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<document id="03568192AD8B7719957E90CF94A54F70" ID-CLB-Dataset="274285" ID-DOI="10.5852/ejt.2023.893.2285" ID-GBIF-Dataset="e441bc9d-bb44-4ee6-b29f-a6724b96592b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8385630" ID-ZooBank="D9DA4879-BC1D-4947-96E8-7D574DE334BA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1695891225180" checkinUser="plazi" docAuthor="Leng, Zhaoqi, Tran, Anh Duc &amp; Ye, Zhen" docDate="2023" docId="045E878DFFF329093A06FBC974DF0DA2" docLanguage="en" docName="EJT.2023.893.1-73.pdf" docOrigin="European Journal of Taxonomy 893 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2285/9863" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyacobates lundbladi" docType="treatment" docVersion="10" lastPageNumber="51" masterDocId="F867FFF5FFDC293B381FFFB270530F1A" masterDocTitle="Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="48" updateTime="1699690649759" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C4DDE6702361F0D7FFC19D60A6C068C1">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="74DCB11690CC96D841475EBB5339F1D2">Leng, Zhaoqi</mods:namePart>
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<mods:affiliation id="53272F26360E353C880C61416E5EC74A">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<heading id="D70081F7FFF329143A06FBC974490B8C" box="[537,1050,1147,1174]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="48" reason="2">
<taxonomicName id="4BF74D18FFF329143A06FBC974490B8C" ID-CoL="4SNHR" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[537,1050,1147,1174]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFF329143A06FBC9737D0B8C" bold="true" box="[537,814,1147,1174]" italics="true" pageId="47" pageNumber="48">Rhyacobates lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFF329143B23FBC974410B8C" author="Hungerford" box="[828,1042,1147,1174]" firstAuthor="Hungerford" pageId="47" pageNumber="48" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
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<paragraph id="8C48369BFFF329143A7FFB2C73870BA3" blockId="47.[537,1050,1147,1209]" box="[608,980,1182,1209]" pageId="47" pageNumber="48">
<figureCitation id="14CC2A1EFFF329143A7FFB2C72ED0BA3" box="[608,702,1182,1209]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[336,1266,297,1835]" captionTargetId="figure-14@4.[255,1333,265,1851]" captionTargetPageId="4" captionText="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385634" httpUri="https://zenodo.org/record/8385634/files/figure.png" pageId="47" pageNumber="48">Figs 2C</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143AD5FB2D72BE0BA3" box="[714,749,1183,1209]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">3L</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143AE5FB2D73460BA3" box="[762,789,1183,1209]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="47" pageNumber="48">4I</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143B3EFB2C73140BA3" box="[801,839,1182,1209]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="47" pageNumber="48">5K</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143B4CFB2D732B0BA3" box="[851,888,1183,1209]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="47" pageNumber="48">7B</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143B9BFB2D73F90BA3" box="[900,938,1183,1209]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="47" pageNumber="48">8H</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143BA8FB2D73870BA3" box="[951,980,1183,1209]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">29</figureCitation>
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<treatmentCitation id="0D56108AFFF3291438A2FB5272BF0BE0" author="Hungerford H. B." box="[189,748,1248,1275]" page="36" pageId="47" pageNumber="48" year="1957">
<taxonomicName id="4BF74D18FFF3291438A2FB5272BF0BE0" ID-CoL="3BGBD" authority="Hungerford, 1957: 33 - 36" authorityName="Hungerford" authorityPageNumber="33 - 36" authorityYear="1957" box="[189,748,1248,1275]" class="Insecta" family="Gerridae" genus="Esakobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFF3291438A2FB5271E90BE0" box="[189,442,1248,1274]" italics="true" pageId="47" pageNumber="48">Esakobates lundbladi</emphasis>
<bibRefCitation id="E8664B6AFFF3291439DDFB5272BF0BE0" author="Hungerford" box="[450,748,1248,1275]" firstAuthor="Hungerford" page="33 - 36" pageId="47" pageNumber="48" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957: 3336</bibRefCitation>
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(original description).
