treatments-xml/data/76/02/87/760287C7CF49FFA30FB4FB79FDA5F8A2.xml

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<mods:title id="D937FE8FD40269FADC59343B879BAF17">Reappraisal of the two axiid genera Manaxius Kensley, 2003 and Calaxidium Sakai, 2014, and description of a new species of Calaxius Sakai &amp; de Saint Laurent, 1989 from French Polynesia, Southwest Pacific (Crustacea: Decapoda: Axiidea)</mods:title>
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<heading id="A55C81BDCF49FFA10FB4FB79FD03FB51" bold="true" box="[151,746,1188,1215]" fontSize="11" level="1" pageId="1" pageNumber="499" reason="1">
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Genus 
<taxonomicName id="39AB4D52CF49FFA10FCDFB78FD03FB51" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" authority="Sakai &amp; de Saint Laurent, 1989" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[238,746,1188,1215]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF49FFA10FCDFB78FEB3FB50" bold="true" box="[238,346,1188,1214]" italics="true" pageId="1" pageNumber="499">Calaxius</emphasis>
Sakai &amp; de 
<bibRefCitation id="9A3A4B20CF49FFA10ED1FB79FD03FB51" author="Saint" box="[498,746,1189,1215]" pageId="1" pageNumber="499" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent, 1989</bibRefCitation>
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<taxonomicName id="39AB4D52CF49FFA10FB4FB30FDB6FAEC" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" authority="Sakai &amp; de Saint Laurent, 1989: 84" authorityName="Sakai &amp; de Saint Laurent" authorityPageNumber="84" authorityYear="1989" box="[151,607,1260,1282]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF49FFA10FB4FB30FF18FAEC" box="[151,241,1260,1282]" italics="true" pageId="1" pageNumber="499">Calaxius</emphasis>
Sakai &amp; de 
<treatmentCitation id="7F0A10C0CF49FFA10E4DFB30FDB6FAEC" author="Saint" box="[366,607,1260,1282]" page="84" pageId="1" pageNumber="499" year="1989">
<bibRefCitation id="9A3A4B20CF49FFA10E4DFB30FDD3FAEC" author="Saint" box="[366,570,1260,1282]" pageId="1" pageNumber="499" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent, 1989</bibRefCitation>
: 84
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</taxonomicName>
; 
<treatmentCitation id="7F0A10C0CF49FFA10D49FB31FCECFAEC" author="Poore" box="[618,773,1260,1282]" page="97" pageId="1" pageNumber="499" year="1994">
<bibRefCitation id="9A3A4B20CF49FFA10D49FB31FD0BFAEC" author="Poore" box="[618,738,1260,1282]" pageId="1" pageNumber="499" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to families and genera. Memoirs of the Museum of Victoria, 54, 79 - 120." type="journal article" year="1994">Poore 1994</bibRefCitation>
: 97
</treatmentCitation>
(key); 
<treatmentCitation id="7F0A10C0CF49FFA10C6DFB30FC1CFAEC" author="Sakai" box="[846,1013,1260,1282]" page="192" pageId="1" pageNumber="499" year="1994">
<bibRefCitation id="9A3A4B20CF49FFA10C6DFB30FC2AFAEC" author="Sakai" box="[846,963,1260,1282]" pageId="1" pageNumber="499" refString="Sakai, K. (1994) Eleven species of Australian Axiidae (Crustacea: Decapoda: Thalassinidea) with descriptions of one new genus and five new species. The Beagle, Occasional Papers of the Northern Territory Museum of Arts and Sciences, 11, 175 - 202." type="journal article" year="1994">Sakai 1994</bibRefCitation>
: 192
</treatmentCitation>
; 
<treatmentCitation id="7F0A10C0CF49FFA10B23FB30FB3AFAEC" author="Clark" box="[1024,1235,1260,1282]" page="64" pageId="1" pageNumber="499" year="2007">
<bibRefCitation id="9A3A4B20CF49FFA10B23FB30FB47FAEC" author="Clark" box="[1024,1198,1260,1282]" pageId="1" pageNumber="499" refString="Clark, P. F., Galil, B. &amp; Poore, G. C. B. (2007) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from West Africa (Crustacea, Decapoda, Axiidae) and synonymy of Manaxius Kensley, 2003. Proceedings of the Biological Society of Washington, 120, 63 - 73. http: // dx. doi. org / 10.2988 / 0006 - 324 X (2007) 120 [63: ANSOCS] 2.0. CO; 2" type="journal article" year="2007">
Clark 
<emphasis id="CCDFEAC3CF49FFA10B1DFB31FB82FAEC" box="[1086,1131,1260,1282]" italics="true" pageId="1" pageNumber="499">et al</emphasis>
. 2007
</bibRefCitation>
: 64
</treatmentCitation>
. 
<taxonomicName id="39AB4D52CF49FFA10FB4FAD7FE21FACE" ID-CoL="40429644-704b-4b92-9ef3-df9c45916044" authority="Kensley, 2003: 367" authorityName="Kensley" authorityPageNumber="367" authorityYear="2003" box="[151,456,1290,1312]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF49FFA10FB4FAD7FF12FACE" box="[151,251,1291,1312]" italics="true" pageId="1" pageNumber="499">Manaxius</emphasis>
<treatmentCitation id="7F0A10C0CF49FFA10E22FAD6FE21FACE" author="Kensley" box="[257,456,1290,1312]" page="367" pageId="1" pageNumber="499" year="2003">
<bibRefCitation id="9A3A4B20CF49FFA10E22FAD6FE7CFACE" author="Kensley" box="[257,405,1290,1312]" pageId="1" pageNumber="499" refString="Kensley, B. (2003) Axioid shrimps from Guam (Crustacea, Decapoda, Thalassinidea). Micronesica, 35 - 36, 359 - 384." type="book chapter" year="2003">Kensley, 2003</bibRefCitation>
: 367
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="7F0A10C0CF49FFA10EF1FAD6FD90FACE" author="Sakai" box="[466,633,1290,1312]" page="131" pageId="1" pageNumber="499" year="2011">
<bibRefCitation id="9A3A4B20CF49FFA10EF1FAD6FDAEFACE" author="Sakai" box="[466,583,1290,1312]" pageId="1" pageNumber="499" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
: 131
</treatmentCitation>
.
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</paragraph>
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<treatmentCitationGroup id="DEBB11FFCF49FFA10FB4FAF5FE2CFAD1" box="[151,453,1321,1343]" pageId="1" pageNumber="499">
<taxonomicName id="39AB4D52CF49FFA10FB4FAF5FE28FAD1" ID-CoL="6cdf28cc-c991-42c0-9c4d-2a5c61d403e7" authority="Sakai, 2014: 887" authorityName="Sakai" authorityPageNumber="887" authorityYear="2014" box="[151,449,1321,1343]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF49FFA10FB4FAF5FEE4FAD1" box="[151,269,1321,1343]" italics="true" pageId="1" pageNumber="499">Calaxidium</emphasis>
<treatmentCitation id="7F0A10C0CF49FFA10E30FAF5FE28FAD1" author="Sakai" box="[275,449,1321,1343]" page="887" pageId="1" pageNumber="499" year="2014">
<bibRefCitation id="9A3A4B20CF49FFA10E30FAF5FE66FAD1" author="Sakai" box="[275,399,1321,1343]" pageId="1" pageNumber="499" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai, 2014</bibRefCitation>
: 887
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</taxonomicName>
.
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</paragraph>
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<paragraph id="FE1436D1CF49FFA10FB4FAB0FB68FA6B" blockId="1.[151,1436,1388,2025]" box="[151,1153,1388,1413]" pageId="1" pageNumber="499">
<emphasis id="CCDFEAC3CF49FFA10FB4FAB0FEC2FA6B" bold="true" box="[151,299,1388,1413]" pageId="1" pageNumber="499">
<typeStatus id="21108873CF49FFA10FB4FAB0FF3BFA6B" box="[151,210,1388,1413]" pageId="1" pageNumber="499">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="39AB4D52CF49FFA10E1AFAB0FC66FA6A" authority="Sakai &amp; de Saint Laurent, 1989" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[313,911,1388,1413]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="499" phylum="Arthropoda" rank="species" species="acutirostris">
<emphasis id="CCDFEAC3CF49FFA10E1AFAB0FDCAFA6A" box="[313,547,1388,1412]" italics="true" pageId="1" pageNumber="499">Calaxius acutirostris</emphasis>
Sakai &amp; de 
<bibRefCitation id="9A3A4B20CF49FFA10D8DFAB0FC66FA6A" author="Saint" box="[686,911,1388,1413]" pageId="1" pageNumber="499" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent, 1989</bibRefCitation>
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, original designation.
</paragraph>
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<paragraph id="FE1436D1CF49FFA10FE4FA53FAE9F806" blockId="1.[151,1436,1388,2025]" pageId="1" pageNumber="499">
<emphasis id="CCDFEAC3CF49FFA10FE4FA53FED3FA46" bold="true" box="[199,314,1423,1448]" pageId="1" pageNumber="499">Diagnosis</emphasis>
. Emended from 
<bibRefCitation id="9A3A4B20CF49FFA10EDEFA4CFCE6FA46" author="Poore" box="[509,783,1423,1448]" pageId="1" pageNumber="499" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae (Crustacea: Decapoda: Axiidea). Memoirs of Museum Victoria, 66, 221 - 287." type="journal article" year="2009">Poore &amp; Collins (2009)</bibRefCitation>
. Carapace smooth or tuberculate; cervical groove visible laterally over third distance to anterolateral margin. Rostrum acutely triangular, with 2 or 3 pairs of lateral spines, usually distinctly longer than eyestalks, not depressed below level of carapace, continuous with definite lateral carinae; supraocular spines prominent; lateral carina with at least 1 spine or denticle; submedian carina present, with 1 or more spines or denticles; median carina a weak ridge, with 1 or few spines or tubercles; postcervical carina absent or restricted to posterior part. First pleomere with acutely pointed pleuron; pleuron of second pleomere acute or rounded, pleura of third to fifth pleomeres acute, sometimes with anteroventral tooth, or rounded. Telson with lateral fixed spines and posterolateral spiniform setae; posterior margin rounded. Eyestalk cylindrical, articulating; cornea pigmented. Antenna with scaphocerite long, acute. Third maxilliped with exopod not clearly bent at base of flagellum. Pleurobranchs present above second to fourth pereopods; podobranchs and arthrobranchs well developed; epipods present on second maxilliped to fourth pereopod. First pereopods (chelipeds) nearly symmetrical to clearly asymmetrical, with propodus compressed laterally; dorsal margin of palm with moderate to strong spines (rarely obsolete), covered with tufts of long setae. Third and fourth pereopods each with transverse rows of spiniform setae on lateral surface of propodus; dactyli tapering, each with longitudinal rows of spiniform setae on lateral surface and flexor margin. Second to fifth pleopods each with appendix interna in both sexes. First pleopod of male absent. Second pleopod of male with rod-like appendix masculina clearly separated from appendix interna. Uropodal endopod with lateral and distolateral spines; exopod with transverse suture.
