcolm J.</mods:namePart>
<mods:affiliation id="AF76EDAF9CA7D9D0913D28E9891D25B6">Harrison Institute, Centre for Systematics and Biodiversity Research, Bowerwood House, St Botolphs Road, Sevenoaks, Kent, TN 13 3 AQ, United Kingdom</mods:affiliation>
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<mods:namePart id="A0F7DEC9BB66308C3E33AF5BF81BF2ED">Hayes, Benjamin</mods:namePart>
<mods:affiliation id="47B7D7CFE5C54F351F55632B743C7926">Monadh, Inveruglas, Kingussie, Inverness-shire, United Kingdom</mods:affiliation>
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<mods:title id="045F4D0950DC9A1EF79568322EF9BBC8">Acta Chiropterologica</mods:title>
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<mods:date id="DB2CA9910CECFCD3E937F9158E60D9FE">2013</mods:date>
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<mods:number id="B89AD18761642BAD896DD2E00F16EB78">15</mods:number>
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<treatment id="837C87DBFF9BC075FE84FBDEFC38FAF2" ID-DOI="http://doi.org/10.5281/zenodo.4332330" ID-GBIF-Taxon="165088909" ID-Zenodo-Dep="4332330" LSID="urn:lsid:plazi:treatment:837C87DBFF9BC075FE84FBDEFC38FAF2" httpUri="http://treatment.plazi.org/id/837C87DBFF9BC075FE84FBDEFC38FAF2" lastPageNumber="327" pageId="15" pageNumber="327">
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<taxonomicName id="CCD54D4EFF9BC075FE84FBDEFD58FBFF" ID-CoL="8LF7P" authority="(Peters, 1852)" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[280,657,1057,1083]" class="Mammalia" family="Vespertilionidae" genus="Neoromicia" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="327" phylum="Chordata" rank="species" species="nana">
<emphasis id="39A1EADFFF9BC075FE84FBDEFE2CFBFF" box="[280,485,1057,1083]" italics="true" pageId="15" pageNumber="327">Neoromicia nana</emphasis>
(Peters, 1852)
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<paragraph id="0B6A36CDFF9BC075FE13FBBDFDDAFB98" blockId="15.[159,755,1057,1143]" box="[399,531,1090,1116]" pageId="15" pageNumber="327">Banana bat</paragraph>
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<treatmentCitationGroup id="2BC511E3FF9BC075FF03FB9DFD3AFBB3" box="[159,755,1122,1143]" pageId="15" pageNumber="327">
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<taxonomicName id="CCD54D4EFF9BC075FF03FB9DFE22FBB3" ID-CoL="8LJ82" authority="Peters, 1852: 63" authorityName="Peters" authorityPageNumber="63" authorityYear="1852" box="[159,491,1122,1143]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="327" phylum="Chordata" rank="species" species="nanus">
<emphasis id="39A1EADFFF9BC075FF03FB9DFE8CFBB3" box="[159,325,1122,1143]" italics="true" pageId="15" pageNumber="327">Vespertilio nanus</emphasis>
<bibRefCitation id="6F444B3CFF9BC075FED0FB9DFE22FBB3" box="[332,491,1122,1143]" pageId="15" pageNumber="327">Peters, 1852: 63</bibRefCitation>
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;
<collectingRegion id="C911F82FFF9BC075FE6AFB9DFDAAFBB3" box="[502,611,1122,1143]" country="Mozambique" name="Inhambane" pageId="15" pageNumber="327">Inhambane</collectingRegion>
,
<collectingCountry id="73C2765DFF9BC075FDF1FB9DFD26FBB3" box="[621,751,1122,1143]" name="Mozambique" pageId="15" pageNumber="327">Mozambique</collectingCountry>
.