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<emphasis id="BE83EA89FFF3291438A2FA93719B0A21" box="[189,456,1313,1339]" italics="true" pageId="47" pageNumber="48">Rhyacobates lundbladi</emphasis>
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<treatmentCitation id="0D56108AFFF3291439FEFA9373D80A21" author="Hungerford H. B. &amp; Matsuda R." box="[481,907,1313,1340]" page="69" pageId="47" pageNumber="48" year="1959">
<bibRefCitation id="E8664B6AFFF3291439FEFA9373D80A21" author="Hungerford &amp; Matsuda" box="[481,907,1313,1340]" firstAuthor="Hungerford" page="69 - 72" pageId="47" pageNumber="48" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda 1959: 6972</bibRefCitation>
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. —
<treatmentCitation id="0D56108AFFF329143BA9FA9374820A21" author="Matsuda R." box="[950,1233,1313,1339]" page="273" pageId="47" pageNumber="48" year="1960">
<bibRefCitation id="E8664B6AFFF329143BA9FA9374820A21" author="Matsuda" box="[950,1233,1313,1339]" firstAuthor="Matsuda" page="273 - 276" pageId="47" pageNumber="48" pagination="25 - 632" refId="ref33267" refString="Matsuda R. 1960. Morphology, evolution and a classification of the Gerridae (Hemiptera-Heteroptera). The University of Kansas Science Bulletin 41: 25 - 632. https: // doi. org / 10.5962 / bhl. part. 15602" type="journal article" year="1960">Matsuda 1960: 273276</bibRefCitation>
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, figs 659, 661, 665, 667, 669672, 677, 679680. —
<treatmentCitation id="0D56108AFFF329143AB7FAF6744A0A44" author="Andersen N. M. &amp; Chen P. P." box="[680,1049,1348,1375]" page="61" pageId="47" pageNumber="48" year="1995">
<bibRefCitation id="E8664B6AFFF329143AB7FAF6744A0A44" author="Andersen &amp; Chen" box="[680,1049,1348,1375]" firstAuthor="Andersen" page="61 - 62" pageId="47" pageNumber="48" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen 1995: 6162</bibRefCitation>
</treatmentCitation>
, figs 2627 (with remarks).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFF3291438A2FA3475270835" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8C48369BFFF3291438A2FA3471650ABA" blockId="47.[189,310,1414,1440]" box="[189,310,1414,1440]" pageId="47" pageNumber="48">
<heading id="D70081F7FFF3291438A2FA3471650ABA" bold="true" box="[189,310,1414,1440]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="BE83EA89FFF3291438A2FA3471650ABA" bold="true" box="[189,310,1414,1440]" pageId="47" pageNumber="48">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFF3291438A2FA0675270835" blockId="47.[189,1401,1460,1839]" pageId="47" pageNumber="48">
Body length of apterous females 11.6212.21, of apterous males 8.028.41. Female: pronotum mainly black with a median brownish-yellow spot; mesonotum and metanotum mainly black with a median brownish-yellow stripe (
<figureCitation id="14CC2A1EFFF3291439C0FA49726C090F" box="[479,575,1531,1557]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Figs 3L</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143A52FA4972D7090F" box="[589,644,1531,1557]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">29A</figureCitation>
); yellowish subtriangular marking on metanotum similar to the marking on pronotum (
<figureCitation id="14CC2A1EFFF3291439C6F9AC72680922" box="[473,571,1566,1592]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Figs 3L</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143A54F9AC72D20922" box="[587,641,1566,1592]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">29A</figureCitation>
); posterior margin of abdominal mediotergite VII with a short median process (
<figureCitation id="14CC2A1EFFF32914399DF9F372410941" box="[386,530,1601,1627]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">Fig. 29A, D</figureCitation>
); abdominal segment VII with four distinct processes, including two lobe-like processes on each connexival corner, consisting of one broader, posteriorly directed outer process and a narrower inner process (
<figureCitation id="14CC2A1EFFF329143A2BF93A729609B8" box="[564,709,1672,1698]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">Fig. 29DF</figureCitation>
