544 lines
132 KiB
XML
544 lines
132 KiB
XML
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<mods:title id="3C953F89ED051A45AF18AB2E89DABC02">Mixtophyes toothlessi sp. n., a new Kinorhyncha species (Anomoirhaga: Neocentrophyidae) from the Clarion-Clipperton Zone, with a revision of terminology and taxonomic features of the family</mods:title>
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<mods:roleTerm id="FDAB5F2D6225CA7714B5892AA94BC42E">Author</mods:roleTerm>
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<mods:namePart id="D78A601E67FC3E80D4683B36059CE7ED">González-Casarrubios, Alberto</mods:namePart>
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<mods:nameIdentifier id="E8047A176B792E82B572080AD172C43D" type="ORCID">0000-0003-1975-6900</mods:nameIdentifier>
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<mods:affiliation id="35E76841098F62CCC0A4B133C014D086">albert 23 @ ucm. es; https: // orcid. org / 0000 - 0003 - 1975 - 6900</mods:affiliation>
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<mods:nameIdentifier id="D6478AEE294CE0B60D3C2B734425783D" type="email">albert23@ucm.es</mods:nameIdentifier>
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<mods:namePart id="43D0309735DFE4BC575F2BA74856E319">Arbizu, Pedro Martínez</mods:namePart>
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<mods:nameIdentifier id="A550D00FD0F0FFA98616CE75FB163BD8" type="ORCID">0000-0002-0891-1154</mods:nameIdentifier>
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<mods:affiliation id="70B9E4C8211AD8417596147D822D4087">German Centre for Marine Biodiversity Research, Senckenberg am Meer, Wilhelmshaven, Germany. pedro. martinez @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 0891 - 1154</mods:affiliation>
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<mods:nameIdentifier id="1460F22B82D355D7AC8365F8F3BCAAA7" type="email">pedro.martinez@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="0899530550E6B68A00C5134CFE396EFB">Sánchez, Nuria</mods:namePart>
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<mods:affiliation id="B46E74B2E33C218E79235D916CCB5CEA">nurisanc @ ucm. es; https: // orcid. org / 0000 - 0003 - 4908 - 8755</mods:affiliation>
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<mods:nameIdentifier id="BCC31109863C5A9D8EF2E785D6F14C65" type="email">nurisanc@ucm.es</mods:nameIdentifier>
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<mods:title id="D9EC9E25DF8409F048FE002AE1189D8D">Zootaxa</mods:title>
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<mods:date id="BEBD6C6CF97F8E09BC955FA138F12EE4">2023</mods:date>
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<treatment id="803AC27CFFD9FF8A18D324CCFBF2F84C" ID-DOI="http://doi.org/10.5281/zenodo.7942463" ID-GBIF-Taxon="217397015" ID-Zenodo-Dep="7942463" LSID="urn:lsid:plazi:treatment:803AC27CFFD9FF8A18D324CCFBF2F84C" httpUri="http://treatment.plazi.org/id/803AC27CFFD9FF8A18D324CCFBF2F84C" lastPageId="8" lastPageNumber="157" pageId="3" pageNumber="152">
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<heading id="5364C406FFD9FF8118D324CCFE54FF1A" bold="true" box="[151,486,223,250]" fontSize="11" level="1" pageId="3" pageNumber="152" reason="1">
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<emphasis id="3AE7AF78FFD9FF8118D324CCFE54FF1A" bold="true" box="[151,486,223,250]" pageId="3" pageNumber="152">
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<taxonomicName id="CF9308E9FFD9FF8118D324CCFE2FFF19" ID-CoL="d1e98507-1cd6-486d-9b76-dbcdc6adca02" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" box="[151,413,223,250]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="3" pageNumber="152" phylum="Kinorhyncha" rank="species" species="toothlessi" status="sp. nov.">
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<emphasis id="3AE7AF78FFD9FF8118D324CCFE2FFF19" bold="true" box="[151,413,223,250]" italics="true" pageId="3" pageNumber="152">Mixtophyes toothlessi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="21D41203FFD9FF8119E024CCFE54FF1A" box="[420,486,223,249]" pageId="3" pageNumber="152" rank="species">sp. n.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="082C736AFFD9FF8118D32558FC85FE86" blockId="3.[151,1437,331,2014]" box="[151,823,331,358]" pageId="3" pageNumber="152">
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<emphasis id="3AE7AF78FFD9FF8118D32558FEA9FE86" bold="true" box="[151,283,331,357]" pageId="3" pageNumber="152">Synonymy.</emphasis>
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<taxonomicName id="CF9308E9FFD9FF8119662558FE6DFE86" box="[290,479,331,358]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="3" pageNumber="152" phylum="Kinorhyncha" rank="species" species="undefined-1">
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<emphasis id="3AE7AF78FFD9FF8119662558FE11FE86" box="[290,419,331,357]" italics="true" pageId="3" pageNumber="152">Mixtophyes</emphasis>
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sp. 1
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</taxonomicName>
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(in Sánchez
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<emphasis id="3AE7AF78FFD9FF811A34255FFD1AFE86" box="[624,680,331,357]" italics="true" pageId="3" pageNumber="152">et al.</emphasis>
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2019, 2022)
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<subSubSection id="408920E1FFD9FF811883257CFA28FDFA" pageId="3" pageNumber="152" type="materials_examined">
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<emphasis id="3AE7AF78FFD9FF811883257CFE01FE6A" bold="true" box="[199,435,367,393]" pageId="3" pageNumber="152">Material examined.</emphasis>
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<typeStatus id="D728CDC8FFD9FF8119FF257CFD94FE6A" box="[443,550,367,393]" pageId="3" pageNumber="152" type="holotype">Holotype</typeStatus>
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, adult male, collected on
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<date id="7C2D55AAFFD9FF811B0A257CFC7FFE69" box="[846,973,367,394]" pageId="3" pageNumber="152" value="2015-03-07">
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<collectingDate id="6C69AC42FFD9FF811B0A257CFC7FFE69" box="[846,973,367,394]" pageId="3" pageNumber="152" value="2015-03-07">07/03/2015</collectingDate>
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at the
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<collectorName id="A56616BCFFD9FF811C5A257CFA2FFE69" box="[1054,1437,367,394]" pageId="3" pageNumber="152">Clarion-Clipperton Fracture Zone</collectorName>
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(central-eastern Pacific Ocean),
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I contract area:
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,
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at
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depth
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; mounted in
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<collectorName id="A56616BCFFD9FF8118D325A4FEF7FE37" box="[151,325,439,468]" pageId="3" pageNumber="152">Fluoromount G</collectorName>
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®, deposited at
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<collectionCode id="6E82EBAFFFD9FF8119B125A4FD99FE32" box="[501,555,439,465]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="3" pageNumber="152" type="Museum">MfN</collectionCode>
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under accession number:
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<collectionCode id="6E82EBAFFFD9FF811B1425ABFC3FFE31" box="[848,909,440,466]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="152" type="Museum">ZMB</collectionCode>
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<specimenCode id="5835DB11FFD9FF811BD025ABFB90FE32" box="[916,1058,439,466]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="152" type="Museum">ZMB 12822</specimenCode>
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.