</paragraph>
</subSubSection>
<subSubSection id="B6B1655ACF4AFFA30FE4FF4BFD9EFC4A" lastPageId="3" lastPageNumber="501" pageId="2" pageNumber="500" type="discussion">
<paragraph id="FE1436D1CF4AFFA20FE4FF4BFE27FE66" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="500">
<emphasis id="CCDFEAC3CF4AFFA20FE4FF4BFEDDFF5E" bold="true" box="[199,308,151,176]" pageId="2" pageNumber="500">Remarks</emphasis>
. The above generic diagnosis is slightly emended from that presented by 
<bibRefCitation id="9A3A4B20CF4AFFA20BA4FF44FA7EFF5E" author="Poore" box="[1159,1431,151,176]" pageId="2" pageNumber="500" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae (Crustacea: Decapoda: Axiidea). Memoirs of Museum Victoria, 66, 221 - 287." type="journal article" year="2009">Poore &amp; Collins (2009)</bibRefCitation>
, incorporating interspecific variation of some characters. For example, 
<taxonomicName id="39AB4D52CF4AFFA20C97FF60FBA2FF3A" box="[948,1099,188,212]" class="Malacostraca" family="Axiidae" genus="Axius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="CCDFEAC3CF4AFFA20C97FF60FBA2FF3A" box="[948,1099,188,212]" italics="true" pageId="2" pageNumber="500">C. inaequalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4AFFA20BA0FF60FB1DFF3A" box="[1155,1268,188,212]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="jenneri">
<emphasis id="CCDFEAC3CF4AFFA20BA0FF60FB1DFF3A" box="[1155,1268,188,212]" italics="true" pageId="2" pageNumber="500">C. jenneri</emphasis>
</taxonomicName>
are unusual in having the rostrum just reaching to the distal corneal margins (
<bibRefCitation id="9A3A4B20CF4AFFA20C71FF03FC1DFF16" author="Rathbun" box="[850,1012,223,248]" pageId="2" pageNumber="500" refString="Rathbun, M. J. (1901) The Brachyura and Macrura of Porto Rico. Bulletin of the U. S. Fish Commission, 20, 1 - 137." type="journal article" year="1901">Rathbun 1901</bibRefCitation>
; 
<bibRefCitation id="9A3A4B20CF4AFFA20B23FF03FB41FF16" author="Williams" box="[1024,1192,223,248]" pageId="2" pageNumber="500" refString="Williams, A. B. (1974) Two new axiids (Crustacea: Decapoda: Thalassinidea: Calocaris) from North Carolina and the Straits of Florida. Proceedings of the Biological Society of Washington, 87, 451 - 464." type="journal article" year="1974">Williams 1974</bibRefCitation>
). The development of the postcervical carina on the carapace varies according to species presently assigned to 
<taxonomicName id="39AB4D52CF4AFFA20BC1FED8FAAFFEF2" box="[1250,1350,260,284]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20BC1FED8FAAFFEF2" box="[1250,1350,260,284]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
. In 
<taxonomicName id="39AB4D52CF4AFFA20AA1FED8FEFEFED1" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="acutirostris">
<emphasis id="CCDFEAC3CF4AFFA20AA1FED8FEFEFED1" italics="true" pageId="2" pageNumber="500">C. acutirostris</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4AFFA20E0BFEF4FE5EFED1" box="[296,439,296,319]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="manningi">
<emphasis id="CCDFEAC3CF4AFFA20E0BFEF4FE5EFED1" box="[296,439,296,319]" italics="true" pageId="2" pageNumber="500">C. manningi</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4AFFA20ED7FEF4FDB8FED1" box="[500,593,296,319]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="tungi">
<emphasis id="CCDFEAC3CF4AFFA20ED7FEF4FDB8FED1" box="[500,593,296,319]" italics="true" pageId="2" pageNumber="500">C. tungi</emphasis>
</taxonomicName>
, the postcervical carina is distinct, running over the entire length of the postcervical carapace (Sakai and de 
<bibRefCitation id="9A3A4B20CF4AFFA20D13FE90FCE2FE8A" author="Saint" box="[560,779,332,357]" pageId="2" pageNumber="500" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent 1989</bibRefCitation>
; 
<bibRefCitation id="9A3A4B20CF4AFFA20C34FE90FC4BFE8A" author="Zhong" box="[791,930,332,357]" pageId="2" pageNumber="500" refString="Zhong, Z. - R. (2000) A new species of Axiidae from the south Chin Sea (Crustacea. Decapoda: Thalassinidea. Acta Zootaxonomica Sinica, 25, 33 - 36. [in Chinese with English abstract]" type="journal article" year="2000">Zhong 2000</bibRefCitation>
; 
<bibRefCitation id="9A3A4B20CF4AFFA20C8CFE90FB64FE8A" author="Kensley" box="[943,1165,332,357]" pageId="2" pageNumber="500" refString="Kensley, B., Lin, F. - J. &amp; Yu, H. - P. (2000) Further records of thalassinidean shrimps from Taiwan (Decapoda: Axiidae and Calocarididae), with descriptions of three new species. Journal of Crustacean Biology, 20 (Special Number 2), 207 - 217. http: // dx. doi. org / 10.1163 / 1937240 X- 90000022" type="journal article" year="2000">
Kensley 
<emphasis id="CCDFEAC3CF4AFFA20B31FE91FBAAFE8A" box="[1042,1091,332,356]" italics="true" pageId="2" pageNumber="500">et al</emphasis>
. 2000
</bibRefCitation>
), whereas it is absent or present in the other species.
</paragraph>
<paragraph id="FE1436D1CF4AFFA20FE4FE48FB7CFD6B" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="500">
<bibRefCitation id="9A3A4B20CF4AFFA20FE4FE48FEBFFE42" author="Sakai" box="[199,342,404,429]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
realized that Kensley’s (2003) 
<taxonomicName id="39AB4D52CF4AFFA20D9AFE49FC5CFE42" box="[697,949,405,428]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="CCDFEAC3CF4AFFA20D9AFE49FC5CFE42" box="[697,949,405,428]" italics="true" pageId="2" pageNumber="500">Manaxius pitatucensis</emphasis>
</taxonomicName>
was not 
<taxonomicName id="39AB4D52CF4AFFA20B38FE48FA68FE42" box="[1051,1409,404,428]" class="Malacostraca" family="Axiidae" genus="Axiopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="pitatucensis" subGenus="Axiopsis">
<emphasis id="CCDFEAC3CF4AFFA20B38FE48FB93FE45" box="[1051,1146,404,427]" italics="true" pageId="2" pageNumber="500">Axiopsis</emphasis>
(
<emphasis id="CCDFEAC3CF4AFFA20BA8FE48FB03FE45" box="[1163,1258,404,427]" italics="true" pageId="2" pageNumber="500">Axiopsis</emphasis>
) 
<emphasis id="CCDFEAC3CF4AFFA20BD8FE49FA68FE42" box="[1275,1409,405,428]" italics="true" pageId="2" pageNumber="500">pitatucensis</emphasis>
</taxonomicName>
, a species which he then redescribed and illustrated from 
<typeStatus id="21108873CF4AFFA20C2DFE64FCD7FE3E" box="[782,830,440,464]" pageId="2" pageNumber="500">type</typeStatus>
material as 
<taxonomicName id="39AB4D52CF4AFFA20CEFFE64FB12FE21" box="[972,1275,440,464]" class="Malacostraca" family="Axiidae" genus="Colemanaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="CCDFEAC3CF4AFFA20CEFFE64FB12FE21" box="[972,1275,440,464]" italics="true" pageId="2" pageNumber="500">Colemanaxius pitatucensis</emphasis>
</taxonomicName>
. 
<bibRefCitation id="9A3A4B20CF4AFFA20A28FE6BFA75FE3E" author="Sakai" box="[1291,1436,439,464]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
invoked ICZN Article 70.3 ‘Misidentified 
<typeStatus id="21108873CF4AFFA20DA8FE01FD52FE1B" box="[651,699,477,501]" pageId="2" pageNumber="500">type</typeStatus>
species’ to clarify the confusion. He chose as 
<typeStatus id="21108873CF4AFFA20BCFFE01FAF5FE1B" box="[1260,1308,477,501]" pageId="2" pageNumber="500">type</typeStatus>
species of 
<taxonomicName id="39AB4D52CF4AFFA20FB4FDDCFEECFDF9" box="[151,261,512,535]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20FB4FDDCFEECFDF9" box="[151,261,512,535]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
the one actually involved in the misidentification, which he named 
<taxonomicName id="39AB4D52CF4AFFA20B2AFDDCFB17FDF9" box="[1033,1278,512,535]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="aganaensis">
<emphasis id="CCDFEAC3CF4AFFA20B2AFDDCFB17FDF9" box="[1033,1278,512,535]" italics="true" pageId="2" pageNumber="500">Manaxius aganaensis</emphasis>
</taxonomicName>
. However, he did not designate name bearing 
<typeStatus id="21108873CF4AFFA20EDAFDF9FDC0FDD3" box="[505,553,549,573]" pageId="2" pageNumber="500">type</typeStatus>
from the three individuals from 
<collectingCountry id="86BC7641CF4AFFA20CB7FDF8FC32FDD2" box="[916,987,548,572]" name="Guam" pageId="2" pageNumber="500">Guam</collectingCountry>
, or the one from Okinawa 
<collectingCountry id="86BC7641CF4AFFA20A20FDF8FAA3FDD3" box="[1283,1354,548,573]" name="Iceland" pageId="2" pageNumber="500">Island</collectingCountry>
, 
<collectingCountry id="86BC7641CF4AFFA20A76FDF9FA71FDD2" box="[1365,1432,549,572]" name="Japan" pageId="2" pageNumber="500">Japan</collectingCountry>
, illustrated by 
<bibRefCitation id="9A3A4B20CF4AFFA20E16FD9BFE0EFD8E" author="Kensley" box="[309,487,583,608]" pageId="2" pageNumber="500" refString="Kensley, B. (2003) Axioid shrimps from Guam (Crustacea, Decapoda, Thalassinidea). Micronesica, 35 - 36, 359 - 384." type="book chapter" year="2003">Kensley (2003)</bibRefCitation>
, as required under ICZN Article 72.3. 