); posterior margin of sternum VII with a small, rounded median process (
<figureCitation id="14CC2A1EFFF32914399BF91971BC09DF" box="[388,495,1707,1733]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">Fig. 29E</figureCitation>
). Male: middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
<figureCitation id="14CC2A1EFFF329143A69F97C728E09F2" box="[630,733,1742,1768]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">Fig. 29J</figureCitation>
); length of middle tibia ca 1.51.6 times length of hind tibia; proctiger relatively large, with angular lobes laterally (
<figureCitation id="14CC2A1EFFF329143B9DF94373B10816" box="[898,994,1777,1804]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="47" pageNumber="48">Figs 7B</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143BF1F94374770811" box="[1006,1060,1777,1803]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">29K</figureCitation>
); paramere relatively slender and sinuate, middle part thickened, distal part tapering towards narrowly rounded apex (
<figureCitation id="14CC2A1EFFF329143CD9F8A675740835" box="[1222,1319,1812,1839]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="47" pageNumber="48">Figs 8H</figureCitation>
,
<figureCitation id="14CC2A1EFFF329143D2BF8A7753B0835" box="[1332,1384,1813,1839]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="47" pageNumber="48">29L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFF3290B38A2F8E471AE0E9D" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8C48369BFFF3291438A2F8E471F9086A" blockId="47.[189,426,1878,1904]" box="[189,426,1878,1904]" pageId="47" pageNumber="48">
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<emphasis id="BE83EA89FFF3291438A2F8E471F9086A" bold="true" box="[189,426,1878,1904]" pageId="47" pageNumber="48">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFF3291438F5F83771B70884" blockId="47.[189,1398,1924,2022]" box="[234,484,1924,1951]" pageId="47" pageNumber="48">
<emphasis id="BE83EA89FFF3291438F5F83771B70884" bold="true" box="[234,484,1924,1951]" pageId="47" pageNumber="48">Non-type specimens</emphasis>
</paragraph>
<paragraph id="8C48369BFFF3290B38A2F81A75320E5C" blockId="47.[189,1398,1924,2022]" lastBlockId="48.[189,1398,265,327]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
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<collectingCountry id="F4E0760BFFF3291438A2F81A714B08D8" box="[189,280,1960,1986]" name="China" pageId="47" pageNumber="48">CHINA</collectingCountry>
<emphasis id="BE83EA89FFF329143923F81A725808DB" bold="true" box="[316,523,1959,1986]" pageId="47" pageNumber="48">
<collectingRegion id="4E33F879FFF329143923F81A725808DB" box="[316,523,1959,1986]" country="China" name="Hunan" pageId="47" pageNumber="48">Hunan Province</collectingRegion>
</emphasis>
• 10 ♁♁,
<specimenCount id="9AF1FD12FFF329143A8DF81A728408D8" box="[658,727,1960,1986]" pageId="47" pageNumber="48" type="female">7 ♀♀</specimenCount>
(apterous);
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,
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;
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,
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;
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<quantity id="4B0F9B7EFFF329143A29F87972D708FF" box="[566,644,1995,2022]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="48" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
;
<date id="F849105BFFF329143AC8F879732108FF" box="[727,882,1995,2021]" pageId="47" pageNumber="48" value="2016-07-27">
<collectingDate id="E80DE9B3FFF329143AC8F879732108FF" box="[727,882,1995,2021]" pageId="47" pageNumber="48" value="2016-07-27">27 Jul. 2016</collectingDate>
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;
<collectorName id="2102534DFFF329143B9FF879744708FF" box="[896,1044,1995,2021]" pageId="47" pageNumber="48">Yan-chen Li</collectorName>
and
<collectorName id="2102534DFFF329143C4CF879756A08FF" box="[1107,1337,1995,2021]" pageId="47" pageNumber="48">Chen-guang Zheng</collectorName>
leg.;
<collectionCode id="EAE6AE5EFFEC290B38A2FEBB71480E39" box="[189,283,265,291]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="48" pageNumber="49">NKUM</collectionCode>
</materialsCitation>
.