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, adult of indeterminate sex, collected on
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<date id="7C2D55AAFFD9FF811AB325C8FCC0FE15" box="[759,882,475,502]" pageId="3" pageNumber="152" value="2015-03-09">
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<collectingDate id="6C69AC42FFD9FF811AB325C8FCC0FE15" box="[759,882,475,502]" pageId="3" pageNumber="152" value="2015-03-09">09/03/2015</collectingDate>
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at the
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,
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<materialsCitation id="B8FB7937FFD9FF811C1D25CFFC02FDFA" ID-GBIF-Occurrence="4436800301" collectionCode="MfN" elevation="4255" latitude="12.596884" longLatPrecision="1" longitude="-116.4935" pageId="3" pageNumber="152" specimenCount="1" typeStatus="paratype">
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I contract area:
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at
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depth
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; mounted for SEM deposited at the
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<collectionCode id="6E82EBAFFFD9FF811B3F25ECFC02FDFA" box="[891,944,511,537]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="3" pageNumber="152" type="Museum">MfN</collectionCode>
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<materialsCitation id="B8FB7937FFD9FF811BF325ECFA24FDFA" ID-GBIF-Occurrence="4436800303" box="[951,1430,511,538]" collectionCode="ZMB" pageId="3" pageNumber="152" specimenCode="ZMB 12823" specimenCount="1" typeStatus="paratype">
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under accession number:
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<collectionCode id="6E82EBAFFFD9FF811C8F2613FAB4FDF9" box="[1227,1286,512,538]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="152" type="Museum">ZMB</collectionCode>
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<specimenCode id="5835DB11FFD9FF811D492613FA24FDFA" box="[1293,1430,511,538]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="152" type="Museum">ZMB 12823</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="408920E1FFD9FF8118832630FCF8FD2E" pageId="3" pageNumber="152" type="etymology">
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<paragraph id="082C736AFFD9FF8118832630FCF8FD2E" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="152">
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<emphasis id="3AE7AF78FFD9FF8118832630FEFCFDDE" bold="true" box="[199,334,547,573]" pageId="3" pageNumber="152">Etymology.</emphasis>
|
||
The species is named after Toothless, The Night Fury, the dragon of the novel series by Cressida Cowell “How to train your dragon” and the film with the same title from DreamWorks Animation, 2010. Several Kinorhyncha species have been described after famous dragons. The new species is reminiscent of Toothless for its middorsal spinose processes, similar to the back protuberances of the dragon, and for having one of the lateral terminal spines broken, like the tail in the animated character.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="408920E1FFD9FF81188326C4FD0CFC62" pageId="3" pageNumber="152" type="diagnosis">
|
||
<paragraph id="082C736AFFD9FF81188326C4FD0CFC62" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="152">
|
||
<emphasis id="3AE7AF78FFD9FF81188326C4FEF0FD12" bold="true" box="[199,322,727,753]" pageId="3" pageNumber="152">Diagnosis.</emphasis>
|
||
<taxonomicName id="CF9308E9FFD9FF81190326C4FE7AFD12" box="[327,456,727,753]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="3" pageNumber="152" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="3AE7AF78FFD9FF81190326C4FE7AFD12" box="[327,456,727,753]" italics="true" pageId="3" pageNumber="152">Mixtophyes</emphasis>
|
||
</taxonomicName>
|
||
with spinose processes on segments
|
||
<quantity id="CF6BDE8FFFD9FF811B1A26C4FC03FD12" box="[862,945,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="2.794" metricValueMin="0.254" pageId="3" pageNumber="152" unit="in" value="6.0" valueMax="11.0" valueMin="1.0">1–11 in</quantity>
|
||
middorsal position and on segments
|
||
<quantity id="CF6BDE8FFFD9FF811D0D26C4FA2EFD12" box="[1353,1436,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="2.54" metricValueMin="0.254" pageId="3" pageNumber="152" unit="in" value="5.5" valueMax="10.0" valueMin="1.0">1–10 in</quantity>
|
||
midlateral position, slightly increasing in length towards the posterior segments; paired setae in paradorsal position on segments 2, 4–6 and 8–9, and unpaired ones on segments 3 and 7. Paired setae in paralateral and ventrolateral positions on segments 2–9. Cuticular ornamentation as a reticulate pattern of net-like ridges present at the anterior margin of segment 1, both dorsally and ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="408920E1FFD9FF8A18832798FBF2F84C" lastPageId="8" lastPageNumber="157" pageId="3" pageNumber="152" type="description">
|
||
<paragraph id="082C736AFFD9FF8118832798FACAFC2A" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="152">
|
||
<emphasis id="3AE7AF78FFD9FF8118832798FEEBFC46" bold="true" box="[199,345,907,933]" pageId="3" pageNumber="152">Description.</emphasis>
|
||
<emphasis id="3AE7AF78FFD9FF8119252798FE2CFC46" box="[353,414,907,933]" italics="true" pageId="3" pageNumber="152">Head</emphasis>
|
||
. With retractable mouth cone and introvert. The specimens have both structures retracted inside the trunk, hence details on the morphology and distribution of oral styles and scalids cannot be provided.