<collectingCountry id="86BC7641CF4AFFA20C8BFD94FC2EFD8E" box="[936,967,584,608]" name="American Samoa" pageId="2" pageNumber="500">As</collectingCountry>
pointed out by 
<bibRefCitation id="9A3A4B20CF4AFFA20BA2FD94FACFFD8E" author="Poore" box="[1153,1318,583,608]" pageId="2" pageNumber="500" refString="Poore, G. C. B. (2015 b) Manaxius aganaensis Sakai, 2011. Accessed through: World Register of Marine Species. Available from: http: // www. marinespecies. org / aphia. php? p = taxdetails &amp; id = 741322 (accessed 29 October 2015)" type="book chapter" year="2015" yearSuffix="b">Poore (2015b)</bibRefCitation>
, the name 
<taxonomicName id="39AB4D52CF4AFFA20FB4FDB1FDFDFD6A" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[151,532,620,645]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="aganaensis">
<emphasis id="CCDFEAC3CF4AFFA20FB4FDB1FE63FD6A" box="[151,394,621,644]" italics="true" pageId="2" pageNumber="500">Manaxius aganaensis</emphasis>
<bibRefCitation id="9A3A4B20CF4AFFA20EB2FDB0FDFDFD6A" author="Sakai" box="[401,532,620,645]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
is not available and can not be 
<typeStatus id="21108873CF4AFFA20C55FDB1FC4FFD6B" box="[886,934,621,645]" pageId="2" pageNumber="500">type</typeStatus>
species of the genus.
</paragraph>
<paragraph id="FE1436D1CF4AFFA20FE4FD53FB69FA96" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="500">
<bibRefCitation id="9A3A4B20CF4AFFA20FE4FD53FEBCFD46" author="Sakai" box="[199,341,655,680]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
further complicated the issue by including 
<taxonomicName id="39AB4D52CF4AFFA20C15FD4CFC4DFD49" box="[822,932,656,679]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20C15FD4CFC4DFD49" box="[822,932,656,679]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
both as a junior synonym of 
<taxonomicName id="39AB4D52CF4AFFA20BCBFD4CFAA5FD46" box="[1256,1356,656,680]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20BCBFD4CFAA5FD46" box="[1256,1356,656,680]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
(p. 84) and as a separate genus (p. 131), but it is assumed that Sakai’s intention was the reinstatement of 
<taxonomicName id="39AB4D52CF4AFFA20BDBFD69FA8FFD22" box="[1272,1382,693,716]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20BDBFD69FA8FFD22" box="[1272,1382,693,716]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
as a valid genus. Sakai’s (2011: 131) 
<taxonomicName id="39AB4D52CF4AFFA20D2EFD04FD92FD01" box="[525,635,728,751]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20D2EFD04FD92FD01" box="[525,635,728,751]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
includes vast majority of species previously assigned to 
<taxonomicName id="39AB4D52CF4AFFA20A26FD04FA80FD1E" box="[1285,1385,728,752]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20A26FD04FA80FD1E" box="[1285,1385,728,752]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, but his logic is very difficult to trace: 
<taxonomicName id="39AB4D52CF4AFFA20D2DFD20FD9BFCFA" box="[526,626,764,788]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20D2DFD20FD9BFCFA" box="[526,626,764,788]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
was compared only with 
<taxonomicName id="39AB4D52CF4AFFA20CADFD20FEEEFCD6" authority="(Herbst, 1796) (Sakai 2011: 84)" authorityName="(Herbst, 1796) (Sakai" authorityPageNumber="84" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Eutrichocheles" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="CCDFEAC3CF4AFFA20CADFD20FB4CFCFA" box="[910,1189,764,788]" italics="true" pageId="2" pageNumber="500">Eutrichocheles modestus</emphasis>
(
<bibRefCitation id="9A3A4B20CF4AFFA20B90FD20FAAFFCFB" author="Herbst" box="[1203,1350,764,789]" pageId="2" pageNumber="500" refString="Herbst, J. F. W. (1796) Versuch einer Naturgeschichte der Krabben und Krese nebst einer systematischen Beschreibung ihrer verschiedenen Arten. Vol. 2. Bei Gottlieb August Lange, Berlin, 226 pp., pls. 22 - 46." type="book chapter" year="1796">Herbst, 1796</bibRefCitation>
) (
<bibRefCitation id="9A3A4B20CF4AFFA20A7FFD20FF17FCD6" author="Sakai" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011: 84</bibRefCitation>
)
</taxonomicName>
, whereas 
<taxonomicName id="39AB4D52CF4AFFA20E59FCFCFE01FCD9" box="[378,488,800,823]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20E59FCFCFE01FCD9" box="[378,488,800,823]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
was compared only with 
<taxonomicName id="39AB4D52CF4AFFA20C36FCFCFAA8FCD6" authority="Wollebaek, 1908 (Sakai 2011: 131)" authorityName="Wollebaek, 1908 (Sakai" authorityPageNumber="131" authorityYear="2011" box="[789,1345,799,824]" class="Malacostraca" family="Axiidae" genus="Calocarides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20C36FCFCFC77FCD6" box="[789,926,800,824]" italics="true" pageId="2" pageNumber="500">Calocarides</emphasis>
<bibRefCitation id="9A3A4B20CF4AFFA20C8BFCC3FB8EFCD6" author="Wollebaek" box="[936,1127,799,824]" pageId="2" pageNumber="500" refString="Wollebaek, A. (1908) Remarks on decapod crustaceans of the North Atlantic and the Norwegian Fiords (I and II). Bergens Museum Arbog Afhandlinger of Arsberetning, 12, 1 - 74." type="journal article" year="1908">Wollebaek, 1908</bibRefCitation>
(
<bibRefCitation id="9A3A4B20CF4AFFA20B5AFCC3FADEFCD6" author="Sakai" box="[1145,1335,799,824]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011: 131</bibRefCitation>
)
</taxonomicName>
. On the other hand, in the key to the genera (p. 17), 
<taxonomicName id="39AB4D52CF4AFFA20DABFC98FD05FCB2" box="[648,748,836,860]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20DABFC98FD05FCB2" box="[648,748,836,860]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
is placed close to 
<taxonomicName id="39AB4D52CF4AFFA20C9EFC98FB06FCB2" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[957,1263,836,861]" class="Malacostraca" family="Axiidae" genus="Albatrossaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20C9EFC98FB8BFCB2" box="[957,1122,836,860]" italics="true" pageId="2" pageNumber="500">Albatrossaxius</emphasis>
<bibRefCitation id="9A3A4B20CF4AFFA20B48FC98FB06FCB2" author="Sakai" box="[1131,1263,836,861]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4AFFA20A05FC99FA7DFCB2" box="[1318,1428,837,860]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20A05FC99FA7DFCB2" box="[1318,1428,837,860]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
. In his key, Sakai uses only the armature of the gastric median carina in differentiating 
<taxonomicName id="39AB4D52CF4AFFA20B4CFCB4FB3AFC6E" box="[1135,1235,872,896]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20B4CFCB4FB3AFC6E" box="[1135,1235,872,896]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
(“Gastric median carina linear”) and the latter two genera (“Gastric median carina tuberculate), but in 
<taxonomicName id="39AB4D52CF4AFFA20BA6FC50FAC2FC4A" box="[1157,1323,908,932]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="acutirostris">
<emphasis id="CCDFEAC3CF4AFFA20BA6FC50FAC2FC4A" box="[1157,1323,908,932]" italics="true" pageId="2" pageNumber="500">C. acutirostris</emphasis>
</taxonomicName>
, the 
<typeStatus id="21108873CF4AFFA20A4FFC51FA75FC4B" box="[1388,1436,909,933]" pageId="2" pageNumber="500">type</typeStatus>
species of 
<taxonomicName id="39AB4D52CF4AFFA20E2DFC6CFE9BFC26" box="[270,370,944,968]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20E2DFC6CFE9BFC26" box="[270,370,944,968]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, the gastric median carina is actually denticulate (cf. Sakai &amp; de 
<bibRefCitation id="9A3A4B20CF4AFFA20B76FC73FAD9FC26" author="Saint" box="[1109,1328,943,968]" pageId="2" pageNumber="500" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent 1989</bibRefCitation>
; 
<bibRefCitation id="9A3A4B20CF4AFFA20A1EFC6CFEC5FC02" author="Poore" pageId="2" pageNumber="500" refString="Poore, G. C. B. &amp; Collins, D. J. (2009) Australian Axiidae (Crustacea: Decapoda: Axiidea). Memoirs of Museum Victoria, 66, 221 - 287." type="journal article" year="2009">Poore &amp; Collins 2009</bibRefCitation>
). Comparison of his (2011) diagnoses of the two genera shows them to differ only in the postcervical carina being present in 
<taxonomicName id="39AB4D52CF4AFFA20E88FC24FDE6FBFE" box="[427,527,1016,1040]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20E88FC24FDE6FBFE" box="[427,527,1016,1040]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, absent in 
<taxonomicName id="39AB4D52CF4AFFA20DB0FC24FCE8FBE1" box="[659,769,1016,1039]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20DB0FC24FCE8FBE1" box="[659,769,1016,1039]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
, the anterolateral margin of the carapace with a spine in 
<taxonomicName id="39AB4D52CF4AFFA20FB4FBC0FF12FBDA" box="[151,251,1052,1076]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20FB4FBC0FF12FBDA" box="[151,251,1052,1076]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, absent in 
<taxonomicName id="39AB4D52CF4AFFA20EA7FBC1FE1BFBDA" box="[388,498,1053,1076]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20EA7FBC1FE1BFBDA" box="[388,498,1053,1076]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
, and the scaphocerite protruded straight forward in 
<taxonomicName id="39AB4D52CF4AFFA20B46FBC0FB20FBDA" box="[1125,1225,1052,1076]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20B46FBC0FB20FBDA" box="[1125,1225,1052,1076]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, ‘horn-shaped’ in 
<taxonomicName id="39AB4D52CF4AFFA20FB4FB9CFEECFBB9" box="[151,261,1088,1111]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20FB4FB9CFEECFBB9" box="[151,261,1088,1111]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