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</paragraph>
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<subSubSection id="43CF6546FF9BC075FF03FB5DFF20FA63" pageId="15" pageNumber="327" type="materials_examined">
<paragraph id="0B6A36CDFF9BC075FF03FB5DFEF7FB78" blockId="15.[159,771,1186,1447]" box="[159,318,1186,1212]" pageId="15" pageNumber="327">
<emphasis id="39A1EADFFF9BC075FF03FB5DFEF7FB78" box="[159,318,1186,1212]" italics="true" pageId="15" pageNumber="327">New material</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9BC075FF59FB3CFF20FA63" blockId="15.[159,771,1186,1447]" pageId="15" pageNumber="327">
<materialsCitation id="BBBD3C90FF9BC075FF59FB3CFD9FFB3B" ID-GBIF-Occurrence="2813185311" collectingDate="1 August, 2012" collectionCode="HZM" latitude="-2.7794333" location="Airport" longLatPrecision="1" longitude="13.5599" pageId="15" pageNumber="327" specimenCode="HZM.321.40186" specimenCount="1" specimenCount-male="1" stateProvince="Lekoumou">
<specimenCode id="5B739EB6FF9BC075FF59FB3CFE42FB19" box="[197,395,1219,1245]" pageId="15" pageNumber="327">HZM.321.40186</specimenCode>
,
<specimenCount id="1DD3FD44FF9BC075FE0AFB3CFE67FB19" box="[406,430,1219,1245]" pageId="15" pageNumber="327" type="male"></specimenCount>
,
<collectingDate id="6F2FE9E5FF9BC075FE26FB3CFDA4FB19" box="[442,621,1219,1245]" pageId="15" pageNumber="327">
<date id="7F6B100DFF9BC075FE26FB3CFDECFB19" box="[442,549,1219,1245]" pageId="15" pageNumber="327">1 August</date>
, 2012
</collectingDate>
,
<location id="0E0A6016FF9BC075FDEBFB3CFD06FB19" LSID="urn:lsid:plazi:treatment:837C87DBFF9BC075FE84FBDEFC38FAF2:0E0A6016FF9BC075FDEBFB3CFD06FB19" box="[631,719,1219,1245]" latitude="-2.7794333" longLatPrecision="1" longitude="13.5599" name="Airport" pageId="15" pageNumber="327" stateProvince="Lekoumou">Airport</location>
,
<collectingRegion id="C911F82FFF9BC075FD47FB3CFECAFB3B" pageId="15" pageNumber="327">Lekoumou</collectingRegion>
,
<geoCoordinate id="6EE1500AFF9BC075FE85FB1AFE68FB3B" box="[281,417,1253,1279]" degrees="2" direction="south" minutes="46.766" orientation="latitude" pageId="15" pageNumber="327" precision="1" value="-2.7794333">2°46.766S</geoCoordinate>
,
<geoCoordinate id="6EE1500AFF9BC075FE25FB1AFD9BFB3B" box="[441,594,1253,1279]" degrees="13" direction="east" minutes="33.594" orientation="longitude" pageId="15" pageNumber="327" precision="1" value="13.5599">13°33.594E</geoCoordinate>
.