|
||
</paragraph>
|
||
<paragraph id="082C736AFFD9FF8118D327C0FF59FBBA" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="152">
|
||
<emphasis id="3AE7AF78FFD9FF8118D327C0FF7DFC0E" box="[151,207,979,1005]" italics="true" pageId="3" pageNumber="152">Neck</emphasis>
|
||
. Four dorsal and three ventral placids. Dorsal placids equal in size, rectangular, robust and concave (ca. 30–34 µm wide and 18–20 µm high); ventral ones with different sizes: midventral placid robust, broad (ca. 53 µm wide and 14 µm high) lateroventral placids wider, weakly sclerotized (ca. 18–19 µm wide and 7–8 µm high) (
|
||
<figureCitation id="90A86FEFFFD9FF811D502008FA2AFBD6" box="[1300,1432,1051,1077]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF8118D3202CFF6CFBBA" box="[151,222,1087,1113]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="3" pageNumber="152">4C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFD9FF8118832070FB86F891" blockId="3.[151,1437,331,2014]" pageId="3" pageNumber="152">
|
||
<emphasis id="3AE7AF78FFD9FF8118832070FEBAFB9E" box="[199,264,1123,1149]" italics="true" pageId="3" pageNumber="152">Trunk</emphasis>
|
||
. With 11 segments. Segments 1 and 11 consisting of one tergal and one sternal plate; segments 2–9 with one tergal and two sternal plates (
|
||
<figureCitation id="90A86FEFFFD9FF811A512094FD29FB42" box="[533,667,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811AE32094FD42FB42" box="[679,752,1159,1185]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3A–H</figureCitation>
|
||
). Tergal plates slightly bulging middorsally, giving the trunk a triangular shape in cross-section. Maximum width at segment 6, progressively tapering at the last trunk segments but almost constant in width (
|
||
<figureCitation id="90A86FEFFFD9FF8119B420DCFDCBFB0A" box="[496,633,1231,1257]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811AC320DCFD7FFB0A" box="[647,717,1231,1257]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3A–H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811A9820DCFCB0FB0A" box="[732,770,1231,1257]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="3" pageNumber="152">4A</figureCitation>
|
||
). Segments 1–9 with middorsal spinose processes slightly increasing in length towards the posterior segments, surpassing the margin of the segment (
|
||
<figureCitation id="90A86FEFFFD9FF811CDA20E0FAB0FAEE" box="[1182,1282,1267,1293]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811D4B20E0FA2AFAEE" box="[1295,1432,1267,1294]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3A, C, E, G</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF8118D32104FF08FAD2" box="[151,186,1303,1329]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="3" pageNumber="152">4B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF8118832104FEE6FAD2" box="[199,340,1303,1330]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="3" pageNumber="152">5A, C, E, H</figureCitation>
|
||
). Hairy midlateral spinose processes present on segments 1–10 (
|
||
<figureCitation id="90A86FEFFFD9FF811C772104FB0BFAD2" box="[1075,1209,1303,1329]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811C822104FAFCFAD1" box="[1222,1358,1303,1330]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3B, D, F, H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811D1F2104FA2AFAD2" box="[1371,1432,1303,1329]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="3" pageNumber="152">4E–F</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF8118D32128FF6AFAB5" box="[151,216,1339,1366]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="3" pageNumber="152">5E–F</figureCitation>
|
||
). Spinose processes with marked notches at the free flap on both sides. No paradorsal intracuticular atria were detected. Minute, dot-shaped glandular cell outlets (cuticular pores in
|
||
<bibRefCitation id="6C020E9BFFD9FF811BEC214CFB37FA9A" author="Sanchez, N. & Pardos, F. & Sorensen, M. V." box="[936,1157,1375,1401]" pageId="3" pageNumber="152" pagination="221 - 239" refId="ref7036" refString="Sanchez, N., Pardos, F. & Sorensen, M. V. (2014) A new kinorhynch genus, Mixtophyes (Kinorhyncha: Homalorhagida), from the Guinea Basin deep-sea, with new data on the family Neocentrophyidae. Helgoland Marine Research, 68 (2), 221 - 239. https: // doi. org / 10.1007 / s 10152 - 014 - 0383 - 6" type="journal article" year="2014">
|
||
Sánchez
|
||
<emphasis id="3AE7AF78FFD9FF811C482173FBF7FA9A" box="[1036,1093,1375,1401]" italics="true" pageId="3" pageNumber="152">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
) as rounded perforations in several positions throughout the cuticle on segments 1–11 (
|
||
<figureCitation id="90A86FEFFFD9FF811B1D2190FC52FA7E" box="[857,992,1411,1437]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811BA92190FBA1FA7D" box="[1005,1043,1411,1438]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="3" pageNumber="152">5G</figureCitation>
|
||
). These glandular cells are usually distributed on segments 2–10 as one or two dorsal pores close to the base of the middorsal spinose processes, one or two pores in the middle region near the free flap, up to three pores around the dorsal muscular scars and up to three pores close to the paralateral setae, located anterior and posterior to them. Ventrally, these structures are distributed as two to four pores around the muscular scars and the middle region of the plates, plus two to three pores surrounding the most lateral sensory spots and one to two pores located over the ventrolateral setae. Segment 1 with a high number of pores distributed over the entire surface. Segment 11 with only two subdorsal pairs of pores and two pairs in the ventrolateral region, all of them near the free flap. Scale-like cuticular hairs as distributed across the trunk on segments 1–11, absent at the muscular scar areas. Muscular scars oval to bolus-shaped, present in laterodorsal and ventromedial positions on segments 1–10 (
|
||
<figureCitation id="90A86FEFFFD9FF811B1022D4FC63F902" box="[852,977,1735,1761]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Fig. 2A–B</figureCitation>
|
||
). Except on segment 1, the ventromedial muscular scars are close to the paraventral position. Longitudinal cuticular ridges in the anteriormost region of the plates on segments 2–11, less conspicuous in segment 2, across all over the segment. Pachycycli and ball and socket joints weakly developed. Primary pectinate fringe simple, with several short, pointed tips. Secondary pectinate fringe and apodemes absent.
|
||
<typeStatus id="D728CDC8FFD9FF8119992344FDF7F892" box="[477,581,1879,1905]" pageId="3" pageNumber="152" type="holotype">Holotype</typeStatus>
|
||
male with sperm in gonads until segment 7.
|
||
</paragraph>
|
||
<paragraph id="082C736AFFD9FF8618832368FDEEFE86" blockId="3.[151,1437,331,2014]" lastBlockId="4.[151,1437,150,357]" lastPageId="4" lastPageNumber="153" pageId="3" pageNumber="152">
|
||
Segment 1. Anterolateral margins of the tergal plate with horn-like extensions (
|
||
<figureCitation id="90A86FEFFFD9FF811C682368FB1DF876" box="[1068,1199,1915,1941]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811CFD2368FB6EF876" box="[1209,1244,1915,1941]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811CA22368FA9CF875" box="[1254,1326,1915,1942]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="3" pageNumber="152">5A–B</figureCitation>
|
||
).Anterior edge of both tergal and sternal plates denticulated, with a notorious reticulate pattern of cuticular wrinkles forming a net-like ornamentation across the anterior margin of the plates (
|
||
<figureCitation id="90A86FEFFFD9FF811BCE23D0FBA3F83E" box="[906,1041,1987,2013]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="3" pageNumber="152">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811C5B23D0FBD7F83E" box="[1055,1125,1987,2013]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="3" pageNumber="152">3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD9FF811C3623D0FB08F83D" box="[1138,1210,1987,2014]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="3" pageNumber="152">5A–B</figureCitation>
|
||
). Spinose processes in middorsal and midlateral positions, surpassing the posterior margin of the segment (
|
||
<figureCitation id="90A86FEFFFDEFF861CC72484FAB9FF52" box="[1155,1291,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861D5D2484FAEDFF52" box="[1305,1375,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861D292485FA3DFF52" box="[1389,1423,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="4" pageNumber="153">5E</figureCitation>
|
||
). Spinose processes of this and following segments consist of an elongate base ending in a flexible, acicular terminal tip, and with a longitudinal band of minute cuticular hairs in the basal part of the structure.