. None of these features are of generic value if real. The development of the postcervical carina appears variable in species Sakai referred to 
<taxonomicName id="39AB4D52CF4AFFA20D0FFBB9FD73FB92" box="[556,666,1125,1148]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20D0FFBB9FD73FB92" box="[556,666,1125,1148]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
: a trace of a postcervical carina is discernible at least in the posterior part in 
<taxonomicName id="39AB4D52CF4AFFA20FCEFB54FE7BFB71" box="[237,402,1160,1183]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="mimasensis">
<emphasis id="CCDFEAC3CF4AFFA20FCEFB54FE7BFB71" box="[237,402,1160,1183]" italics="true" pageId="2" pageNumber="500">C. mimasensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4AFFA20EF3FB54FDA4FB71" box="[464,589,1160,1183]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20EF3FB54FDA4FB71" box="[464,589,1160,1183]" italics="true" pageId="2" pageNumber="500">C. izuensis</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="9A3A4B20CF4AFFA20DABFB5BFC89FB4E" author="Komai" box="[648,864,1159,1184]" pageId="2" pageNumber="500" refString="Komai, T., Ohtsuka, S., Nakaguchi, K. &amp; Go, A. (2002) Decapod crustaceans collected from the southern part of the Sea of Japan in 2000 - 2001 using TRV Toyoshio-maru. Natural History Research, 7, 19 - 73." type="journal article" year="2002">
Komai 
<emphasis id="CCDFEAC3CF4AFFA20DFCFB55FCFAFB4E" box="[735,787,1160,1184]" italics="true" pageId="2" pageNumber="500">et al</emphasis>
. 2002
</bibRefCitation>
; 
<bibRefCitation id="9A3A4B20CF4AFFA20C4CFB5BFBEBFB4E" author="Komai" box="[879,1026,1159,1184]" pageId="2" pageNumber="500" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai 2011</bibRefCitation>
). The extent of a protrusion on the anterolateral margin is also variable and the scaphocerite is straight to slightly curved downwards (the contrast between “protruded forward” versus “horn-shaped” does not make sense). The lateral spine on the cheliped palm that characterizes all species of 
<emphasis id="CCDFEAC3CF4AFFA20D22FB28FD35FAE2" box="[513,732,1268,1292]" italics="true" pageId="2" pageNumber="500">Calaxius-Manaxius</emphasis>
complex was not mentioned in either diagnosis. 
<bibRefCitation id="9A3A4B20CF4AFFA20A28FB28FA75FAE3" author="Sakai" box="[1291,1436,1268,1293]" pageId="2" pageNumber="500" refString="Sakai, K. (2015) Two new species, Colemanaxius andamanensis sp. nov. and Bruceaxius thailandensis sp. nov. from the Andaman Sea, Thailand (infraorder Axiidea De Saint Laurent, 1979). Crustaceana, 88, 867 - 880. http: // dx. doi. org / 10.1163 / 15685403 - 00003440" type="journal article" year="2015">Sakai (2015)</bibRefCitation>
finally recognized 
<taxonomicName id="39AB4D52CF4AFFA20E5AFAC4FE0EFAC1" box="[377,487,1304,1327]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20E5AFAC4FE0EFAC1" box="[377,487,1304,1327]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="39AB4D52CF4AFFA20C0BFAC4FC65FADE" box="[808,908,1304,1328]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20C0BFAC4FC65FADE" box="[808,908,1304,1328]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, but his argumentation was based only on comparison of the respective 
<typeStatus id="21108873CF4AFFA20EC9FAE1FDF3FABB" box="[490,538,1341,1365]" pageId="2" pageNumber="500">type</typeStatus>
species (he did not realize that his 
<taxonomicName id="39AB4D52CF4AFFA20C94FAE1FBA6FABA" box="[951,1103,1341,1364]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="aganaesis">
<emphasis id="CCDFEAC3CF4AFFA20C94FAE1FBA6FABA" box="[951,1103,1341,1364]" italics="true" pageId="2" pageNumber="500">M. aganaesis</emphasis>
</taxonomicName>
was not available). Here the synonymy of 
<taxonomicName id="39AB4D52CF4AFFA20E12FABCFE76FA99" box="[305,415,1376,1399]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20E12FABCFE76FA99" box="[305,415,1376,1399]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
with 
<taxonomicName id="39AB4D52CF4AFFA20EFCFABCFDAAFA96" box="[479,579,1376,1400]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20EFCFABCFDAAFA96" box="[479,579,1376,1400]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
proposed by 
<bibRefCitation id="9A3A4B20CF4AFFA20DF9FA83FC40FA96" box="[730,937,1375,1400]" pageId="2" pageNumber="500" refString="Clark, P. F., Galil, B. &amp; Poore, G. C. B. (2007) A new species of Calaxius Sakai &amp; de Saint Laurent, 1989, from West Africa (Crustacea, Decapoda, Axiidae) and synonymy of Manaxius Kensley, 2003. Proceedings of the Biological Society of Washington, 120, 63 - 73. http: // dx. doi. org / 10.2988 / 0006 - 324 X (2007) 120 [63: ANSOCS] 2.0. CO; 2" type="journal article">
Clark 
<emphasis id="CCDFEAC3CF4AFFA20C3CFABDFCB1FA96" box="[799,856,1376,1400]" italics="true" pageId="2" pageNumber="500">et al.</emphasis>
(2007)
</bibRefCitation>
is fully confirmed.
</paragraph>
<paragraph id="FE1436D1CF4AFFA20FE4FA58FA91F8A2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="500">
<bibRefCitation id="9A3A4B20CF4AFFA20FE4FA58FE89FA72" author="Sakai" box="[199,352,1412,1437]" pageId="2" pageNumber="500" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai (2014)</bibRefCitation>
established a new monotypic genus 
<taxonomicName id="39AB4D52CF4AFFA20C0CFA58FC5BFA72" box="[815,946,1412,1436]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20C0CFA58FC5BFA72" box="[815,946,1412,1436]" italics="true" pageId="2" pageNumber="500">Calaxidium</emphasis>
</taxonomicName>
to accommodate 
<taxonomicName id="39AB4D52CF4AFFA20BB1FA58FAB7FA72" box="[1170,1374,1412,1436]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20BB1FA58FAB7FA72" box="[1170,1374,1412,1436]" italics="true" pageId="2" pageNumber="500">Calaxius izuensis</emphasis>
</taxonomicName>
. His argumentation (
<bibRefCitation id="9A3A4B20CF4AFFA20E6EFA7BFDFBFA2E" author="Sakai" box="[333,530,1447,1472]" pageId="2" pageNumber="500" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai 2014: 887</bibRefCitation>
) is directly quoted below: “However, the characters presented for 
<taxonomicName id="39AB4D52CF4AFFA20A1BFA74FF19FA0A" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20A1BFA74FF19FA0A" italics="true" pageId="2" pageNumber="500">Calaxius izuensis</emphasis>
</taxonomicName>
by Komai indicate that 
<taxonomicName id="39AB4D52CF4AFFA20D37FA10FD7AFA0A" box="[532,659,1484,1508]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20D37FA10FD7AFA0A" box="[532,659,1484,1508]" italics="true" pageId="2" pageNumber="500">C. izuensis</emphasis>
</taxonomicName>
should not be included in the genus 
<taxonomicName id="39AB4D52CF4AFFA20B71FA10FB5FFA0A" box="[1106,1206,1484,1508]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20B71FA10FB5FFA0A" box="[1106,1206,1484,1508]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
, because the genus 
<taxonomicName id="39AB4D52CF4AFFA20FB4FA2CFF12F9E6" box="[151,251,1520,1544]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20FB4FA2CFF12F9E6" box="[151,251,1520,1544]" italics="true" pageId="2" pageNumber="500">Calaxius</emphasis>
</taxonomicName>
is characterized as follows (
<bibRefCitation id="9A3A4B20CF4AFFA20D1BFA33FD2BF9E6" author="Sakai" box="[568,706,1519,1544]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
): (1) the P1 are subequal in size and shape, both chelae are rather elongate, and the fingers of both larger and smaller chelipeds are longer than the palm (vs. in Komai’s 
<taxonomicName id="39AB4D52CF4AFFA20A1BF9C8FF19F9A1" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20A1BF9C8FF19F9A1" italics="true" pageId="2" pageNumber="500">Calaxius izuensis</emphasis>
</taxonomicName>
the P1 are unequal, both chelae are stout, and the fingers of the larger cheliped are shorter than the palm, and those of the smaller cheliped are as long as the palm); (2) the abdominal pleura 1-6 are sharply pointed ventrally (vs. the abdominal pleuron 1 is sharply pointed, the abdominal pleuron 2 is truncate, and the abdominal pleura 3-6 are rounded ventrally); (3) the male Plp1 is absent, the Plp2 endopod bears mesially articulated appendices masculina and interna at the proximal third (Poore &amp; Collins, 2010 [sic], fig. 19h) (vs. the male Plp1 is absent, the Plp2 endopod bears mesially an appendix masculina fused proximally with an appendix interna 
<bibRefCitation id="9A3A4B20CF4AFFA20A63F930FEACF8C6" author="Komai" pageId="2" pageNumber="500" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011" yearSuffix="f">(Komai, 2011, fig. 21f))</bibRefCitation>
; and (4) the postcervical carina is distinct (vs. the postcervical carina is not distinct, but its trace is discernible). Hence, a new genus, 
<taxonomicName id="39AB4D52CF4AFFA20D34F8E8FD73F8A2" box="[535,666,1844,1868]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CCDFEAC3CF4AFFA20D34F8E8FD73F8A2" box="[535,666,1844,1868]" italics="true" pageId="2" pageNumber="500">Calaxidium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7EC57B8CF4AFFA20D83F8E9FCEFF8A2" box="[672,774,1845,1869]" pageId="2" pageNumber="500" rank="genus">gen. nov.</taxonomicNameLabel>
is here established for 
<taxonomicName id="39AB4D52CF4AFFA20B2AF8E8FA80F8A2" authority="Komai, 2011" authorityName="Komai" authorityYear="2011" box="[1033,1385,1844,1869]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4AFFA20B2AF8E8FB24F8A2" box="[1033,1229,1844,1868]" italics="true" pageId="2" pageNumber="500">Calaxius izuensis</emphasis>
<bibRefCitation id="9A3A4B20CF4AFFA20BF0F8E8FA80F8A2" author="Komai" box="[1235,1385,1844,1869]" pageId="2" pageNumber="500" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai, 2011</bibRefCitation>
</taxonomicName>
.”
