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<document id="87E19A72CDADDFEFEC61603800B476C9" ID-CLB-Dataset="45907" ID-DOI="10.5281/zenodo.279290" ID-GBIF-Dataset="bfde6d67-dd00-40e7-a7de-80fad78042d5" ID-ISSN="1175-5326" ID-Zenodo-Dep="279290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413659433" checkinUser="plazi" docAuthor="Lapolla, John S., Hawkes, Peter G. &amp; Fisher, Brian L." docDate="2011" docId="773B4C70FFF4C135FF56FE49FF4AFA08" docLanguage="en" docName="zt03110p036.pdf" docOrigin="Zootaxa 3110" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nylanderia umbella LaPolla and Fisher, sp. nov." docType="treatment" docVersion="12" lastPageNumber="24" masterDocId="8B023408FFFAC13BFFC1FFA3FF81FFC2" masterDocTitle="Monograph of Nylanderia (Hymenoptera: Formicidae) of the World, Part I: Nylanderia in the Afrotropics" masterLastPageNumber="36" masterPageNumber="10" pageNumber="24" updateTime="1698328294110" updateUser="ExternalLinkService">
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<mods:title id="AC935BA9354867E9F8938E0C76397876">Monograph of Nylanderia (Hymenoptera: Formicidae) of the World, Part I: Nylanderia in the Afrotropics</mods:title>
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<mods:namePart id="A66164B4A0C72F9E94CA9279B40D4A98">Lapolla, John S.</mods:namePart>
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<mods:namePart id="18BE713E53D5C5A359B832D67AF8E351">Hawkes, Peter G.</mods:namePart>
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<mods:namePart id="2247789E35F86A45EBBFB062AABF9DAC">Fisher, Brian L.</mods:namePart>
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<mods:date id="C627A8EAE79DC53A1521E07DF0A6A6B6">2011</mods:date>
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<treatment id="773B4C70FFF4C135FF56FE49FF4AFA08" ID-DOI="http://doi.org/10.5281/zenodo.6190839" ID-GBIF-Taxon="119424248" ID-Zenodo-Dep="6190839" LSID="urn:lsid:plazi:treatment:773B4C70FFF4C135FF56FE49FF4AFA08" httpUri="http://treatment.plazi.org/id/773B4C70FFF4C135FF56FE49FF4AFA08" lastPageNumber="24" pageId="14" pageNumber="24">
<subSubSection id="B788AEEDFFF4C135FF56FE49FE40FDE7" pageId="14" pageNumber="24" type="nomenclature">
<paragraph id="FF2DFD66FFF4C135FF56FE49FD69FDC6" blockId="14.[151,744,490,549]" box="[151,744,490,516]" pageId="14" pageNumber="24">
<heading id="A4654A0AFFF4C135FF56FE49FD69FDC6" bold="true" box="[151,744,490,516]" fontSize="11" level="1" pageId="14" pageNumber="24" reason="1">
<emphasis id="CDE62174FFF4C135FF56FE49FD69FDC6" bold="true" box="[151,744,490,516]" pageId="14" pageNumber="24">
<taxonomicName id="389286E5FFF4C135FF56FE49FDFDFDC6" authority="LaPolla and Fisher" authorityName="LaPolla and Fisher" box="[151,636,490,516]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="umbella" status="sp. nov.">
<emphasis id="CDE62174FFF4C135FF56FE49FE06FDC6" bold="true" box="[151,391,490,516]" italics="true" pageId="14" pageNumber="24">Nylanderia umbella</emphasis>
LaPolla and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="D6D59C0FFFF4C135FD4AFE49FD69FDC6" box="[651,744,490,516]" pageId="14" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF56FDAEFE40FDE7" blockId="14.[151,744,490,549]" box="[151,449,525,549]" pageId="14" pageNumber="24">
(
<figureCitation id="67A9E1E3FFF4C135FF5EFDAEFEA6FDE7" box="[159,295,525,549]" captionStart="FIGURES 46 60" captionStartId="19.[151,264,1883,1906]" captionTargetBox="[167,1436,193,1834]" captionTargetId="figure@19.[151,1436,193,1834]" captionTargetPageId="19" captionText="FIGURES 46 60. Afrotropical Nylanderia species in various views: scintilla (worker, 46 48, CASENT 0179579, Ivory Coast); silvula (worker, 49 51, LACMENT 272717; queen 52 54, LACMENT 272721; male, 55 57, LACMENT 272719; all from Kenya); umbella (worker, 58 60, CASENT 0179583, Cameroon)" httpUri="https://zenodo.org/record/279294/files/figure.png" pageId="14" pageNumber="24">Figs. 5860</figureCitation>
,