|
||
<typeStatus id="D728CDC8FFDEFF861CEA24CCFB54FF1A" box="[1198,1254,223,249]" pageId="4" pageNumber="153">Type</typeStatus>
|
||
1 sensory spots in subdorsal (two pairs), laterodorsal, paralateral and ventromedial (two pairs) positions. Sensory spots of this and following segments formed by numerous small papillae around a central pore or two pores (
|
||
<figureCitation id="90A86FEFFFDEFF861CDE2534FB4CFEA2" box="[1178,1278,295,321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="4" pageNumber="153">Figs. 4G</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861D4E2535FA9DFEA2" box="[1290,1327,294,321]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="4" pageNumber="153">5D</figureCitation>
|
||
). No cilia were observed emerging from the pores.
|
||
</paragraph>
|
||
<caption id="5CEC23E2FFDEFF8618DC2587FB1CFE16" ID-Table-UUID="5CEC23E2FFDEFF8618DC2587FB1CFE16" httpUri="http://table.plazi.org/id/5CEC23E2FFDEFF8618DC2587FB1CFE16" pageId="4" pageNumber="153" startId="4.[152,244,404,428]" targetBox="[166,981,516,1426]" targetIsTable="true" targetPageId="4" targetType="table">
|
||
<paragraph id="082C736AFFDEFF8618DC2587FB1CFE16" blockId="4.[151,1437,404,501]" pageId="4" pageNumber="153">
|
||
<emphasis id="3AE7AF78FFDEFF8618DC2587FEB5FE4F" bold="true" box="[152,263,404,428]" pageId="4" pageNumber="153">TABLE 1</emphasis>
|
||
. Measurements of male holotype (ZMB 12822) of
|
||
<taxonomicName id="CF9308E9FFDEFF861B622587FBBEFE4F" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" box="[806,1036,404,428]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="4" pageNumber="153" phylum="Kinorhyncha" rank="species" species="toothlessi" status="sp. nov.">
|
||
<emphasis id="3AE7AF78FFDEFF861B622587FBBEFE4F" box="[806,1036,404,428]" italics="true" pageId="4" pageNumber="153">Mixtophyes toothlessi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AE7AF78FFDEFF861C572586FBFCFE4E" bold="true" box="[1043,1102,405,429]" pageId="4" pageNumber="153">
|
||
<taxonomicNameLabel id="21D41203FFDEFF861C572586FBFCFE4E" box="[1043,1102,405,429]" pageId="4" pageNumber="153" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations: CL, cumulative length; LTS, lateral terminal spine; MDS, middorsal spine; MSW, maximum sternal width; S, segment; SW, standard sternal width; TL, total trunk length. Numbers after abbreviations indicate the respective segment.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="082C736AFFDEFF8618E22617FC04FA72" pageId="4" pageNumber="153">
|
||
<table id="7A9381CAFFDE007D18E22617FC67FA71" box="[166,981,516,1426]" gridcols="2" gridrows="23" pageId="4" pageNumber="153">
|
||
<tr id="B6A37128FFDE007D18E22617FC67FDFF" box="[166,981,516,540]" gridrow="0" pageId="4" pageNumber="153">
|
||
<th id="F5721854FFDE007D18E22617FEF5FDFF" box="[166,327,516,540]" gridcol="0" gridrow="0" pageId="4" pageNumber="153">
|
||
<emphasis id="3AE7AF78FFDEFF8618E22617FEA7FDFF" bold="true" box="[166,277,516,540]" pageId="4" pageNumber="153">Character</emphasis>
|
||
</th>
|
||
<th id="F5721854FFDE007D1B362617FC67FDFF" box="[882,981,516,540]" gridcol="1" gridrow="0" pageId="4" pageNumber="153">
|
||
<emphasis id="3AE7AF78FFDEFF861B362617FC67FDFF" bold="true" box="[882,981,516,540]" pageId="4" pageNumber="153">Holotype</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2263EFC67FDA6" box="[166,981,557,581]" gridrow="1" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2263EFEF5FDA6" box="[166,327,557,581]" gridcol="0" gridrow="1" pageId="4" pageNumber="153">TL (µm)</td>
|
||
<td id="F5721854FFDE007D1B36263EFC67FDA6" box="[882,981,557,581]" gridcol="1" gridrow="1" pageId="4" pageNumber="153">778</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22646FC67FD8E" box="[166,981,597,621]" gridrow="2" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22646FEF5FD8E" box="[166,327,597,621]" gridcol="0" gridrow="2" pageId="4" pageNumber="153">MSW-6 (µm)</td>
|
||
<td id="F5721854FFDE007D1B362646FC67FD8E" box="[882,981,597,621]" gridcol="1" gridrow="2" pageId="4" pageNumber="153">170</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2266EFC67FD76" box="[166,981,637,661]" gridrow="3" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2266EFEF5FD76" box="[166,327,637,661]" gridcol="0" gridrow="3" pageId="4" pageNumber="153">MSW-6/TL (%)</td>
|
||
<td id="F5721854FFDE007D1B36266EFC67FD76" box="[882,981,637,661]" gridcol="1" gridrow="3" pageId="4" pageNumber="153">21,81</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E226B6FC67FD5E" box="[166,981,677,701]" gridrow="4" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E226B6FEF5FD5E" box="[166,327,677,701]" gridcol="0" gridrow="4" pageId="4" pageNumber="153">SW-10 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3626B6FC67FD5E" box="[882,981,677,701]" gridcol="1" gridrow="4" pageId="4" pageNumber="153">-</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E226DDFC67FD05" box="[166,981,718,742]" gridrow="5" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E226DDFEF5FD05" box="[166,327,718,742]" gridcol="0" gridrow="5" pageId="4" pageNumber="153">SW-10/TL (%)</td>
|
||
<td id="F5721854FFDE007D1B3626DDFC67FD05" box="[882,981,718,742]" gridcol="1" gridrow="5" pageId="4" pageNumber="153">-</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E226E5FC67FCED" box="[166,981,758,782]" gridrow="6" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E226E5FEF5FCED" box="[166,327,758,782]" gridcol="0" gridrow="6" pageId="4" pageNumber="153">S1 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3626E5FC67FCED" box="[882,981,758,782]" gridcol="1" gridrow="6" pageId="4" pageNumber="153">128</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2270DFC67FCD5" box="[166,981,798,822]" gridrow="7" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2270DFEF5FCD5" box="[166,327,798,822]" gridcol="0" gridrow="7" pageId="4" pageNumber="153">S2 (µm)</td>
|
||
<td id="F5721854FFDE007D1B36270DFC67FCD5" box="[882,981,798,822]" gridcol="1" gridrow="7" pageId="4" pageNumber="153">64</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22755FC67FCBD" box="[166,981,838,862]" gridrow="8" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22755FEF5FCBD" box="[166,327,838,862]" gridcol="0" gridrow="8" pageId="4" pageNumber="153">S3 (µm)</td>
|
||