</paragraph>
<paragraph id="FE1436D1CF4AFFA30FE4F884FE15FE8A" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1437,151,1869]" lastPageId="3" lastPageNumber="501" pageId="2" pageNumber="500">
<collectingCountry id="86BC7641CF4AFFA20FE4F884FF0FF89E" box="[199,230,1880,1904]" name="American Samoa" pageId="2" pageNumber="500">As</collectingCountry>
noted above, three of the four features cited by 
<bibRefCitation id="9A3A4B20CF4AFFA20DDDF88BFC67F89E" author="Sakai" box="[766,910,1879,1904]" pageId="2" pageNumber="500" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai (2014)</bibRefCitation>
for diagnosing 
<taxonomicName id="39AB4D52CF4AFFA20B1DF884FB28F89E" box="[1086,1217,1880,1904]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20B1DF884FB28F89E" box="[1086,1217,1880,1904]" italics="true" pageId="2" pageNumber="500">Calaxidium</emphasis>
</taxonomicName>
are seen also in the species transferred to 
<taxonomicName id="39AB4D52CF4AFFA20EB7F8A1FDEBF87A" box="[404,514,1917,1940]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20EB7F8A1FDEBF87A" box="[404,514,1917,1940]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
</taxonomicName>
by 
<bibRefCitation id="9A3A4B20CF4AFFA20D11F8A0FD2FF87A" author="Sakai" box="[562,710,1916,1941]" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
, although 
<bibRefCitation id="9A3A4B20CF4AFFA20C63F8A0FC3AF87A" author="Sakai" box="[832,979,1916,1941]" pageId="2" pageNumber="500" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai (2014)</bibRefCitation>
did not mention 
<taxonomicName id="39AB4D52CF4AFFA20BBEF8A1FF09F856" authority="Sakai (2011)" authorityName="Sakai" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20BBEF8A1FAE5F87A" box="[1181,1292,1917,1940]" italics="true" pageId="2" pageNumber="500">Manaxius</emphasis>
sensu 
<bibRefCitation id="9A3A4B20CF4AFFA20A7DF8A0FF09F856" author="Sakai" pageId="2" pageNumber="500" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
</taxonomicName>
in comparison with 
<taxonomicName id="39AB4D52CF4AFFA20EE5F87CFDA0F856" box="[454,585,1952,1976]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="500" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4AFFA20EE5F87CFDA0F856" box="[454,585,1952,1976]" italics="true" pageId="2" pageNumber="500">Calaxidium</emphasis>
</taxonomicName>
at all. With regard to the character of the appendix masculina on the second pleopod, the basal articulation of it has never been considered as of taxonomic significance in taxa with a nonmodified second pleopod. In fact, the lack of the basal articulation in Komai’s (
<bibRefCitation id="9A3A4B20CF4BFFA30B08FF4BFAC5FF5E" author="Komai" box="[1067,1324,151,176]" pageId="3" pageNumber="501" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai 2011: Fig. 21F</bibRefCitation>
) figure is merely an artistic omission. 
<bibRefCitation id="9A3A4B20CF4BFFA30EFDFF60FD97FF3A" author="Komai" box="[478,638,188,213]" pageId="3" pageNumber="501" refString="Komai, T. (2011) Deep-sea shrimps and lobsters (Crustacea: Decapoda: Dendrobranchiata and Pleocyemata) from the Sagami Sea and Izu Islands, central Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 47, 279 - 337." type="journal article" year="2011">Komai (2011)</bibRefCitation>
discussed on the close similarities between 
<taxonomicName id="39AB4D52CF4BFFA30B5DFF60FB12FF3A" box="[1150,1275,188,212]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4BFFA30B5DFF60FB12FF3A" box="[1150,1275,188,212]" italics="true" pageId="3" pageNumber="501">C. izuensis</emphasis>
</taxonomicName>
and the other three 
<taxonomicName id="39AB4D52CF4BFFA30FFAFF3CFED4FF16" box="[217,317,224,248]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FFAFF3CFED4FF16" box="[217,317,224,248]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
species, 
<taxonomicName id="39AB4D52CF4BFFA30E8AFF3CFDCEFF16" box="[425,551,224,248]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="CCDFEAC3CF4BFFA30E8AFF3CFDCEFF16" box="[425,551,224,248]" italics="true" pageId="3" pageNumber="501">C. kensleyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30D1AFF3CFD0DFF16" box="[569,740,224,248]" class="Malacostraca" family="Axiidae" genus="Axius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pailoloensis">
<emphasis id="CCDFEAC3CF4BFFA30D1AFF3CFD0DFF16" box="[569,740,224,248]" italics="true" pageId="3" pageNumber="501">C. pailoloensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4BFFA30C02FF3CFC75FF16" box="[801,924,224,248]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="CCDFEAC3CF4BFFA30C02FF3CFC75FF16" box="[801,924,224,248]" italics="true" pageId="3" pageNumber="501">C. sibogae</emphasis>
</taxonomicName>
, which were all transferred to 
<taxonomicName id="39AB4D52CF4BFFA30A24FF3CFA9CFF19" box="[1287,1397,224,247]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30A24FF3CFA9CFF19" box="[1287,1397,224,247]" italics="true" pageId="3" pageNumber="501">Manaxius</emphasis>
</taxonomicName>
by 
<bibRefCitation id="9A3A4B20CF4BFFA30FB4FED8FEC1FEF2" author="Sakai" box="[151,296,260,285]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
, but 
<bibRefCitation id="9A3A4B20CF4BFFA30E7DFED8FE06FEF2" author="Sakai" box="[350,495,260,285]" pageId="3" pageNumber="501" refString="Sakai, K. (2014) A review of Ambiaxiopsis altimanus Komai, 2011 and Calaxius izuensis Komai, 2011, with a new genus, Calaxidium gen. nov. (Decapoda, Pleocyemata, Callianassidea). Crustaceana, 87, 886 - 889. http: // dx. doi. org / 10.1163 / 15685403 - 00003313" type="journal article" year="2014">Sakai (2014)</bibRefCitation>
completely ignored Komai’s (2011) indication. 
<taxonomicName id="39AB4D52CF4BFFA30B29FED8FB64FEF2" box="[1034,1165,260,284]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30B29FED8FB64FEF2" box="[1034,1165,260,284]" italics="true" pageId="3" pageNumber="501">Calaxidium</emphasis>
</taxonomicName>
cannot be distinguished from 
<taxonomicName id="39AB4D52CF4BFFA30FF5FEF4FDC0FEAE" authority="Sakai (2011)" authorityName="Sakai" authorityYear="2011" box="[214,553,295,320]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FF5FEF4FEADFED1" box="[214,324,296,319]" italics="true" pageId="3" pageNumber="501">Manaxius</emphasis>
sensu 
<bibRefCitation id="9A3A4B20CF4BFFA30EB7FEFBFDC0FEAE" author="Sakai" box="[404,553,295,320]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
</taxonomicName>
. In conclusion, 
<taxonomicName id="39AB4D52CF4BFFA30DFCFEF4FC8BFEAE" box="[735,866,296,320]" class="Malacostraca" family="Axiidae" genus="Calaxidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30DFCFEF4FC8BFEAE" box="[735,866,296,320]" italics="true" pageId="3" pageNumber="501">Calaxidium</emphasis>
</taxonomicName>
is also here synonymized under 
<taxonomicName id="39AB4D52CF4BFFA30BFEFEF4FAA8FEAE" box="[1245,1345,296,320]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30BFEFEF4FAA8FEAE" box="[1245,1345,296,320]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
, and 
<taxonomicName id="39AB4D52CF4BFFA30AA1FEF4FF19FE8A" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4BFFA30AA1FEF4FF19FE8A" italics="true" pageId="3" pageNumber="501">C. izuensis</emphasis>
</taxonomicName>
is returned to 
<taxonomicName id="39AB4D52CF4BFFA30EB2FE90FE1CFE8A" box="[401,501,332,356]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30EB2FE90FE1CFE8A" box="[401,501,332,356]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FE1436D1CF4BFFA30FE4FEACFD9EFC4A" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="501">
Among 11 species referred to 
<taxonomicName id="39AB4D52CF4BFFA30D06FEACFD7AFE69" box="[549,659,368,391]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30D06FEACFD7AFE69" box="[549,659,368,391]" italics="true" pageId="3" pageNumber="501">Manaxius</emphasis>
</taxonomicName>
by 
<bibRefCitation id="9A3A4B20CF4BFFA30DE0FEB3FA8AFE66" author="Sakai" box="[707,1379,367,392]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">
Sakai (2011; except for the nomen nudum 
<taxonomicName id="39AB4D52CF4BFFA30B91FEADFAB1FE69" box="[1202,1368,368,392]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="aganaensis">
<emphasis id="CCDFEAC3CF4BFFA30B91FEADFAB1FE69" box="[1202,1368,368,392]" italics="true" pageId="3" pageNumber="501">M. aganaensis</emphasis>
</taxonomicName>
)
</bibRefCitation>
, two species do not fit the generic diagnosis of 
<taxonomicName id="39AB4D52CF4BFFA30DA2FE48FD0CFE42" box="[641,741,404,428]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30DA2FE48FD0CFE42" box="[641,741,404,428]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
. “ 
<taxonomicName id="39AB4D52CF4BFFA30C21FE49FC99FE42" box="[770,880,405,428]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30C21FE49FC99FE42" box="[770,880,405,428]" italics="true" pageId="3" pageNumber="501">Manaxius</emphasis>
</taxonomicName>
” 