</materialsCitation>
<collectorName id="A620531BFF9BC075FDF5FB1AFD19FB3B" box="[617,720,1253,1279]" pageId="15" pageNumber="327">Previous</collectorName>
records are included in
<figureCitation id="93EE2A48FF9BC075FE3CFAF8FDE0FAE5" box="[416,553,1287,1313]" captionStart="APPENDIX" captionStartId="21.[99,120,486,512]" captionTargetPageId="21" captionText="APPENDIX I A list of the forty-three bat species currently recorded from Congo, including citations in the literature and locality data. CS (Conservation status NT: Near threatened; DD: Data deficient; LC: Least concern) is based on IUCN Red List of Threatened Species retrieved on 8 May, 2013. Two species listed here have not been assessed (na) by IUCN for their conservation status but rather have been included within other taxa: Triaenops afer in Triaenops persicus and Chaerephon leucogaster in Tadarida pumila. Superscripts associated with localities in column 4 refer to the superscripts for the literature cited in column 3" figureDoi="http://doi.org/10.5281/zenodo.3943589" httpUri="https://zenodo.org/record/3943589/files/figure.png" pageId="15" pageNumber="327">
<collectorName id="A620531BFF9BC075FE3CFAF8FDE0FAE5" box="[416,553,1287,1313]" pageId="15" pageNumber="327">Appendix I</collectorName>
</figureCitation>
; the known distribution corresponds to localities 17, 29, and
<quantity id="CC2D9B28FF9BC075FD5FFAD7FCCAFA86" box="[707,771,1320,1346]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="15" pageNumber="327" unit="in" value="39.0">39 in</quantity>
<figureCitation id="93EE2A48FF9BC075FF03FAB5FF2CFAA0" box="[159,229,1354,1380]" captionStart="FIG" captionStartId="3.[159,172,1686,1707]" captionTargetBox="[166,1418,221,1664]" captionTargetId="figure@3.[135,1537,218,1670]" captionTargetPageId="3" captionText="FIG. 1. Localities from which bats have been collected in Congo based on the literature and current study (see Appendix I). Size of red dot is proportional in area to the number of species collected from that locality. Habitat types are based on Bergmans (1979). Localities (listed in Appendix III): (1) Pointe-Noire; (2) Hinda; (3) Loandjili; (4) Tchissanga; (5) Koubotchi; (6) Ménengué; (7) NGongo; (8) Dinguembo; (9) Béna; (10) Goumina; (11) Sounda, Gorge of; (12) Makaba; (13) Bouloungi; (14) Dimonika; (15) Mayombe; (16) Bamba, col du Mont; (17ac) Loudima-Kimongo (includes Cave of Doumboula, Cave of Loudima, and Cave of Viaduc); (18a,b) Kila NTari and Mouyondzi; (19ah) Bangou Massif (includes Bouaboumpono, Cave of Kimanika, Cave of Matouridi, Meya / Cave of Meya-Nzouari, Cave of Meya-Nwadi, Mountembessa, Cave of Mpasa, and Cave of Mpoka); (20) Voka; (21) Kinkala; (22) Ile MBamou; (23) La Région du Pool (Stanley Pool); (24) Brazzaville; (25) Grand Bois; (26) Sibiti; (27) Komono; (28) Divenié; (29) Lekoumou; (30) Lesio-Louna Reserve; (31) Ngabé; (32) Plateau Konkouya; (33) Lékana; (34) Djambala; (35) Ewo; (36) Konda; (37) Ubangi (Oubangi); (38) Mouth of Ubangi River; (39) Kellé; (40) Odzala/Odzala National Park; (41) Kango; (42) Impfondo; (43) Epena. Scale as marked. C.A.R.= Central African Republic; D.R. Congo = Democratic Republic of Congo" figureDoi="http://doi.org/10.5281/zenodo.3943565" httpUri="https://zenodo.org/record/3943565/files/figure.png" pageId="15" pageNumber="327">Fig. 1</figureCitation>
. This species is apparently common wherever banana plants are abundant (