<td id="F5721854FFDE007D1B362755FC67FCBD" box="[882,981,838,862]" gridcol="1" gridrow="8" pageId="4" pageNumber="153">62</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2277CFC67FC64" box="[166,981,879,903]" gridrow="9" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2277CFEF5FC64" box="[166,327,879,903]" gridcol="0" gridrow="9" pageId="4" pageNumber="153">S4 (µm)</td>
|
||
<td id="F5721854FFDE007D1B36277CFC67FC64" box="[882,981,879,903]" gridcol="1" gridrow="9" pageId="4" pageNumber="153">73</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22784FC67FC4C" box="[166,981,919,943]" gridrow="10" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22784FEF5FC4C" box="[166,327,919,943]" gridcol="0" gridrow="10" pageId="4" pageNumber="153">S5 (µm)</td>
|
||
<td id="F5721854FFDE007D1B362784FC67FC4C" box="[882,981,919,943]" gridcol="1" gridrow="10" pageId="4" pageNumber="153">78</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E227ACFC67FC34" box="[166,981,959,983]" gridrow="11" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E227ACFEF5FC34" box="[166,327,959,983]" gridcol="0" gridrow="11" pageId="4" pageNumber="153">S6 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3627ACFC67FC34" box="[882,981,959,983]" gridcol="1" gridrow="11" pageId="4" pageNumber="153">79</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E227F4FC67FC1C" box="[166,981,999,1023]" gridrow="12" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E227F4FEF5FC1C" box="[166,327,999,1023]" gridcol="0" gridrow="12" pageId="4" pageNumber="153">S7 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3627F4FC67FC1C" box="[882,981,999,1023]" gridcol="1" gridrow="12" pageId="4" pageNumber="153">87</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22003FC67FBCB" box="[166,981,1040,1064]" gridrow="13" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22003FEF5FBCB" box="[166,327,1040,1064]" gridcol="0" gridrow="13" pageId="4" pageNumber="153">S8 (µm)</td>
|
||
<td id="F5721854FFDE007D1B362003FC67FBCB" box="[882,981,1040,1064]" gridcol="1" gridrow="13" pageId="4" pageNumber="153">88</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2202BFC67FBB3" box="[166,981,1080,1104]" gridrow="14" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2202BFEF5FBB3" box="[166,327,1080,1104]" gridcol="0" gridrow="14" pageId="4" pageNumber="153">S9 (µm)</td>
|
||
<td id="F5721854FFDE007D1B36202BFC67FBB3" box="[882,981,1080,1104]" gridcol="1" gridrow="14" pageId="4" pageNumber="153">83</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22073FC67FB9B" box="[166,981,1120,1144]" gridrow="15" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22073FEF5FB9B" box="[166,327,1120,1144]" gridcol="0" gridrow="15" pageId="4" pageNumber="153">S10 (µm)</td>
|
||
<td id="F5721854FFDE007D1B362073FC67FB9B" box="[882,981,1120,1144]" gridcol="1" gridrow="15" pageId="4" pageNumber="153">68</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2209BFC67FB43" box="[166,981,1160,1184]" gridrow="16" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2209BFEF5FB43" box="[166,327,1160,1184]" gridcol="0" gridrow="16" pageId="4" pageNumber="153">S11 (µm)</td>
|
||
<td id="F5721854FFDE007D1B36209BFC67FB43" box="[882,981,1160,1184]" gridcol="1" gridrow="16" pageId="4" pageNumber="153">57</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E220A3FC67FB2A" box="[166,981,1200,1225]" gridrow="17" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E220A3FEF5FB2A" box="[166,327,1200,1225]" gridcol="0" gridrow="17" pageId="4" pageNumber="153">MDS10 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3620A3FC67FB2A" box="[882,981,1200,1225]" gridcol="1" gridrow="17" pageId="4" pageNumber="153">116</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E220CAFC67FB12" box="[166,981,1241,1265]" gridrow="18" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E220CAFEF5FB12" box="[166,327,1241,1265]" gridcol="0" gridrow="18" pageId="4" pageNumber="153">MDS11 (µm)</td>
|
||
<td id="F5721854FFDE007D1B3620CAFC67FB12" box="[882,981,1241,1265]" gridcol="1" gridrow="18" pageId="4" pageNumber="153">-</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22112FC67FAFA" box="[166,981,1281,1305]" gridrow="19" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22112FEF5FAFA" box="[166,327,1281,1305]" gridcol="0" gridrow="19" pageId="4" pageNumber="153">LTS (µm)</td>
|
||
<td id="F5721854FFDE007D1B362112FC67FAFA" box="[882,981,1281,1305]" gridcol="1" gridrow="19" pageId="4" pageNumber="153">483</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2213AFC67FAA2" box="[166,981,1321,1345]" gridrow="20" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2213AFEF5FAA2" box="[166,327,1321,1345]" gridcol="0" gridrow="20" pageId="4" pageNumber="153">LTS/TL (%)</td>
|
||
<td id="F5721854FFDE007D1B36213AFC67FAA2" box="[882,981,1321,1345]" gridcol="1" gridrow="20" pageId="4" pageNumber="153">62,10</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E22142FC67FA8A" box="[166,981,1361,1385]" gridrow="21" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E22142FEF5FA8A" box="[166,327,1361,1385]" gridcol="0" gridrow="21" pageId="4" pageNumber="153">CL (µm)</td>
|
||
<td id="F5721854FFDE007D1B362142FC67FA8A" box="[882,981,1361,1385]" gridcol="1" gridrow="21" pageId="4" pageNumber="153">867</td>
|
||
</tr>
|
||
<tr id="B6A37128FFDE007D18E2216AFC67FA71" box="[166,981,1401,1426]" gridrow="22" pageId="4" pageNumber="153">
|
||
<td id="F5721854FFDE007D18E2216AFEF5FA71" box="[166,327,1401,1426]" gridcol="0" gridrow="22" pageId="4" pageNumber="153">CL/TL (%)</td>
|
||
<td id="F5721854FFDE007D1B36216AFC67FA71" box="[882,981,1401,1426]" gridcol="1" gridrow="22" pageId="4" pageNumber="153">111,52</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDEFF86188321D2FBC6F9C7" blockId="4.[151,1437,1473,2004]" pageId="4" pageNumber="153">
|
||
Segment 2. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs in the
|
||
<typeStatus id="D728CDC8FFDEFF861C5721F6FBCAFA1C" box="[1043,1144,1509,1535]" pageId="4" pageNumber="153" type="holotype">holotype</typeStatus>
|
||
, one pair in the
|
||
<typeStatus id="D728CDC8FFDEFF861D6821F6FA23FA1C" box="[1324,1425,1509,1535]" pageId="4" pageNumber="153" type="paratype">paratype</typeStatus>
|
||
), laterodorsal (two pairs) and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDEFF861B27221AFC5AF9C0" box="[867,1000,1545,1571]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861BB7221AFB8BF9C0" box="[1011,1081,1545,1571]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861C00221AFBD4F9C7" box="[1092,1126,1545,1572]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="4" pageNumber="153">5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDEFF861883223EFC8AF973" blockId="4.[151,1437,1473,2004]" pageId="4" pageNumber="153">