<taxonomicName id="39AB4D52CF4BFFA30CA5FE48FB45FE42" authority="Coelho, 1973" authorityName="Coelho" authorityYear="1973" box="[902,1196,404,429]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="CCDFEAC3CF4BFFA30CA5FE48FC1DFE42" box="[902,1012,404,428]" italics="true" pageId="3" pageNumber="501">angulatus</emphasis>
(
<bibRefCitation id="9A3A4B20CF4BFFA30B25FE48FB4DFE42" author="Coelho" box="[1030,1188,404,429]" pageId="3" pageNumber="501" refString="Coelho, P. A. (1973) Descricao preliminar de Calastacus angulatus, n. sp., e de C. spinosus, n. sp., do norte do Brasil (Crustacea, Decapoda, Callianassidae). Ciencia e Cultura, 25 (Supplement 6), 344 - 345." type="journal article" year="1973">Coelho, 1973</bibRefCitation>
)
</taxonomicName>
, originally described from 
<collectingCountry id="86BC7641CF4BFFA30FFBFE6BFEF5FE3E" box="[216,284,439,464]" name="Brazil" pageId="3" pageNumber="501">Brazil</collectingCountry>
under 
<taxonomicName id="39AB4D52CF4BFFA30E51FE64FD65FE3E" authority="Faxon, 1893" authorityName="Faxon" authorityYear="1893" box="[370,652,440,464]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30E51FE64FE07FE3E" box="[370,494,440,464]" italics="true" pageId="3" pageNumber="501">Calastacus</emphasis>
<bibRefCitation id="9A3A4B20CF4BFFA30EDAFE64FD65FE3E" author="Faxon" box="[505,652,440,464]" pageId="3" pageNumber="501" refString="Faxon, W. (1893) Reports on the dredging operations off the west coast of Central America to the Galapagos, to the west coast of Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried on by the U. S. Fish Commission Steamer Albatross during 1891, lieut. Commander Z. L. Tanner, U. S. N., commanding, 6: Preliminary description of new species of Crustacea. Bulletin of the Museum of Comparative Zoology at Harvard College, 24, 149 - 220." type="journal article" year="1893">Faxon, 1893</bibRefCitation>
</taxonomicName>
, is not consistent with the generic diagnosis of 
<taxonomicName id="39AB4D52CF4BFFA30BE3FE64FACDFE3E" box="[1216,1316,440,464]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30BE3FE64FACDFE3E" box="[1216,1316,440,464]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
in having elongate eyestalks overreaching the rostral apex (
<bibRefCitation id="9A3A4B20CF4BFFA30DE3FE00FBE6FE1A" author="Coelho" box="[704,1039,476,501]" pageId="3" pageNumber="501" refString="Coelho, P. A. &amp; Ramos-Porto, M. (1991) Sinopse dos crustaceos decapodos brasileiros (familias Scyllaridae, Palinuridae, Nephropidae, Parastacidae e Axiidae). Anais da Universidade Federal Rural de Pernambuco, Ciencias Biologicas, 8 / 10, 47 - 88." type="journal article" year="1991">Coelho &amp; Ramos-Porto 1991</bibRefCitation>
). 
<bibRefCitation id="9A3A4B20CF4BFFA30B01FE00FAECFE1A" box="[1058,1285,476,501]" pageId="3" pageNumber="501" refString="Coelho, P. A., de Almeida, A. O., Bezerra, L. E. A. &amp; de Souza-Filho, J. F. (2007) An updated checklist of decapod crustaceans (infraorders Astacidea, Thalassinidea, Polychelida, Palinura, and Anomura) from the northern and northeastern Brazilian coast. Zootaxa, 1519, 1 - 16." type="journal article">
Coelho 
<emphasis id="CCDFEAC3CF4BFFA30B59FE01FB45FE1A" box="[1146,1196,476,500]" italics="true" pageId="3" pageNumber="501">et al</emphasis>
. (2007)
</bibRefCitation>
had assigned this species to 
<taxonomicName id="39AB4D52CF4BFFA30E73FDDCFC99FDF6" authority="Sakai &amp; de Saint Laurent, 1989" authorityName="Sakai &amp; de Saint Laurent" authorityYear="1989" box="[336,880,511,536]" class="Malacostraca" family="Axiidae" genus="Acanthaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30E73FDDCFE32FDF6" box="[336,475,512,536]" italics="true" pageId="3" pageNumber="501">Acanthaxius</emphasis>
Sakai &amp; de 
<bibRefCitation id="9A3A4B20CF4BFFA30DA2FE23FC99FDF6" author="Saint" box="[641,880,511,536]" pageId="3" pageNumber="501" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article" year="1989">Saint Laurent, 1989</bibRefCitation>
</taxonomicName>
. Reexamination of the 
<typeStatus id="21108873CF4BFFA30BACFDDCFB56FDF6" box="[1167,1215,512,536]" pageId="3" pageNumber="501">type</typeStatus>
material, the sole representative of the species, is necessary to determine the generic status of 
<taxonomicName id="39AB4D52CF4BFFA30B07FDF8FAF3FDD2" box="[1060,1306,548,572]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="CCDFEAC3CF4BFFA30B07FDF8FAF3FDD2" box="[1060,1306,548,572]" italics="true" pageId="3" pageNumber="501">Calastacus angulatus</emphasis>
</taxonomicName>
. 
<taxonomicName id="39AB4D52CF4BFFA30A0EFDF9FE52FD8E" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="CCDFEAC3CF4BFFA30A0EFDF9FEC5FD8E" italics="true" pageId="3" pageNumber="501">Manaxius thailandensis</emphasis>
<bibRefCitation id="9A3A4B20CF4BFFA30E16FD9BFE52FD8E" author="Sakai" box="[309,443,583,608]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
was founded on the basis of two specimens (
<typeStatus id="21108873CF4BFFA30CEFFD9BFBD9FD8E" box="[972,1072,583,608]" pageId="3" pageNumber="501" type="holotype">holotype</typeStatus>
male and one 
<typeStatus id="21108873CF4BFFA30BFEFD94FAD4FD8E" box="[1245,1341,584,608]" pageId="3" pageNumber="501" type="paratype">paratype</typeStatus>
female) from the Gulf of 
<collectingCountry id="86BC7641CF4BFFA30E46FDB0FE22FD6B" box="[357,459,620,645]" name="Thailand" pageId="3" pageNumber="501">Thailand</collectingCountry>
, originally identified as 
<taxonomicName id="39AB4D52CF4BFFA30DC8FDB0FC7AFD6A" box="[747,915,620,644]" class="Malacostraca" family="Axiidae" genus="Eutrichocheles" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30DC8FDB0FC7AFD6A" box="[747,915,620,644]" italics="true" pageId="3" pageNumber="501">Eutrichocheles</emphasis>
</taxonomicName>
sp. by 
<bibRefCitation id="9A3A4B20CF4BFFA30CCCFDB0FB6FFD6A" author="Sakai" box="[1007,1158,620,645]" pageId="3" pageNumber="501" refString="Sakai, K. (1992) Axiid collections of the Zoological Museum, Copenhagen, with the description of one new genus and six new species (Axiidae, Thalassinidea, Crustacea). Zoologica Scripta, 21, 157 - 180. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00318. x" type="journal article" year="1992">Sakai (1992)</bibRefCitation>
. This species better fits 
<taxonomicName id="39AB4D52CF4BFFA30FB4FD4CFE2DFD46" authority="de Man, 1905" authorityName="de Man" authorityYear="1905" box="[151,452,655,680]" class="Malacostraca" family="Axiidae" genus="Paraxiopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FB4FD4CFEF4FD49" box="[151,285,656,679]" italics="true" pageId="3" pageNumber="501">Paraxiopsis</emphasis>
de 
<bibRefCitation id="9A3A4B20CF4BFFA30E6AFD4CFE2DFD46" author="Man" box="[329,452,655,680]" pageId="3" pageNumber="501" refString="Man, J. G. de (1905) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 9, 587 - 614." type="journal article" year="1905">
<collectingCountry id="86BC7641CF4BFFA30E6AFD4CFE96FD49" box="[329,383,656,679]" name="Isle of Man" pageId="3" pageNumber="501">Man</collectingCountry>
, 
<quantity id="39539B34CF4BFFA30EAFFD53FE2DFD46" box="[396,452,655,680]" metricMagnitude="1" metricUnit="m" metricValue="4.8387" pageId="3" pageNumber="501" unit="in" value="1905.0">1905</quantity>
</bibRefCitation>
</taxonomicName>
in having a small, bispinose antennal scaphocerite and the slender cheliped with nonsetose chela lacking dorsal spines on the palm (cf. 
<bibRefCitation id="9A3A4B20CF4BFFA30DF0FD68FC9BFD23" author="Kensley" box="[723,882,692,717]" pageId="3" pageNumber="501" refString="Kensley, B. (1996) The genus Paraxiopsis De Man, with descriptions of new species from the Western Atlantic (Crustacea: Decapoda: Axiidae). Bulletin of Marine Science, 58, 709 - 729." type="journal article" year="1996">Kensley 1996</bibRefCitation>
). Sakai’s (2011) interpretation on the structure of the antennal scaphocerite is not consistent with his previous description (
<bibRefCitation id="9A3A4B20CF4BFFA30CFEFD0BFB8BFD1E" author="Sakai" box="[989,1122,727,752]" pageId="3" pageNumber="501" refString="Sakai, K. (1992) Axiid collections of the Zoological Museum, Copenhagen, with the description of one new genus and six new species (Axiidae, Thalassinidea, Crustacea). Zoologica Scripta, 21, 157 - 180. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00318. x" type="journal article" year="1992">Sakai 1992</bibRefCitation>
). The lack of a supraocular spine also excludes 
<taxonomicName id="39AB4D52CF4BFFA30EAFFD21FD77FCFA" box="[396,670,764,788]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="CCDFEAC3CF4BFFA30EAFFD21FD77FCFA" box="[396,670,764,788]" italics="true" pageId="3" pageNumber="501">Manaxius thailandensis</emphasis>
</taxonomicName>
from 
<taxonomicName id="39AB4D52CF4BFFA30DD2FD20FCBCFCFA" box="[753,853,764,788]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30DD2FD20FCBCFCFA" box="[753,853,764,788]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
. 