<figureCitation id="67A9E1E3FFF4C135FEF3FDAEFECFFDE7" box="[306,334,525,549]" captionStart="FIGURES 73 88" captionStartId="21.[151,264,1317,1340]" captionTargetBox="[151,1436,195,1286]" captionTargetId="figure@21.[151,1436,195,1295]" captionTargetPageId="21" captionText="FIGURES 73 88. Scapes of Afrotropical Nylanderia species (workers only): boltoni (73, CASENT 0179580, Gabon); bourbonica (74, CASENT 0101309, Reunion); brevisetula (75, CASENT 0090304, Central African Republic); impolita (76, CASENT 0179589, Gabon); incallida (77, CASENT 0179576, Ivory Coast); jaegerskioeldi (78, CASENT 0179577, Kenya); lepida (79, CASENT 0179591, Gabon); luteafra (80, CASENT 0179584, Gabon); mendica (81, CASENT 0179590, Gabon); natalensis (82, CASENT 0396096, South Africa); scintilla (83, CASENT 0179579, Ivory Coast); silvula (84, LACMENT 272717, Kenya); umbella (85, CASENT 0179583, Cameroon); usambarica (86, CASENT 0235343, Tanzania); vaga (87, CASENT 0104253, U. S. A); vividula (88, CASENT 0103260, U. S. A.)" httpUri="https://zenodo.org/record/279296/files/figure.png" pageId="14" pageNumber="24">85</figureCitation>
[worker])
</paragraph>
</subSubSection>
<subSubSection id="B788AEEDFFF4C135FF56FDECFEE6FD48" pageId="14" pageNumber="24" type="materials_examined">
<paragraph id="FF2DFD66FFF4C135FF56FDECFEE6FD48" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
<typeStatus id="202943C4FFF4C135FF56FDECFF7EFDA5" box="[151,255,591,615]" pageId="14" pageNumber="24" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="8785BDF6FFF4C135FEA5FDECFD88FDA5" box="[356,521,591,615]" name="Cameroon" pageId="14" pageNumber="24">CAMEROON</collectingCountry>
: Mbalmayo,
<date id="8B2CDBA6FFF4C135FD5DFDECFD71FDA5" box="[668,752,591,615]" pageId="14" pageNumber="24" value="1993-11">xi.1993</date>
(N. Stork) (
<collectionCode id="998365A3FFF4C135FCBBFDECFC55FDA5" box="[890,980,591,615]" pageId="14" pageNumber="24">BMNH</collectionCode>
); 3
<typeStatus id="202943C4FFF4C135FC3FFDF3FBDFFDAA" box="[1022,1118,592,616]" pageId="14" pageNumber="24" type="paratype">paratype</typeStatus>
workers, same label data as
<typeStatus id="202943C4FFF4C135FF56FDD1FF78FD48" box="[151,249,626,650]" pageId="14" pageNumber="24" type="holotype">holotype</typeStatus>
(
<collectionCode id="998365A3FFF4C135FEC6FDD1FEDEFD48" box="[263,351,626,650]" pageId="14" pageNumber="24">BMNH</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="B788AEEDFFF4C135FF06FD36FC2FFB15" pageId="14" pageNumber="24" type="description">
<paragraph id="FF2DFD66FFF4C135FF06FD36FECFFD30" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF06FD36FE29FD6C" bold="true" box="[199,424,661,686]" pageId="14" pageNumber="24">Worker diagnosis:</emphasis>
Head width less than
<quantity id="386A5083FFF4C135FD79FD36FCA5FD6F" box="[696,804,661,685]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="14" pageNumber="24" unit="mm" value="0.51">0.51 mm</quantity>
; head, mesosoma, and gaster dark brown, contrasting sharply with yellowish-brown antennae, mandibles, mesocoxae, metacoxae, and legs; very short, angular, propodeal dorsal face.
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF06FD5EFD22FCD7" blockId="14.[151,1437,591,1483]" box="[199,675,765,790]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF06FD5EFEFAFCD4" bold="true" box="[199,379,765,790]" pageId="14" pageNumber="24">Compare with:</emphasis>
<taxonomicName id="389286E5FFF4C135FE43FD5DFE73FCD7" box="[386,498,765,789]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="boltoni">
<emphasis id="CDE62174FFF4C135FE43FD5DFE73FCD7" box="[386,498,765,789]" italics="true" pageId="14" pageNumber="24">N. boltoni</emphasis>
</taxonomicName>
and
<taxonomicName id="389286E5FFF4C135FDE8FD5DFD22FCD7" box="[553,675,765,789]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="scintilla">
<emphasis id="CDE62174FFF4C135FDE8FD5DFD22FCD7" box="[553,675,765,789]" italics="true" pageId="14" pageNumber="24">N. scintilla</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF06FCBCFD5DFC98" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
WORKER.