<materialsCitation id="3C9F3CC6FFEC290B3924FEBB70AD0E5C" ID-GBIF-Occurrence="4429869392" collectingDate="1935-07-16" collectionCode="IZAS" collectorName="O. Piel" county="Gu-ling County" location="Lu-shan City" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Jiangxi">
<emphasis id="BE83EA89FFEC290B3924FEBB72580E39" bold="true" box="[315,523,265,291]" pageId="48" pageNumber="49">
<collectingRegion id="4E33F879FFEC290B3924FEBB72580E39" box="[315,523,265,291]" country="China" name="Jiangxi" pageId="48" pageNumber="49">Jiangxi Province</collectingRegion>
</emphasis>
<specimenCount id="9AF1FD12FFEC290B3A3BFEBB72030E39" box="[548,592,265,291]" pageId="48" pageNumber="49" type="female">1 ♀</specimenCount>
(apterous);
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,
<collectingCounty id="65294E17FFEC290B3B9BFEBB74130E39" box="[900,1088,265,291]" pageId="48" pageNumber="49">Gu-ling County</collectingCounty>
;
<date id="F849105BFFEC290B3C53FEBB748C0E3E" box="[1100,1247,265,292]" pageId="48" pageNumber="49" value="1935-07-16">
<collectingDate id="E80DE9B3FFEC290B3C53FEBB748C0E3E" box="[1100,1247,265,292]" pageId="48" pageNumber="49" value="1935-07-16">16 Jul. 1935</collectingDate>
</date>
;
<collectorName id="2102534DFFEC290B3CF4FEBB756F0E39" box="[1259,1340,265,291]" pageId="48" pageNumber="49">O. Piel</collectorName>
leg.;
<collectionCode id="EAE6AE5EFFEC290B38A2FE9E70AD0E5C" box="[189,254,300,326]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="48" pageNumber="49">IZAS</collectionCode>
</materialsCitation>
<materialsCitation id="3C9F3CC6FFEC290B3908FE9E750D0E5C" ID-GBIF-Occurrence="4429869354" box="[279,1374,298,327]" collectingDate="1935-07-16" collectionCode="IZAS" collectorName="O. Piel" county="Gu-ling County" location="Lu-shan City" pageId="48" pageNumber="49" specimenCount="1" stateProvince="Jiangxi">
1 ♁ (dealated macropterous);
<location id="89286040FFEC290B3A69FE9E73410E5C" LSID="urn:lsid:plazi:treatment:045E878DFFF329093A06FBC974DF0DA2:89286040FFEC290B3A69FE9E73410E5C" box="[630,786,300,326]" county="Gu-ling County" name="Lu-shan City" pageId="48" pageNumber="49" stateProvince="Jiangxi">Lu-shan City</location>
,
<collectingCounty id="65294E17FFEC290B3B02FE9E738A0E5C" box="[797,985,300,326]" pageId="48" pageNumber="49">Gu-ling County</collectingCounty>
;
<date id="F849105BFFEC290B3BFAFE9E742A0E5D" box="[997,1145,300,327]" pageId="48" pageNumber="49" value="1935-07-16">
<collectingDate id="E80DE9B3FFEC290B3BFAFE9E742A0E5D" box="[997,1145,300,327]" pageId="48" pageNumber="49" value="1935-07-16">16 Jul. 1935</collectingDate>
</date>
;
<collectorName id="2102534DFFEC290B3C9AFE9E74840E5C" box="[1157,1239,300,326]" pageId="48" pageNumber="49">O. Piel</collectorName>
leg.;
<collectionCode id="EAE6AE5EFFEC290B3D06FE9E750D0E5C" box="[1305,1374,300,326]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="48" pageNumber="49">IZAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8C48369BFFEC290B38A2FEDF71AE0E9D" blockId="48.[189,509,365,391]" box="[189,509,365,391]" pageId="48" pageNumber="49">
<heading id="D70081F7FFEC290B38A2FEDF71AE0E9D" bold="true" box="[189,509,365,391]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="BE83EA89FFEC290B38A2FEDF71AE0E9D" bold="true" box="[189,509,365,391]" pageId="48" pageNumber="49">Supplemental description</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFEC290B38F5FE2E74B008D2" pageId="48" pageNumber="49" type="description">
<paragraph id="8C48369BFFEC290B38F5FE2E71EB0EAC" blockId="48.[185,1399,412,685]" box="[234,440,412,438]" pageId="48" pageNumber="49">
<emphasis id="BE83EA89FFEC290B38F5FE2E71EB0EAC" bold="true" box="[234,440,412,438]" pageId="48" pageNumber="49">Apterous female</emphasis>
</paragraph>
<paragraph id="8C48369BFFEC290B38A2FE7271AC0DB7" blockId="48.[185,1399,412,685]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2FE72713D0EC2" baselines="467,467" box="[189,366,448,474]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="48" pageNumber="49">MEASUREMENTS</smallCapsWord>
. Body length 11.6212.21, width 2.603.08, head width 1.67, interocular width 0.81, eye length (dorsal view) 0.66; relative lengths of antennal segments IIV: 5.11: 1.24: 1.67: 1.10; pronotum: length 0.91, width 1.78; mesonotum: length 2.94, width 3.34; metanotum: length 0.81, width 3.08; abdomen length (ventral view) 5.68; abdominal sternum VII: length 1.95, width 1.53; abdominal mediotergite I: length 0.52, width 1.41; relative lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 5.28:4.51: 3.05: 1.10, middle leg: 13.71:7.95: 3.68: 0.51, hind leg: 14.10: 6.73: 0.17: 0.21.