<bibRefCitation id="6F444B3CFF9BC075FE69FA93FF14FA63" author="Happold and Happold" firstAuthor="Happold" pageId="15" pageNumber="327" refId="ref13898" refString="HAPPOLD, M., and D. C. D. HAPPOLD (eds.). 2013. Mammals of Africa Volume IV: Hedgehogs, shrews, and bats. Bloomsbury Publishing, London, 800 pp." type="book" year="2013">Happold and Happold, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="43CF6546FF9BC075FF03FA2FFC27FB80" pageId="15" pageNumber="327" type="description">
<paragraph id="0B6A36CDFF9BC075FF03FA2FFEE3FA2E" blockId="15.[159,772,1488,2019]" box="[159,298,1488,1514]" pageId="15" pageNumber="327">
<emphasis id="39A1EADFFF9BC075FF03FA2FFEE3FA2E" box="[159,298,1488,1514]" italics="true" pageId="15" pageNumber="327">Description</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9BC075FF59FA0DFC27FB80" blockId="15.[159,772,1488,2019]" lastBlockId="15.[816,1429,855,1092]" pageId="15" pageNumber="327">
A very small, nondescript species with a forearm length of
<quantity id="CC2D9B28FF9BC075FE89F9EBFE48F9EA" box="[277,385,1556,1582]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="15" pageNumber="327" unit="mm" value="31.1">31.1 mm</quantity>
(
<tableCitation id="46570376FF9BC075FE0FF9EBFE38F9EA" box="[403,497,1556,1582]" captionStart="TABLE" captionStartId="7.[159,172,220,241]" captionTargetPageId="7" captionText="TABLE 2. External measurements (mm) and bacular length (mm) of 14 bat species recently collected at Lekoumou, Congo.Acronyms explained in the Materials and Methods section include means, ranges, and, where relevant, standard deviations. Numbers in parentheses refer to sample size where they differ from those stated in column 2" httpUri="http://table.plazi.org/id/5FAA6645FF93C07DFF03FF23FC6EFEEE" pageId="15" pageNumber="327" tableUuid="5FAA6645FF93C07DFF03FF23FC6EFEEE">Table 2</tableCitation>
). The soles of the feet and base of the thumb are slightly swollen to assist the bat when moving inside smooth banana leaves. In the ear, the tragus has a virtually straight anterior border, a rounded tip, and an angular posterior border, without a basal notch (
<figureCitation id="93EE2A48FF9BC075FE6AF943FD9EF912" box="[502,599,1724,1750]" captionStart="FIG" captionStartId="11.[159,172,1911,1932]" captionTargetBox="[287,1299,1270,1885]" captionTargetId="figure@11.[211,1406,1263,1892]" captionTargetPageId="11" captionText="FIG. 8. Right tragus of (A): Glauconycteris alboguttata, HZM.1.40172, ♀; (B): G. beatrix, HZM.1.40181, ♀; (C): Hypsugo crassulus, HZM.1.40190, ♂; (D): Neoromicia brunnea, HZM.1.40191, ♂; (E): N. nana, HZM.321.40186, ♂; (F): N. tenuipinnis, HZM.7.40170, ♀; (G): Pipistrellus nanulus, HZM.1.40189, ♂; (H): Mimetillus moloneyi, HZM.10.40177, ♂; (I): Miniopterus inflatus, HZM.19.40182, ♂. Scale = 2 mm" figureDoi="http://doi.org/10.5281/zenodo.3943579" httpUri="https://zenodo.org/record/3943579/files/figure.png" pageId="15" pageNumber="327">Fig. 8E</figureCitation>
). The skull is small (GTL =
<quantity id="CC2D9B28FF9BC075FED5F922FE0BF933" box="[329,450,1757,1783]" metricMagnitude="-2" metricUnit="m" metricValue="1.172" pageId="15" pageNumber="327" unit="mm" value="11.72">11.72 mm</quantity>
) (
<tableCitation id="46570376FF9BC075FE45F922FDFDF933" box="[473,564,1757,1783]" captionStart="TABLE" captionStartId="8.[99,112,725,746]" captionTargetPageId="8" captionText="TABLE 3. Cranial and dental measurements (mm) of 14 bat species recently collected at Lekoumou, Congo.Acronyms explained in the Materials and Methods section include means, ranges, and, where relevant,standard deviations. Numbers in parentheses refer to sample size where they differ from those stated in column 2" httpUri="http://table.plazi.org/id/5FAA6645FF9CC072FFFFFD2AFAA7FCC7" pageId="15" pageNumber="327" tableUuid="5FAA6645FF9CC072FFFFFD2AFAA7FCC7">Table 3</tableCitation>