|
||
Segment 3. Spinose processes in middorsal and midlateral positions. Unpaired, paradorsal seta and paired setae in paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDEFF861A632266FD1EF96C" box="[551,684,1653,1679]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861AFC2266FD4FF96C" box="[696,765,1653,1679]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF861B4D2266FC99F973" box="[777,811,1653,1680]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="4" pageNumber="153">5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDEFF861883228AFDACF918" blockId="4.[151,1437,1473,2004]" pageId="4" pageNumber="153">
|
||
Segment 4. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDEFF86194822F2FE22F918" box="[268,400,1761,1787]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF8619D822F2FE53F918" box="[412,481,1761,1787]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF8619A922F2FDA3F918" box="[493,529,1761,1787]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="4" pageNumber="153">4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDEFF8618832316FE5DF884" blockId="4.[151,1437,1473,2004]" pageId="4" pageNumber="153">
|
||
Segment 5. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDEFF861948235EFE22F884" box="[268,400,1869,1895]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF8619D8235EFE51F884" box="[412,483,1869,1895]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDEFF8618832362FC18F830" blockId="4.[151,1437,1473,2004]" pageId="4" pageNumber="153">
|
||
Segment 6. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDEFF8618DB23AAFE96F830" box="[159,292,1977,2003]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="4" pageNumber="153">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDEFF86197423AAFEC2F830" box="[304,368,1977,2003]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="4" pageNumber="153">3E–F</figureCitation>
|
||
). Paradorsal sensory spots could not be confirmed.
|
||
</paragraph>
|
||
<caption id="5CEC23E2FFDFFF8718D322A3FD1AF89E" ID-DOI="http://doi.org/10.5281/zenodo.7935925" ID-Zenodo-Dep="7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="5" pageNumber="154" startId="5.[151,250,1712,1736]" targetBox="[155,1433,184,1686]" targetPageId="5" targetType="figure">
|
||
<paragraph id="082C736AFFDFFF8718D322A3FD1AF89E" blockId="5.[151,1437,1712,1917]" pageId="5" pageNumber="154">
|
||
<emphasis id="3AE7AF78FFDFFF8718D322A3FEA3F92A" bold="true" box="[151,273,1712,1737]" pageId="5" pageNumber="154">FIGURE 2.</emphasis>
|
||
Line art illustrations of adults of
|
||
<taxonomicName id="CF9308E9FFDFFF871A1F22A2FCF0F92A" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" box="[603,834,1713,1737]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="5" pageNumber="154" phylum="Kinorhyncha" rank="species" species="thoothlessi" status="sp. nov.">
|
||
<emphasis id="3AE7AF78FFDFFF871A1F22A2FCF0F92A" box="[603,834,1713,1737]" italics="true" pageId="5" pageNumber="154">Mixtophyes thoothlessi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AE7AF78FFDFFF871B0322A2FCCCF92A" bold="true" box="[839,894,1713,1737]" pageId="5" pageNumber="154">
|
||
<taxonomicNameLabel id="21D41203FFDFFF871B0322A2FCCCF92A" box="[839,894,1713,1737]" pageId="5" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length. Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5CEC23E2FFDCFF8418D32337FAECF812" ID-DOI="http://doi.org/10.5281/zenodo.7935929" ID-Zenodo-Dep="7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="6" pageNumber="155" startId="6.[151,250,1828,1852]" targetBox="[280,1307,186,1799]" targetPageId="6" targetType="figure">
|
||
<paragraph id="082C736AFFDCFF8418D32337FAECF812" blockId="6.[151,1437,1828,2033]" pageId="6" pageNumber="155">
|
||
<emphasis id="3AE7AF78FFDCFF8418D32337FEA1F8DF" bold="true" box="[151,275,1828,1852]" pageId="6" pageNumber="155">FIGURE 3.</emphasis>
|
||
Light micrographs of male holotype of
|
||
<taxonomicName id="CF9308E9FFDCFF841AE82337FC3AF8DF" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" box="[684,904,1828,1852]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="6" pageNumber="155" phylum="Kinorhyncha" rank="species" species="toothlessi" status="sp. nov.">
|
||
<emphasis id="3AE7AF78FFDCFF841AE82337FC3AF8DF" box="[684,904,1828,1852]" italics="true" pageId="6" pageNumber="155">Mixtophyes toothlessi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AE7AF78FFDCFF841BCB2336FC7BF8DE" bold="true" box="[911,969,1829,1853]" pageId="6" pageNumber="155">
|
||
<taxonomicNameLabel id="21D41203FFDCFF841BCB2336FC7BF8DE" box="[911,969,1829,1853]" pageId="6" pageNumber="155" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A–H: 20 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="082C736AFFDDFF8518832484FCB7FF1A" blockId="7.[151,1437,151,465]" pageId="7" pageNumber="156">
|
||
Segment 7. Spinose processes in middorsal and midlateral positions. Unpaired, paradorsal seta and paired setae in paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDDFF851A6324CCFD1EFF1A" box="[551,684,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="7" pageNumber="156">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF851AFC24CCFD4AFF1A" box="[696,760,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="7" pageNumber="156">3E–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDDFF8518832510FD32FE86" blockId="7.[151,1437,151,465]" pageId="7" pageNumber="156">
|
||
Segment 8. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) positions (
|
||
<figureCitation id="90A86FEFFFDDFF8519482558FE22FE86" box="[268,400,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="7" pageNumber="156">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF8519D82558FE6BFE86" box="[412,473,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="7" pageNumber="156">3E–F</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF8519A02558FD90FE86" box="[484,546,331,357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="7" pageNumber="156">4E–F</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF851A692559FDC1FE86" box="[557,627,330,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="7" pageNumber="156">5C, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFDDFF851883257CFDCDFE32" blockId="7.[151,1437,151,465]" pageId="7" pageNumber="156">