</paragraph>
<paragraph id="8C48369BFFEC290B38A6FD6675220C61" blockId="48.[185,1399,724,891]" pageId="48" pageNumber="49">
C OLORATION. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum and metanotum mainly black with a median brownish-yellow stripe (
<figureCitation id="14CC2A1EFFEC290B38DAFCA8717B0C2E" box="[197,296,794,820]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="48" pageNumber="49">Figs 3L</figureCitation>
,
<figureCitation id="14CC2A1EFFEC290B3927FCA871230C2E" box="[312,368,794,820]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">29A</figureCitation>
). Yellowish subtriangular marking on metanotum similar with the one on pronotum. Connexivum dorsally brownish-yellow.Abdominal tergum I with a median yellow marking.Mesosternum chiefly blackish with a median subtriangular yellowish spot (
<figureCitation id="14CC2A1EFFEC290B3B9BFCD373BC0C61" box="[900,1007,865,891]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29B</figureCitation>
). Abdominal venter light yellow.
</paragraph>
<paragraph id="8C48369BFFEC290B38A2FC1074AF0BA9" blockId="48.[186,1399,930,1203]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2FC10717F0CA1" baselines="949,950" box="[189,300,930,956]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="48" pageNumber="49">ABDOMEN</smallCapsWord>
. Abdomen relatively long, sightly curved dorsad towards end (
<figureCitation id="14CC2A1EFFEC290B3C32FC1074CF0CA6" box="[1069,1180,930,956]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29C</figureCitation>
). Connexiva erect on abdominal segments IVI, subparallel to each other above mediotergites, reflexed over terminal mediotergites, nearly meeting each other above mediotergite VII (
<figureCitation id="14CC2A1EFFEC290B3BE8FC5B74380B19" box="[1015,1131,1001,1027]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29D</figureCitation>
). Posterior margin of abdominal mediotergite VII with a short median process. Abdominal segment VII elongate, moderately curved dorsad towards end or nearly straight, about as long as two preceding abdominal segments together (
<figureCitation id="14CC2A1EFFEC290B38DDFBE071770B77" box="[194,292,1106,1133]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="48" pageNumber="49">Figs 5K</figureCitation>
,
<figureCitation id="14CC2A1EFFEC290B392FFBE071350B76" box="[304,358,1106,1132]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">29C</figureCitation>
). Posterior margin of abdominal segment VII with four distinct processes, including two lobe-like processes on each connexival corner (
<figureCitation id="14CC2A1EFFEC290B3AFEFBC773380B8A" box="[737,875,1141,1168]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29DF</figureCitation>
), one broader, posteriorly-directed outer lobe and the other narrower pointed inner lobe, ventrally with a small, median process (
<figureCitation id="14CC2A1EFFEC290B3C9AFB2B74BC0BA9" box="[1157,1263,1177,1203]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29E</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFEC290B38F5FB6871F20BEE" blockId="48.[185,1399,1242,1515]" box="[234,417,1242,1268]" pageId="48" pageNumber="49">
<emphasis id="BE83EA89FFEC290B38F5FB6871F20BEE" bold="true" box="[234,417,1242,1268]" pageId="48" pageNumber="49">Apterous male</emphasis>
</paragraph>
<paragraph id="8C48369BFFEC290B38A2FB4C71AC0AF1" blockId="48.[185,1399,1242,1515]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2FB4C713D0A0C" baselines="1297,1297" box="[189,366,1278,1304]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="48" pageNumber="49">MEASUREMENTS</smallCapsWord>
. Body length 8.028.41, width 2.002.60, head width 1.54, interocular width 0.83, eye length (dorsal view) 0.60; relative lengths of antennal segments IIV: 4.12: 1.02: 1.35: 0.94; pronotum: length 0.75, width 1.66; mesonotum: length 2.33, width 2.41; metanotum: length 0.61, width 2.06; abdomen length (ventral view) 3.08; abdominal sternum VII: length 0.59, width 1.22; abdominal mediotergite I: length 0.40, width 0.87; relative lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 4.15: 3.41: 1.38: 0.70, middle leg: 11.32: 6.44: 2.63: 0.43, hind leg: 10.71: 4.08: 0.14: 0.20.
</paragraph>
<paragraph id="8C48369BFFEC290B38A2F9A074EB09C6" blockId="48.[189,1399,1554,1756]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2F9A0711B0930" baselines="1573,1573" box="[189,328,1554,1580]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="48" pageNumber="49">COLORATION</smallCapsWord>
. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum with two color forms: one without (
<figureCitation id="14CC2A1EFFEC290B3A6DF9EA72B30968" box="[626,736,1624,1650]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29G</figureCitation>
) and the other with (
<figureCitation id="14CC2A1EFFEC290B3BFCF9EA746E0968" box="[995,1085,1624,1650]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="48" pageNumber="49">Figs 4I</figureCitation>
,
<figureCitation id="14CC2A1EFFEC290B3C53F9EA74D30968" box="[1100,1152,1624,1650]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">29H</figureCitation>
) yellow marking on metanotum and abdominal tergum I. Individuals of the
<specimenCount id="9AF1FD12FFEC290B3B6CF9CE73B8098C" box="[883,1003,1660,1686]" pageId="48" pageNumber="49" type="generic" typeStatus="types">two types</specimenCount>
were found mixed in the same population. Connexivum dorsally blackish. Mesosternum chiefly blackish with a median subtriangular yellowish spot, narrower than that of female (
<figureCitation id="14CC2A1EFFEC290B3ACBF970736509C6" box="[724,822,1730,1756]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29I</figureCitation>
). Abdominal venter light yellow.
</paragraph>
<paragraph id="8C48369BFFEC290B38A2F8B67139085B" blockId="48.[187,1398,1795,1857]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2F8B670B80806" baselines="1815,1815" box="[189,235,1796,1822]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="48" pageNumber="49">LEG</smallCapsWord>
. Middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
<figureCitation id="14CC2A1EFFEC290B38E6F894710E085B" box="[249,349,1830,1857]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">Fig. 29J</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFEC290B38A2F8DA74B008D2" blockId="48.[189,1399,1896,1993]" pageId="48" pageNumber="49">
<smallCapsWord id="8AAEA047FFEC290B38A2F8DA7162089B" baselines="1915,1916" box="[189,305,1896,1922]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="48" pageNumber="49">GENITALIA</smallCapsWord>
. Abdominal segment VIII ventro-laterally impressed. Pygophore large, ovate. Proctiger relatively large, with angular lobes laterally (
<figureCitation id="14CC2A1EFFEC290B3AC5F839736F08BC" box="[730,828,1931,1958]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="48" pageNumber="49">Figs 7B</figureCitation>
,
<figureCitation id="14CC2A1EFFEC290B3B54F83973D108BF" box="[843,898,1931,1957]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">29K</figureCitation>
). Paramere relatively slender and sinuate, middle part thickened, distal part tapering towards narrowly rounded apex (
<figureCitation id="14CC2A1EFFEC290B3C2AF81C74C508D2" box="[1077,1174,1966,1992]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="48" pageNumber="49">Figs 8H</figureCitation>
,
<figureCitation id="14CC2A1EFFEC290B3CBCF81C748408D2" box="[1187,1239,1966,1992]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="48" pageNumber="49">29L</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D8886613FFED290A38A2F8AF755908F2" ID-DOI="http://doi.org/10.5281/zenodo.8385698" ID-Zenodo-Dep="8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1821,1847]" targetBox="[265,1312,272,1759]" targetPageId="49">
<paragraph id="8C48369BFFED290A38A2F8AF755908F2" blockId="49.[187,1399,1821,2024]" pageId="49" pageNumber="50">
<emphasis id="BE83EA89FFED290A38A2F8AF714B082D" bold="true" box="[189,280,1821,1847]" pageId="49" pageNumber="50">Fig. 29.</emphasis>
Morphological features of
<taxonomicName id="4BF74D18FFED290A3A52F8AC74130822" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[589,1088,1821,1848]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFED290A3A52F8AC730B0822" box="[589,856,1822,1848]" italics="true" pageId="49" pageNumber="50">Rhyacobates lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFED290A3B7AF8AC746B0822" author="Hungerford" box="[869,1080,1821,1848]" firstAuthor="Hungerford" pageId="49" pageNumber="50" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFED290A3C50F8AF7437082D" bold="true" box="[1103,1124,1821,1847]" pageId="49" pageNumber="50">A</emphasis>
. Body of female, dorsal view.
<emphasis id="BE83EA89FFED290A391CF8F371440841" bold="true" box="[259,279,1857,1883]" pageId="49" pageNumber="50">B</emphasis>
. Body of female, excluding head and prothorax, ventral view.
<emphasis id="BE83EA89FFED290A3BE3F8F374420841" bold="true" box="[1020,1041,1857,1883]" pageId="49" pageNumber="50">C</emphasis>
. Body of female, lateral view.
<emphasis id="BE83EA89FFED290A38A1F8D670800864" bold="true" box="[190,211,1892,1918]" pageId="49" pageNumber="50">D</emphasis>
. Abdominal end of female, dorsal view.
<emphasis id="BE83EA89FFED290A3AB4F8D672EC0864" bold="true" box="[683,703,1892,1918]" pageId="49" pageNumber="50">E</emphasis>
. Abdominal end of female, ventral view.
<emphasis id="BE83EA89FFED290A3CBDF8D674E70864" bold="true" box="[1186,1204,1892,1918]" pageId="49" pageNumber="50">F</emphasis>
. Abdominal end of female, lateral view.
<emphasis id="BE83EA89FFED290A39D4F835725B08BB" bold="true" box="[459,520,1927,1953]" pageId="49" pageNumber="50">GH</emphasis>
. Body of male, dorsal view.
<emphasis id="BE83EA89FFED290A3B4EF835730F08BB" bold="true" box="[849,860,1927,1953]" pageId="49" pageNumber="50">I</emphasis>
. Body of male, excluding head and prothorax, ventral view.
<emphasis id="BE83EA89FFED290A3942F819713808DF" bold="true" box="[349,363,1963,1989]" pageId="49" pageNumber="50">J</emphasis>
. Basal part of right middle leg of male, ventral view.
<emphasis id="BE83EA89FFED290A3BF8F81973AE08DF" bold="true" box="[999,1021,1963,1989]" pageId="49" pageNumber="50">K</emphasis>
. Proctiger of male, dorsal view.
<emphasis id="BE83EA89FFED290A38A1F87C708208F2" bold="true" box="[190,209,1998,2024]" pageId="49" pageNumber="50">L</emphasis>
. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C4ED6510FFEE290938A2FEBA73F80E48" box="[189,939,264,338]" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8C48369BFFEE290938A2FEBA71050E38" blockId="50.[189,342,264,290]" box="[189,342,264,290]" pageId="50" pageNumber="51">
<heading id="D70081F7FFEE290938A2FEBA71050E38" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="BE83EA89FFEE290938A2FEBA71050E38" bold="true" box="[189,342,264,290]" pageId="50" pageNumber="51">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFEE290938A2FE8573F80E48" blockId="50.[189,939,311,338]" box="[189,939,311,338]" pageId="50" pageNumber="51">
<collectingCountry id="F4E0760BFFEE290938A2FE85715B0E4B" box="[189,264,311,337]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
:
<collectingRegion id="4E33F879FFEE2909390CFE8A71320E48" box="[275,353,312,338]" country="China" name="Hunan" pageId="50" pageNumber="51">Hunan</collectingRegion>
(first record);
<collectingRegion id="4E33F879FFEE29093A14FE8572200E4B" box="[523,627,311,337]" country="China" name="Zhejiang" pageId="50" pageNumber="51">Zhejiang</collectingRegion>
(
<bibRefCitation id="E8664B6AFFEE29093A9DFE8573CC0E48" author="Andersen &amp; Chen" box="[642,927,311,338]" firstAuthor="Andersen" pageId="50" pageNumber="51" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFEE290938A2FECA74DF0DA2" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8C48369BFFEE290938A2FECA71F90E89" blockId="50.[189,426,376,403]" box="[189,426,376,403]" pageId="50" pageNumber="51">
<heading id="D70081F7FFEE290938A2FECA71F90E89" bold="true" box="[189,426,376,403]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="BE83EA89FFEE290938A2FECA71F90E89" bold="true" box="[189,426,376,403]" pageId="50" pageNumber="51">Comparative notes</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFEE290938A2FE1574DF0DA2" blockId="50.[188,1399,423,696]" pageId="50" pageNumber="51">
<taxonomicName id="4BF74D18FFEE290938A2FE15719D0EDB" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[189,462,423,449]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFEE290938A2FE15719D0EDB" box="[189,462,423,449]" italics="true" pageId="50" pageNumber="51">Rhyacobates lundbladi</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4BF74D18FFEE29093ACEFE1A73210EDB" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[721,882,423,449]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFEE29093ACEFE1A73210EDB" box="[721,882,423,449]" italics="true" pageId="50" pageNumber="51">R. svenhedini</emphasis>
</taxonomicName>
due to their large body sizes, the bilobate processes on the connexival corners of abdominal segment VII and the median process on posterior margin of abdominal mediotergite VII (
<figureCitation id="14CC2A1EFFEE29093ABEFE5F73770D12" box="[673,804,493,520]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="50" pageNumber="51">Figs 3L, P</figureCitation>
,
<figureCitation id="14CC2A1EFFEE29093B2BFE5C733A0D12" box="[820,873,494,520]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="50" pageNumber="51">29D</figureCitation>
,
<figureCitation id="14CC2A1EFFEE29093B66FE5C73E30D12" box="[889,944,494,520]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="50" pageNumber="51">34D</figureCitation>
). However,
<taxonomicName id="4BF74D18FFEE29093C5AFE5C74890D1D" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[1093,1242,493,519]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFEE29093C5AFE5C74890D1D" box="[1093,1242,493,519]" italics="true" pageId="50" pageNumber="51">R. lundbladi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4BF74D18FFEE290938A2FDA3710D0D31" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[189,350,529,555]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFEE290938A2FDA3710D0D31" box="[189,350,529,555]" italics="true" pageId="50" pageNumber="51">R. svenhedini</emphasis>
</taxonomicName>
by the combination of following characteristics of the female: (1) in
<taxonomicName id="4BF74D18FFEE29093CB2FDA375130D31" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[1197,1344,529,555]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFEE29093CB2FDA375130D31" box="[1197,1344,529,555]" italics="true" pageId="50" pageNumber="51">R. lundbladi</emphasis>
</taxonomicName>
, the subtriangular yellowish marking on metanotum is wide, with larger apex angle (
<figureCitation id="14CC2A1EFFEE29093C7DFD8674930D54" box="[1122,1216,564,590]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="50" pageNumber="51">Figs 3L</figureCitation>
,
<figureCitation id="14CC2A1EFFEE29093CD3FD8675500D54" box="[1228,1283,564,590]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="50" pageNumber="51">29D</figureCitation>
), whereas in
<taxonomicName id="4BF74D18FFEE290938C3FDEA71290D6B" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[220,378,599,625]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFEE290938C3FDEA71290D6B" box="[220,378,599,625]" italics="true" pageId="50" pageNumber="51">R. svenhedini</emphasis>
</taxonomicName>
, the subtriangular yellowish marking on metanotum is relatively thin, with sharp apex angle (
<figureCitation id="14CC2A1EFFEE29093912FDC871340D8E" box="[269,359,634,660]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="50" pageNumber="51">Figs 3P</figureCitation>
,
<figureCitation id="14CC2A1EFFEE2909396CFDC871F90D8E" box="[371,426,634,660]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="50" pageNumber="51">34D</figureCitation>
); (2) in
<taxonomicName id="4BF74D18FFEE29093A18FDC972CB0D8E" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[519,664,634,660]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFEE29093A18FDC972CB0D8E" box="[519,664,634,660]" italics="true" pageId="50" pageNumber="51">R. lundbladi</emphasis>
</taxonomicName>
the inner processes on the connexivum of abdominal segment VII are elongate (
<figureCitation id="14CC2A1EFFEE29093993FD2F71AA0DA2" box="[396,505,669,696]" captionStart="Fig" captionStartId="49.[189,232,1821,1847]" captionTargetBox="[265,1312,272,1759]" captionTargetId="figure-12@49.[238,1350,265,1776]" captionTargetPageId="49" captionText="Fig. 29.Morphological features of Rhyacobates lundbladi (Hungerford, 1957). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385698" httpUri="https://zenodo.org/record/8385698/files/figure.png" pageId="50" pageNumber="51">Fig. 29D</figureCitation>
), whereas that in
<taxonomicName id="4BF74D18FFEE29093AD9FD2C73370DAD" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[710,868,669,695]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFEE29093AD9FD2C73370DAD" box="[710,868,669,695]" italics="true" pageId="50" pageNumber="51">R. svenhedini</emphasis>
</taxonomicName>
is very small (
<figureCitation id="14CC2A1EFFEE29093C0DFD2F742C0DA2" box="[1042,1151,669,696]" captionStart="Fig" captionStartId="61.[189,232,1788,1814]" captionTargetBox="[312,1309,275,1739]" captionTargetId="figure-11@61.[242,1345,265,1745]" captionTargetPageId="61" captionText="Fig. 34. Morphological features of Rhyacobates svenhedini (Lundblad, 1934). A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, dorsal view. E. Abdominal end of female, ventral view. F. Abdominal end of female, lateral view. GH. Body of male, apterous form (G) and macropterous form (H), dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: AJ = 1 mm; KL = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385708" httpUri="https://zenodo.org/record/8385708/files/figure.png" pageId="50" pageNumber="51">Fig. 34D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>