); the braincase is distinctly elevated above the rostrum, with a marked concavity in the interorbital region (
<figureCitation id="93EE2A48FF9BC075FDCDF8DFFD76F8FE" box="[593,703,1824,1850]" captionStart="FIG" captionStartId="16.[159,172,1289,1310]" captionTargetBox="[390,1220,230,1257]" captionTargetId="figure@16.[352,1259,219,1273]" captionTargetPageId="16" captionText="FIG. 12. Skulls of three Neoromicia, including lateral view of right upper incisors, and occlusal view of left mandibular dentition [left] and left maxillary dentition [right] of (A): N. brunnea, HZM.1.40191, ♂; (B): N. nana, HZM.321.40186, ♂ and (C) N. tenuipinnis, HZM.7.40170, ♀. Scales for skulls and occlusal views of toothrows (left and right) = 2 mm; scale for upper incisors (centre) = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3943587" httpUri="https://zenodo.org/record/3943587/files/figure.png" pageId="15" pageNumber="327">Fig. 12B</figureCitation>
). The first upper incisor (I
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) is narrow and small with a notch (indicative of a secondary cusp) at about two-thirds its height; the second upper incisor (I
<superScript id="FCA09B85FF9BC075FD48F87EFD17F857" attach="left" box="[724,734,1921,1939]" fontSize="8" pageId="15" pageNumber="327">3</superScript>
) is also narrow, unicuspid and about three-quarters the height of I
<superScript id="FCA09B85FF9BC075FEB9F83BFEE6F812" attach="left" box="[293,303,1988,2006]" fontSize="8" pageId="15" pageNumber="327">2</superScript>
(
<figureCitation id="93EE2A48FF9BC075FED9F836FE7EF827" box="[325,439,1993,2019]" captionStart="FIG" captionStartId="16.[159,172,1289,1310]" captionTargetBox="[390,1220,230,1257]" captionTargetId="figure@16.[352,1259,219,1273]" captionTargetPageId="16" captionText="FIG. 12. Skulls of three Neoromicia, including lateral view of right upper incisors, and occlusal view of left mandibular dentition [left] and left maxillary dentition [right] of (A): N. brunnea, HZM.1.40191, ♂; (B): N. nana, HZM.321.40186, ♂ and (C) N. tenuipinnis, HZM.7.40170, ♀. Scales for skulls and occlusal views of toothrows (left and right) = 2 mm; scale for upper incisors (centre) = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3943587" httpUri="https://zenodo.org/record/3943587/files/figure.png" pageId="15" pageNumber="327">Fig. 12B</figureCitation>
). The first upper premolar (P
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) is minute and situated slightly internal to the toothrow but is visible when the skull is viewed laterally (
<figureCitation id="93EE2A48FF9BC075FC3DFC5FFBC4FC7E" box="[929,1037,928,954]" captionStart="FIG" captionStartId="16.[159,172,1289,1310]" captionTargetBox="[390,1220,230,1257]" captionTargetId="figure@16.[352,1259,219,1273]" captionTargetPageId="16" captionText="FIG. 12. Skulls of three Neoromicia, including lateral view of right upper incisors, and occlusal view of left mandibular dentition [left] and left maxillary dentition [right] of (A): N. brunnea, HZM.1.40191, ♂; (B): N. nana, HZM.321.40186, ♂ and (C) N. tenuipinnis, HZM.7.40170, ♀. Scales for skulls and occlusal views of toothrows (left and right) = 2 mm; scale for upper incisors (centre) = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3943587" httpUri="https://zenodo.org/record/3943587/files/figure.png" pageId="15" pageNumber="327">Fig. 12B</figureCitation>
). The first lower premolar (P
<subScript id="97513488FF9BC075FAF9FC50FAA6FC05" attach="left" box="[1381,1391,943,961]" fontSize="8" pageId="15" pageNumber="327">2</subScript>