<taxonomicName id="39AB4D52CF4BFFA30C49FD21FB99FCFA" box="[874,1136,764,788]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="ohsumiensis">
<emphasis id="CCDFEAC3CF4BFFA30C49FD21FB99FCFA" box="[874,1136,764,788]" italics="true" pageId="3" pageNumber="501">Manaxius ohsumiensis</emphasis>
</taxonomicName>
, described from Ohsumi Islands, located to the north of the Ryukyu Islands, 
<collectingCountry id="86BC7641CF4BFFA30DD1FCFCFCDCFCD9" box="[754,821,800,823]" name="Japan" pageId="3" pageNumber="501">Japan</collectingCountry>
, appears very similar to 
<taxonomicName id="39AB4D52CF4BFFA30B76FCFCFEA0FCB2" authority="Kensley (2003)" authorityName="Kensley" authorityYear="2003" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="CCDFEAC3CF4BFFA30B76FCFCFABDFCD9" box="[1109,1364,800,823]" italics="true" pageId="3" pageNumber="501">Manaxius pitatucensis</emphasis>
sensu 
<bibRefCitation id="9A3A4B20CF4BFFA30FB4FC98FEA0FCB2" author="Kensley" box="[151,329,836,861]" pageId="3" pageNumber="501" refString="Kensley, B. (2003) Axioid shrimps from Guam (Crustacea, Decapoda, Thalassinidea). Micronesica, 35 - 36, 359 - 384." type="book chapter" year="2003">Kensley (2003)</bibRefCitation>
</taxonomicName>
, and it is likely that the two taxa are conspecific. Nevertheless, it is advisable to reexamine the material studied by 
<bibRefCitation id="9A3A4B20CF4BFFA30EA1FCBBFDDDFC6E" author="Kensley" box="[386,564,871,896]" pageId="3" pageNumber="501" refString="Kensley, B. (2003) Axioid shrimps from Guam (Crustacea, Decapoda, Thalassinidea). Micronesica, 35 - 36, 359 - 384." type="book chapter" year="2003">Kensley (2003)</bibRefCitation>
and 
<bibRefCitation id="9A3A4B20CF4BFFA30D57FCBBFCEEFC6E" author="Sakai" box="[628,775,871,896]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
to reach final conclusion. Here I tentatively include 
<taxonomicName id="39AB4D52CF4BFFA30A5EFCB5FEC9FC4A" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="ohsumiensis">
<emphasis id="CCDFEAC3CF4BFFA30A5EFCB5FEC9FC4A" italics="true" pageId="3" pageNumber="501">M. ohsumiensis</emphasis>
</taxonomicName>
as a valid species in 
<taxonomicName id="39AB4D52CF4BFFA30D2FFC50FD99FC4A" box="[524,624,908,932]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30D2FFC50FD99FC4A" box="[524,624,908,932]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B6B1655ACF4BFFA30FE4FC6CFDA5F8A2" pageId="3" pageNumber="501" type="reference_group">
<paragraph id="FE1436D1CF4BFFA30FE4FC6CFDCBFADE" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="501">
<taxonomicName id="39AB4D52CF4BFFA30FE4FC6CFDE8FC26" ID-CoL="43ee3471-58ca-4fcb-b6a1-78bbfc7b60a9" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[199,513,943,968]" class="Malacostraca" family="Axiidae" genus="Coelhocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FE4FC6CFE9CFC26" box="[199,373,944,968]" italics="true" pageId="3" pageNumber="501">Coelhocalaxius</emphasis>
<bibRefCitation id="9A3A4B20CF4BFFA30E58FC73FDE8FC26" author="Sakai" box="[379,513,943,968]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
, which was established to accommodate 
<taxonomicName id="39AB4D52CF4BFFA30CE8FC6CFB47FC29" box="[971,1198,944,968]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="CCDFEAC3CF4BFFA30CE8FC6CFB47FC29" box="[971,1198,944,968]" italics="true" pageId="3" pageNumber="501">Calastacus spinosus</emphasis>
</taxonomicName>
with 
<taxonomicName id="39AB4D52CF4BFFA30BCEFC6CFA97FC26" ID-CoL="ef3ffec1-e80c-4a07-84cc-dbf557f541d4" box="[1261,1406,944,968]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="oxypleura">
<emphasis id="CCDFEAC3CF4BFFA30BCEFC6CFA97FC26" box="[1261,1406,944,968]" italics="true" pageId="3" pageNumber="501">C. oxypleura</emphasis>
</taxonomicName>
as a junior synonym, was said to differ from 
<taxonomicName id="39AB4D52CF4BFFA30D59FC08FD37FC02" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[634,734,980,1004]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30D59FC08FD37FC02" box="[634,734,980,1004]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
in having a narrow rather than a triangular rostrum and more elongate eyestalks. However, the rostrum varies considerably between species. Sakai &amp; de 
<bibRefCitation id="9A3A4B20CF4BFFA30B8DFC2BFA75FBFE" box="[1198,1436,1015,1040]" pageId="3" pageNumber="501" refString="Sakai, K. &amp; Saint Laurent, M. de (1989) A check list of Axiidae (Decapoda, Crustacea, Thalassinidea, Anomula), with remarks and in addition descriptions of one new subfamily, eleven new genera and two new species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 3, 1 - 104." type="journal article">Saint Laurent (1989)</bibRefCitation>
noted that the eyestalks of species of 
<taxonomicName id="39AB4D52CF4BFFA30D65FBC0FD43FBDA" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[582,682,1052,1076]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30D65FBC0FD43FBDA" box="[582,682,1052,1076]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
were ‘rather elongate’, another variable character of little generic value. Spination of the cheliped palm is the same in both genera. The only species included, 
<taxonomicName id="39AB4D52CF4BFFA30B93FB9CFA7DFBB9" box="[1200,1428,1088,1112]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="CCDFEAC3CF4BFFA30B93FB9CFA7DFBB9" box="[1200,1428,1088,1112]" italics="true" pageId="3" pageNumber="501">Calastacus spinosus</emphasis>
</taxonomicName>
, has a postcervical carina, said by 
<bibRefCitation id="9A3A4B20CF4BFFA30D36FBB8FD4CFB92" author="Sakai" box="[533,677,1124,1149]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
to set it apart from 
<taxonomicName id="39AB4D52CF4BFFA30CAAFBB8FC04FB92" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[905,1005,1124,1148]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30CAAFBB8FC04FB92" box="[905,1005,1124,1148]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
but as shown above it is not the case. Nevertheless, 
<taxonomicName id="39AB4D52CF4BFFA30E16FB54FDF1FB71" box="[309,536,1160,1184]" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="CCDFEAC3CF4BFFA30E16FB54FDF1FB71" box="[309,536,1160,1184]" italics="true" pageId="3" pageNumber="501">Calastacus spinosus</emphasis>
</taxonomicName>
seems to differ from any species assigned to 
<taxonomicName id="39AB4D52CF4BFFA30B33FB54FB9DFB4E" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[1040,1140,1160,1184]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30B33FB54FB9DFB4E" box="[1040,1140,1160,1184]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
in the lack of conspicuous supraocular spines and the presence of two pairs of lateral longitudinal carinae on the second to fifth pleomeres (
<bibRefCitation id="9A3A4B20CF4BFFA30FBDFB13FEC6FB06" author="Sakai" box="[158,303,1231,1256]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
). Consequently, 
<taxonomicName id="39AB4D52CF4BFFA30D20FB0CFD58FB06" ID-CoL="43ee3471-58ca-4fcb-b6a1-78bbfc7b60a9" box="[515,689,1232,1256]" class="Malacostraca" family="Axiidae" genus="Coelhocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30D20FB0CFD58FB06" box="[515,689,1232,1256]" italics="true" pageId="3" pageNumber="501">Coelhocalaxius</emphasis>
</taxonomicName>
is maintained as a valid genus. The synonymy between 
<taxonomicName id="39AB4D52CF4BFFA30FB4FB28FE46FAE2" ID-CoL="32ea5e37-4514-436c-89cc-c8207d076327" box="[151,431,1268,1292]" class="Malacostraca" family="Axiidae" genus="Coelhocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="CCDFEAC3CF4BFFA30FB4FB28FE46FAE2" box="[151,431,1268,1292]" italics="true" pageId="3" pageNumber="501">Coelhocalaxius spinosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4BFFA30ECFFB28FD69FAE2" ID-CoL="ef3ffec1-e80c-4a07-84cc-dbf557f541d4" box="[492,640,1268,1292]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="oxypleura">
<emphasis id="CCDFEAC3CF4BFFA30ECFFB28FD69FAE2" box="[492,640,1268,1292]" italics="true" pageId="3" pageNumber="501">C. oxypleura</emphasis>
</taxonomicName>
is not very convincing, because the latter differs from the former in having broader telson and uropods.