<emphasis id="CDE62174FFF4C135FE90FC82FDD3FCFA" box="[337,594,800,824]" italics="true" pageId="14" pageNumber="24">Measurements (n = 3)</emphasis>
: TL: 2.12.4; HW: 0.490.51; HL: 0.590.61; EL: 0.140.16; SL: 0.62 0.64; PW: 0.350.38; WL: 0.620.69; GL: 0.921.2
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF04FCC6FD49FCBF" blockId="14.[151,1437,591,1483]" box="[197,712,869,893]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF04FCC6FE9EFCBF" box="[197,287,869,893]" italics="true" pageId="14" pageNumber="24">Indices:</emphasis>
CI: 8084; REL: 2327; SI: 123130
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF04FC24FB8CFB77" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">Overall dark brown, with lighter brown to yellow mandibles, antennae, and legs; cuticle smooth and shining; procoxae dark brown, with mesocoxae and metacoxae yellow-brown to yellow in color. Head with a dense layer of pubescence; posterior margin with rounded posterolateral corners, slightly emarginate medially; scapes surpass posterior margin by about the length of the first two funicular segments; scapes with scattered erect macrosetae and a dense layer of pubescence (SMC = 1316). Mesosoma with erect macrosetae of varying lengths concentrated on posterior pronotum and mesonotum (PMC = 35; MMC = 2); layer of pubescence covers mesosomal notum; metanotal area compact; dorsal face of propodeum angular and low (lower than mesonotum) with very short dorsal face and longer declivitous face; propodeum dorsum with pubescence; declivity smooth and shining with no pubescence. Gaster with abundant erect macrosetae and a dense layer of pubescence.</paragraph>
<paragraph id="FF2DFD66FFF4C135FF06FB1CFC2FFB15" blockId="14.[151,1437,591,1483]" box="[199,942,1215,1240]" pageId="14" pageNumber="24">The queen and male castes are currently unknown for this species.</paragraph>
</subSubSection>
<subSubSection id="B788AEEDFFF4C135FF06FB40FE9EFADF" pageId="14" pageNumber="24" type="etymology">
<paragraph id="FF2DFD66FFF4C135FF06FB40FE9EFADF" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF06FB40FECFFB39" bold="true" box="[199,334,1251,1275]" pageId="14" pageNumber="24">Etymology.</emphasis>
The species epithet
<taxonomicName id="389286E5FFF4C135FDF0FB40FD0AFB39" box="[561,651,1251,1275]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="umbella">
<emphasis id="CDE62174FFF4C135FDF0FB40FD0AFB39" box="[561,651,1251,1275]" italics="true" pageId="14" pageNumber="24">umbella</emphasis>
</taxonomicName>
(L. = shade or shadow) is in reference to the dark brown coloration of this species.
</paragraph>
</subSubSection>
<subSubSection id="B788AEEDFFF4C135FF06FA8BFF4AFA08" pageId="14" pageNumber="24" type="materials_examined">
<paragraph id="FF2DFD66FFF4C135FF06FA8BFDAFFAA0" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF06FA8BFDB2FA82" bold="true" box="[199,563,1319,1344]" pageId="14" pageNumber="24">
Non-type material.
<collectingCountry id="8785BDF6FFF4C135FE6EFA84FDAFFA82" box="[431,558,1319,1344]" name="Uganda" pageId="14" pageNumber="24">UGANDA</collectingCountry>
:
</emphasis>
Murchison Falls National Park, Rabongo Forest, 02°15ʹ N, 31°48ʹ E*,
<quantity id="386A5083FFF4C135FA8FFA84FA16FAFD" box="[1358,1431,1319,1343]" metricMagnitude="2" metricUnit="m" metricValue="9.58" pageId="14" pageNumber="24" unit="m" value="958.0">958 m</quantity>
,
<date id="8B2CDBA6FFF4C135FF56FAE9FEA7FAA0" box="[151,294,1354,1378]" pageId="14" pageNumber="24" value="2009-07-11">11 July 2009</date>
(W. Freund &amp; T. Klug)
</paragraph>
<paragraph id="FF2DFD66FFF4C135FF06FACEFF4AFA08" blockId="14.[151,1437,591,1483]" pageId="14" pageNumber="24">
<emphasis id="CDE62174FFF4C135FF06FACEFE90FA47" bold="true" box="[199,273,1389,1413]" pageId="14" pageNumber="24">Notes.</emphasis>
This is among the darkest of the Afrotropical species, with strongly contrasting antennae, coxae, and legs. Based on overall similarities in size and shape,
<taxonomicName id="389286E5FFF4C135FD2EFA32FCEAFA6A" box="[751,875,1424,1448]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="umbella">
<emphasis id="CDE62174FFF4C135FD2EFA32FCEAFA6A" box="[751,875,1424,1448]" italics="true" pageId="14" pageNumber="24">N. umbella</emphasis>
</taxonomicName>
appears closely related to
<taxonomicName id="389286E5FFF4C135FB5BFA32FA8AFA6A" box="[1178,1291,1424,1448]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="boltoni">
<emphasis id="CDE62174FFF4C135FB5BFA32FA8AFA6A" box="[1178,1291,1424,1448]" italics="true" pageId="14" pageNumber="24">N. boltoni</emphasis>
</taxonomicName>
and
<taxonomicName id="389286E5FFF4C135FA85FA32FF45FA09" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="24" phylum="Arthropoda" rank="species" species="scintilla">
<emphasis id="CDE62174FFF4C135FA85FA32FF45FA09" italics="true" pageId="14" pageNumber="24">N. scintilla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>