) is about two-thirds the crown area and height of the second (P
<subScript id="97513488FF9BC075FC32FC0BFC71FBC2" attach="left" box="[942,952,1012,1030]" fontSize="8" pageId="15" pageNumber="327">4</subScript>
). The baculum has a short, slightly curved shaft, a flattened, broader tip and a bilobate base (
<figureCitation id="93EE2A48FF9BC075FCE9FBD5FC29FB80" box="[885,992,1066,1092]" captionStart="FIG" captionStartId="15.[159,172,710,731]" captionTargetBox="[329,1234,229,686]" captionTargetId="figure@15.[313,1265,219,692]" captionTargetPageId="15" captionText="FIG. 11. Bacula of four pipistrelles. (A): Hypsugo crassulus, HZM.1.40190; (B): Neoromicia brunnea, HZM.1.40191; (C): N. nana, HZM.321.40186; and (D): Pipistrellus nanulus, HZM.1.40189. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3943585" httpUri="https://zenodo.org/record/3943585/files/figure.png" pageId="15" pageNumber="327">Fig. 11C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="43CF6546FF9BC075FCACFB90FC38FAF2" pageId="15" pageNumber="327" type="discussion">
<paragraph id="0B6A36CDFF9BC075FCACFB90FC3FFB4D" blockId="15.[816,1429,1135,1334]" box="[816,1014,1135,1161]" pageId="15" pageNumber="327">
<emphasis id="39A1EADFFF9BC075FCACFB90FC3FFB4D" box="[816,1014,1135,1161]" italics="true" pageId="15" pageNumber="327">Taxonomic notes</emphasis>
</paragraph>
<paragraph id="0B6A36CDFF9BC075FCCAFB6DFC38FAF2" blockId="15.[816,1429,1135,1334]" pageId="15" pageNumber="327">
Geographical variation in
<taxonomicName id="CCD54D4EFF9BC075FB06FB6DFB34FB68" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[1178,1277,1170,1196]" class="Mammalia" family="Vespertilionidae" genus="Neoromicia" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="327" phylum="Chordata" rank="species" species="nana">
<emphasis id="39A1EADFFF9BC075FB06FB6DFB34FB68" box="[1178,1277,1170,1196]" italics="true" pageId="15" pageNumber="327">N. nana</emphasis>
</taxonomicName>
seems little understood (
<bibRefCitation id="6F444B3CFF9BC075FC4DFB4BFB52FB0A" author="Rosevear" box="[977,1179,1204,1230]" firstAuthor="Rosevear" pageId="15" pageNumber="327" refId="ref14501" refString="ROSEVEAR, D. R. 1965. The bats of West Africa. British Museum (Natural History), London, xvii + 418 pp." type="book" year="1965">Rosevear, 1965</bibRefCitation>
;
<bibRefCitation id="6F444B3CFF9BC075FB2EFB4BFCA5FB35" author="Monadjem" etAl="et al." firstAuthor="Monadjem" pageId="15" pageNumber="327" refId="ref14301" refString="MONADJEM, A., P. J. TAYLOR, F. P. D. COTTERILL, and M. C. SCHOEMAN. 2010. Bats of Central and Southern Africa: a biogeographic and taxonomic synthesis. WITS University Press, Johannesburg, 596 pp." type="book" year="2010">
Monadjem
<emphasis id="39A1EADFFF9BC075FADAFB4BFA4CFB0A" box="[1350,1413,1204,1230]" italics="true" pageId="15" pageNumber="327">et al</emphasis>
., 2010
</bibRefCitation>
) and currently all specimens from
<collectingCountry id="73C2765DFF9BC075FA84FB28FAAFFB35" box="[1304,1382,1239,1265]" name="Democratic Republic of the Congo" pageId="15" pageNumber="327">Congo</collectingCountry>
are referred to the nominate subspecies (
<bibRefCitation id="6F444B3CFF9BC075FB6BFB06FC2CFAF2" author="Happold and Happold" firstAuthor="Happold" pageId="15" pageNumber="327" refId="ref13898" refString="HAPPOLD, M., and D. C. D. HAPPOLD (eds.). 2013. Mammals of Africa Volume IV: Hedgehogs, shrews, and bats. Bloomsbury Publishing, London, 800 pp." type="book" year="2013">Happold and Happold, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>