|
||
Segment 9. Spinose processes in middorsal and midlateral positions. Paired setae in paradorsal, paralateral and ventrolateral positions. Sensory spots in paradorsal, subdorsal (two pairs), laterodorsal and ventromedial (two pairs) position (
|
||
<figureCitation id="90A86FEFFFDDFF85194525A4FE37FE32" box="[257,389,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="7" pageNumber="156">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF8519D525A4FE6AFE32" box="[401,472,439,465]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="7" pageNumber="156">3G–H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF8519A725A4FD93FE32" box="[483,545,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="7" pageNumber="156">4E–F</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFDDFF851A6825A5FDC0FE32" box="[556,626,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="7" pageNumber="156">5C, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="5CEC23E2FFDDFF8518D322A0FB8DF80F" ID-DOI="http://doi.org/10.5281/zenodo.7935935" ID-Zenodo-Dep="7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="7" pageNumber="156" startId="7.[151,250,1715,1739]" targetBox="[151,1436,490,1691]" targetPageId="7" targetType="figure">
|
||
<paragraph id="082C736AFFDDFF8518D322A0FB8DF80F" blockId="7.[151,1437,1715,2028]" pageId="7" pageNumber="156">
|
||
<emphasis id="3AE7AF78FFDDFF8518D322A0FEA6F928" bold="true" box="[151,276,1715,1739]" pageId="7" pageNumber="156">FIGURE 4.</emphasis>
|
||
Light micrographs of male holotype of
|
||
<taxonomicName id="CF9308E9FFDDFF851AF622A0FC22F928" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" box="[690,912,1715,1739]" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="7" pageNumber="156" phylum="Kinorhyncha" rank="species" species="toothlessi" status="sp. nov.">
|
||
<emphasis id="3AE7AF78FFDDFF851AF622A0FC22F928" box="[690,912,1715,1739]" italics="true" pageId="7" pageNumber="156">Mixtophyes toothlessi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AE7AF78FFDDFF851BD322A0FC63F928" bold="true" box="[919,977,1715,1739]" pageId="7" pageNumber="156">
|
||
<taxonomicNameLabel id="21D41203FFDDFF851BD322A0FC63F928" box="[919,977,1715,1739]" pageId="7" pageNumber="156" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5CEC23E2FFD2FF8A18D3206CFCB0FA93" ID-DOI="http://doi.org/10.5281/zenodo.7935939" ID-Zenodo-Dep="7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="8" pageNumber="157" startId="8.[151,250,1151,1175]" targetBox="[151,1436,181,1126]" targetPageId="8" targetType="figure">
|
||
<paragraph id="082C736AFFD2FF8A18D3206CFCB0FA93" blockId="8.[151,1437,1151,1392]" pageId="8" pageNumber="157">
|
||
<emphasis id="3AE7AF78FFD2FF8A18D3206CFEA4FB74" bold="true" box="[151,278,1151,1175]" pageId="8" pageNumber="157">FIGURE 5.</emphasis>
|
||
Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of
|
||
<taxonomicName id="CF9308E9FFD2FF8A1D63206CFF4AFB58" authority="González-Casarrubios & Arbizu & Sánchez, 2023" authorityName="González-Casarrubios & Arbizu & Sánchez" authorityYear="2023" class="Allomalorhagida" family="Neocentrophyidae" genus="Mixtophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="8" pageNumber="157" phylum="Kinorhyncha" rank="species" species="toothlessi" status="sp. nov.">
|
||
<emphasis id="3AE7AF78FFD2FF8A1D63206CFF4AFB58" italics="true" pageId="8" pageNumber="157">Mixtophyes toothlessi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3AE7AF78FFD2FF8A18BA20B0FE84FB58" bold="true" box="[254,310,1187,1211]" pageId="8" pageNumber="157">
|
||
<taxonomicNameLabel id="21D41203FFD2FF8A18BA20B0FE84FB58" box="[254,310,1187,1211]" pageId="8" pageNumber="157" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="082C736AFFD2FF8A1883218DFE69F9AB" blockId="8.[151,1437,1437,1968]" pageId="8" pageNumber="157">
|
||
Segment 10. Middorsal spinose process whose base ends in the middle region of the plate. This structure has a pointed, thin free region, more flexible than the ones on the preceding segments Spinose processes in midlateral position, surpassing the margin of the following segment. Two pairs of sensory spots in subdorsal and ventromedial positions. Straight posterior margin of the segment, without notches in the middle region of the tergal plate (
|
||
<figureCitation id="90A86FEFFFD2FF8A1D20221AFF4DF9AB" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="8" pageNumber="157">Figs. 2A–B, D</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A194E223DFEE0F9AB" box="[266,338,1582,1608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="8" pageNumber="157">3G–H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A1919223DFE30F9AB" box="[349,386,1582,1608]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="8" pageNumber="157">4H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A19C9223EFE7CF9AB" box="[397,462,1581,1608]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="8" pageNumber="157">5F, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="082C736AFFD2FF8A18832241FBF2F84C" blockId="8.[151,1437,1437,1968]" pageId="8" pageNumber="157">
|
||
Segment 11. Middorsal spinose process emerging from the posterior region of the segment. Midlateral spinose processes absent. Small, midterminal process and minute, conical spines present in ventrolateral position, both structures only confirmed in the LM specimen (
|
||
<figureCitation id="90A86FEFFFD2FF8A1AEF228AFC9DF957" box="[683,815,1689,1716]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="8" pageNumber="157">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A1B7E2289FCECF957" box="[826,862,1690,1716]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="8" pageNumber="157">3H</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A1B2E2289FC34F957" box="[874,902,1690,1716]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="8" pageNumber="157">4J</figureCitation>
|
||
). Paired sensory spots in ventromedial position;
|
||
<typeStatus id="D728CDC8FFD2FF8A18D322ADFF75F93B" box="[151,199,1726,1752]" pageId="8" pageNumber="157">type</typeStatus>
|
||
3 sensory spots present in subdorsal and ventromedial positions (
|
||
<figureCitation id="90A86FEFFFD2FF8A1BEA22AEFBA1F93B" box="[942,1043,1725,1752]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="8" pageNumber="157">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A1C5B22AEFB89F93B" box="[1055,1083,1725,1752]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="8" pageNumber="157">5J</figureCitation>