</paragraph>
<paragraph id="FE1436D1CF4BFFA30FE4FAE0FE19F979" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="501">
<taxonomicName id="39AB4D52CF4BFFA30FE4FAE0FDE8FABA" ID-CoL="4ea4cff3-bb44-415d-b783-f7e1cae50d44" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[199,513,1340,1365]" class="Malacostraca" family="Axiidae" genus="Heterocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FE4FAE0FE98FABA" box="[199,369,1340,1364]" italics="true" pageId="3" pageNumber="501">Heterocalaxius</emphasis>
<bibRefCitation id="9A3A4B20CF4BFFA30E5AFAE0FDE8FABA" author="Sakai" box="[377,513,1340,1365]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
, which was established for 
<taxonomicName id="39AB4D52CF4BFFA30C1EFAE0FC14FABA" ID-CoL="a54df6a2-39e4-4de4-95cc-6dd1970d1280" box="[829,1021,1340,1364]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="carneyi">
<emphasis id="CCDFEAC3CF4BFFA30C1EFAE0FC14FABA" box="[829,1021,1340,1364]" italics="true" pageId="3" pageNumber="501">Calaxius carneyi</emphasis>
</taxonomicName>
, was said to differ from 
<taxonomicName id="39AB4D52CF4BFFA30A39FAE0FA97FABA" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[1306,1406,1340,1364]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30A39FAE0FA97FABA" box="[1306,1406,1340,1364]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
in the lack of an appendix masculina on the male second pleopod, the lack of a postcervical carina on the carapace, the short scaphocerite, the major cheliped palm and carpus weakly tuberculate dorsally, the minor cheliped palm and carpus more tuberculate dorsally than in the major cheliped, the cheliped fingers being shorter than the palms, and the triangular telson (
<bibRefCitation id="9A3A4B20CF4BFFA30EE0FA10FDA3FA0A" author="Sakai" box="[451,586,1484,1509]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai 2011</bibRefCitation>
). Of these characters, the development of the postcervical carina and the length of the cheliped fingers are variable characters of little generic value, as discussed above, but the other characters seem to be still significant in recognizing 
<taxonomicName id="39AB4D52CF4BFFA30DC9F9C8FC7DF9C2" ID-CoL="4ea4cff3-bb44-415d-b783-f7e1cae50d44" box="[746,916,1556,1580]" class="Malacostraca" family="Axiidae" genus="Heterocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30DC9F9C8FC7DF9C2" box="[746,916,1556,1580]" italics="true" pageId="3" pageNumber="501">Heterocalaxius</emphasis>
</taxonomicName>
. With regard to the statement on the armature of the cheliped palms and carpi by 
<bibRefCitation id="9A3A4B20CF4BFFA30D1EF9EBFD39F9BE" author="Sakai" box="[573,720,1591,1616]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
is not specific; the lack of conspicuous spines on the dorsal margins of the cheliped palms and carpi, not the condition of tuberculation, is significant in differentiating 
<taxonomicName id="39AB4D52CF4BFFA30FB4F95CFEA8F976" ID-CoL="4ea4cff3-bb44-415d-b783-f7e1cae50d44" box="[151,321,1664,1688]" class="Malacostraca" family="Axiidae" genus="Heterocalaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30FB4F95CFEA8F976" box="[151,321,1664,1688]" italics="true" pageId="3" pageNumber="501">Heterocalaxius</emphasis>
</taxonomicName>
from 
<taxonomicName id="39AB4D52CF4BFFA30EA6F95CFE00F976" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[389,489,1664,1688]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30EA6F95CFE00F976" box="[389,489,1664,1688]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FE1436D1CF4BFFA30FE4F979FDA5F8A2" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="501">
In conclusion, the following 14 specific taxa are recognized in 
<taxonomicName id="39AB4D52CF4BFFA30CADF978FC1BF952" ID-CoL="53098ce0-843a-4a54-92f8-6b2b1087f095" box="[910,1010,1700,1724]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="CCDFEAC3CF4BFFA30CADF978FC1BF952" box="[910,1010,1700,1724]" italics="true" pageId="3" pageNumber="501">Calaxius</emphasis>
</taxonomicName>
for the time being: 
<taxonomicName id="39AB4D52CF4BFFA30BF1F978FA9DF952" ID-CoL="f7ca06e2-ab5e-4741-a484-d738bc7fed34" box="[1234,1396,1700,1724]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="acutirostris">
<emphasis id="CCDFEAC3CF4BFFA30BF1F978FA9DF952" box="[1234,1396,1700,1724]" italics="true" pageId="3" pageNumber="501">C. acutirostris</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30AA1F978FEDDF90E" class="Malacostraca" family="Calocarididae" genus="Calastacus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="euophthalmus">
<emphasis id="CCDFEAC3CF4BFFA30AA1F978FEDDF90E" italics="true" pageId="3" pageNumber="501">C. euophthalmus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30E61F914FE1EF90E" ID-CoL="f7400038-cbde-473e-9eea-901cbe3e01c2" box="[322,503,1736,1760]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="CCDFEAC3CF4BFFA30E61F914FE1EF90E" box="[322,503,1736,1760]" italics="true" pageId="3" pageNumber="501">C. galapagensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30D26F914FD73F90E" ID-CoL="1046e3f9-1e0c-436e-b488-fa205bf6fefb" box="[517,666,1736,1760]" class="Malacostraca" family="Axiidae" genus="Axius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="CCDFEAC3CF4BFFA30D26F914FD73F90E" box="[517,666,1736,1760]" italics="true" pageId="3" pageNumber="501">C. inaequalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30D8AF914FCCAF931" ID-CoL="9e4f1187-2e9b-44c9-820d-52ba4494b459" box="[681,803,1736,1759]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="izuensis">
<emphasis id="CCDFEAC3CF4BFFA30D8AF914FCCAF931" box="[681,803,1736,1759]" italics="true" pageId="3" pageNumber="501">C. izuensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30C12F914FC48F931" box="[817,929,1736,1759]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="jenneri">
<emphasis id="CCDFEAC3CF4BFFA30C12F914FC48F931" box="[817,929,1736,1759]" italics="true" pageId="3" pageNumber="501">C. jenneri</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30C8CF914FBC3F90E" box="[943,1066,1736,1760]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="kensleyi">
<emphasis id="CCDFEAC3CF4BFFA30C8CF914FBC3F90E" box="[943,1066,1736,1760]" italics="true" pageId="3" pageNumber="501">C. kensleyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30B1AF914FB2DF931" ID-CoL="ce165e4e-12b0-4b9f-aec1-5e2e0a7f1b2e" box="[1081,1220,1736,1759]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="manningi">
<emphasis id="CCDFEAC3CF4BFFA30B1AF914FB2DF931" box="[1081,1220,1736,1759]" italics="true" pageId="3" pageNumber="501">C. manningi</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30BF1F914FA9DF931" box="[1234,1396,1736,1759]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="mimasensis">
<emphasis id="CCDFEAC3CF4BFFA30BF1F914FA9DF931" box="[1234,1396,1736,1759]" italics="true" pageId="3" pageNumber="501">C. mimasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30AA1F914FF05F8EA" ID-CoL="b0d94978-780b-498d-a004-915f98f13f05" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="poupini" status="sp. nov.">
<emphasis id="CCDFEAC3CF4BFFA30AA1F914FF05F8EA" italics="true" pageId="3" pageNumber="501">C. poupini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7EC57B8CF4BFFA30FD7F930FEDDF8EB" box="[244,308,1772,1797]" pageId="3" pageNumber="501" rank="species">
<emphasis id="CCDFEAC3CF4BFFA30FD7F930FEDDF8EB" bold="true" box="[244,308,1772,1797]" pageId="3" pageNumber="501">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="39AB4D52CF4BFFA30E67F930FE06F8EA" ID-CoL="90401c93-3848-4ce5-91c5-f9848993d870" box="[324,495,1772,1796]" class="Malacostraca" family="Axiidae" genus="Manaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="ohsumiensis" status="comb. nov.">
<emphasis id="CCDFEAC3CF4BFFA30E67F930FE06F8EA" box="[324,495,1772,1796]" italics="true" pageId="3" pageNumber="501">C. ohsumiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7EC57B8CF4BFFA30ED4F931FDB4F8EB" box="[503,605,1772,1797]" pageId="3" pageNumber="501">n. comb.</taxonomicNameLabel>
, 
<taxonomicName id="39AB4D52CF4BFFA30D4AF930FCFBF8EA" ID-CoL="be49664e-8048-4d42-8056-c7595e170ad2" box="[617,786,1772,1796]" class="Malacostraca" family="Axiidae" genus="Axius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="pailoloensis">
<emphasis id="CCDFEAC3CF4BFFA30D4AF930FCFBF8EA" box="[617,786,1772,1796]" italics="true" pageId="3" pageNumber="501">C. pailoloensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39AB4D52CF4BFFA30C01F930FC72F8EA" box="[802,923,1772,1796]" class="Malacostraca" family="Calocarididae" genus="Calocaris" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="CCDFEAC3CF4BFFA30C01F930FC72F8EA" box="[802,923,1772,1796]" italics="true" pageId="3" pageNumber="501">C. sibogae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="39AB4D52CF4BFFA30CF8F930FBDCF8EA" ID-CoL="d942c861-3369-4155-98f8-2a0285b40e07" box="[987,1077,1772,1796]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="tungi">
<emphasis id="CCDFEAC3CF4BFFA30CF8F930FBDCF8EA" box="[987,1077,1772,1796]" italics="true" pageId="3" pageNumber="501">C. tungi</emphasis>
</taxonomicName>
. 
<collectingCountry id="86BC7641CF4BFFA30B66F930FB8DF8EA" box="[1093,1124,1772,1796]" name="American Samoa" pageId="3" pageNumber="501">As</collectingCountry>
discussed by 
<bibRefCitation id="9A3A4B20CF4BFFA30A26F930FA7EF8EA" author="Sakai" box="[1285,1431,1772,1797]" pageId="3" pageNumber="501" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). Crustaceana Monographs, 13, 1 - 616. http: // dx. doi. org / 10.1163 / 9789047424185" type="journal article" year="2011">Sakai (2011)</bibRefCitation>
, however, relationship between 
<taxonomicName id="39AB4D52CF4BFFA30ED0F8CCFD94F8C9" ID-CoL="ce165e4e-12b0-4b9f-aec1-5e2e0a7f1b2e" box="[499,637,1808,1831]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="manningi">
<emphasis id="CCDFEAC3CF4BFFA30ED0F8CCFD94F8C9" box="[499,637,1808,1831]" italics="true" pageId="3" pageNumber="501">C. manningi</emphasis>
</taxonomicName>
and 
<taxonomicName id="39AB4D52CF4BFFA30D91F8CCFCE4F8C9" ID-CoL="d942c861-3369-4155-98f8-2a0285b40e07" box="[690,781,1808,1831]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="501" phylum="Arthropoda" rank="species" species="tungi">
<emphasis id="CCDFEAC3CF4BFFA30D91F8CCFCE4F8C9" box="[690,781,1808,1831]" italics="true" pageId="3" pageNumber="501">C. tungi</emphasis>
</taxonomicName>
is not clear, and further comparison would be necessary to establish the identity of these two taxa.
</paragraph>
</subSubSection>
</treatment>
</document>