|
||
). Posterior margin of the tergal plate with subdorsal and laterodorsal notches together with a pair of extensions over the lateral terminal spines. Sternal plates with straight posterior margin and a pair of extensions over the lateral terminal spines (
|
||
<figureCitation id="90A86FEFFFD2FF8A1D562316FA2AF8C3" box="[1298,1432,1797,1824]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="8" pageNumber="157">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="90A86FEFFFD2FF8A18D32339FF6DF8A7" box="[151,223,1834,1860]" captionStart="FIGURE 3" captionStartId="6.[151,250,1828,1852]" captionTargetBox="[280,1307,186,1799]" captionTargetId="figure-28@6.[275,1312,181,1804]" captionTargetPageId="6" captionText="FIGURE 3. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing the main trunk characters. A. Dorsal view of segments 1–2. B. Ventral view of segments 1–2. C. Dorsal view of segments 3–5. D. Ventral view of segments 3–5. E. Dorsal view of segments 6–8. F. Ventral view of segments 6–8. G. Dorsal view of segments 9–11. H. Ventral view of segments 9–11. Abbreviations: lts, lateral terminal spine; mdp, middorsal spinose process; mlp, midlateral spinose process; msp, minute spine; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars:A–H: 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7935929" httpUri="https://zenodo.org/record/7935929/files/figure.png" pageId="8" pageNumber="157">3G–H</figureCitation>
|
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,
|
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<figureCitation id="90A86FEFFFD2FF8A18AD2339FEB0F8A7" box="[233,258,1834,1860]" captionStart="FIGURE 4" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[151,1436,490,1691]" captionTargetId="figure-127@7.[151,1437,489,1692]" captionTargetPageId="7" captionText="FIGURE 4. Light micrographs of male holotype of Mixtophyes toothlessi sp. n. showing trunk overviews and detailed trunk characters. A. Dorsal overview. B. Detail of dorsal spinose process and setae of segment 4. C. Detail on dorsal placids. D. Detail on ventral placis. E. Detail of lateral position of segment 8 showing the paralateral seta. F. Detail of lateral position of segment 8 showing paralateral and ventrolateral setae. G. Close-up of a sensory spot and a glandular cell outlet of segment 3. H. Detail of the middorsal spinose process of segment 10. I. Detailed dorsal view of segment 11. J. Detail on the midterminal process of segment 11. Abbreviations: gco, glandular cell outlet; lts, lateral terminal spine; mdp, middorsal spinose process; mds, middorsal spine; mlp, midlateral spinose process; mtp, midterminal process; pdse, paradorsal seta; plse, paralateral seta; ss, sensory spot; ss3, type 3 sensory spot; vlse, ventrolateral seta. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; C–D: 20 μm; B, E–J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7935935" httpUri="https://zenodo.org/record/7935935/files/figure.png" pageId="8" pageNumber="157">4I</figureCitation>
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,
|
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<figureCitation id="90A86FEFFFD2FF8A1949233AFEFFF8A7" box="[269,333,1833,1860]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="8" pageNumber="157">5F, H</figureCitation>
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). Long, slender lateral terminal spines (LTS:TL: 62 %) (
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<figureCitation id="90A86FEFFFD2FF8A1BFF233AFBA1F8A7" box="[955,1043,1833,1860]" captionStart="FIGURE 2" captionStartId="5.[151,250,1712,1736]" captionTargetBox="[155,1433,184,1686]" captionTargetId="figure-19@5.[151,1436,181,1689]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations of adults of Mixtophyes thoothlessi sp. n. A. Male, ventral view. B. Male, dorsal view. C. Dorsal overview showing the LTS length.Abbreviations: co, cuticular ornamentation; cr, cuticular ridge; gco, glandular cell outlet; mlp, midlateral spinose process; ldss, laterodorsal sensory spot; lts, lateral terminal spine; mdp, middorsal process; mlp, midlateral process; ms, muscular scar; msp, minute spine; mtp, midterminal process; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; sdss, subdorsal sensory spot; ss3, type 3 sensory spot; tsj, tergosternal junction; vlse, ventrolateral seta; vmss, ventromedial sensory spot. Scalebar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7935925" httpUri="https://zenodo.org/record/7935925/files/figure.png" pageId="8" pageNumber="157">Fig. 2C</figureCitation>
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). The SEM specimen showed some elongated, rounded structures on the sternal plates of segments 10 and 11 (
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||
<figureCitation id="90A86FEFFFD2FF8A1BB4235EFBF2F88B" box="[1008,1088,1869,1896]" captionStart="FIGURE 5" captionStartId="8.[151,250,1151,1175]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure-28@8.[151,1436,181,1127]" captionTargetPageId="8" captionText="FIGURE 5. Scanning electron micrographs of Scanning electron micrographs of adult paratype of paratype of Mixtophyes toothlessi sp. n. A. Lateral overview. B. Detail on the anterior region of segment 1. C. Detail of the middorsal spinose process of segment 9. D. Detail of a sensory spot. E. View of the right half of the tergal plates of segments 1–3. F. Dorsal view of segments 10–11. G. Glandular cell outlet. H. View of the right half of the tergal plates of segments 10–11. I. Putative bacterial epibionts. J. Type 3 sensory spots of segment 11. Abbreviations: lts, lateral terminal spines; mdp, middorsal spinose process; mlp, midlateral spinose process. Numbers after abbreviations indicate the corresponding segment. Sensory spots marked as dashed circles. Scalebars: A: 100 μm; B–C, E–F, H: 10 μm; D, G, I–J: 1 μm." figureDoi="http://doi.org/10.5281/zenodo.7935939" httpUri="https://zenodo.org/record/7935939/files/figure.png" pageId="8" pageNumber="157">Fig. 5I</figureCitation>
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). These structures could be an artefact of the fixative or a result of the critical point, being a cluster of crystals, but the option that they could be small outcrops of epibiont bacilli or even some kind of fungus cannot be dismissed.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |