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<document id="76BA03BB648F6DBABAC331D1DA704280" ID-DOI="10.5281/zenodo.156319" ID-GBIF-Dataset="4f0aa4a5-9be8-4f8d-aba0-88b4940a9ec6" ID-ISSN="1175­5326" ID-Zenodo-Dep="156319" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459605738350" checkinUser="plazi" docAuthor="Edgecombe, Gregory D." docDate="2003" docId="03D5C216FFD8FF9FFEDD07E1B2EEFCC7" docLanguage="en" docName="zt00193.pdf" docOrigin="Zootaxa 193" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Paralamyctes wellingtonensis Edgecombe, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="FFECBA6EFFD9FF95FFD5050DB03AFFA1" masterDocTitle="A new species of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae) from southern Chile" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1698202849325" updateUser="plazi">
<mods:mods id="B1C9C61769078A51F16F46CCF043764D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3826F7A3F31D63983A04CA40D18B8625">
<mods:title id="74B564ED6D03DA8EE62CA4C64E234BE3">A new species of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae) from southern Chile</mods:title>
</mods:titleInfo>
<mods:name id="654C650D64FB95095F23BEDBD6A1D0E1" type="personal">
<mods:role id="87D9F38DE4E99317A1231D3537B4D3B3">
<mods:roleTerm id="6295EB30BDD7A32D3732BA4904C51A7C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="611D5F5D85D20458F4ABE606ACDB7126">Edgecombe, Gregory D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="61843A48A9C987BB1F5160DECC1A18FA">text</mods:typeOfResource>
<mods:relatedItem id="D5BD362AC06A8D1299D71BAC8B56DD63" type="host">
<mods:titleInfo id="CEF900113E04480FE428820DC694F7CD">
<mods:title id="A1CDD32AC501D6B873D8793342756AF7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="134323849180D325FC81C95B6FE2842A">
<mods:date id="2820FFB4EA0C3540BD027C551C7ABF1F">2003</mods:date>
<mods:detail id="C10EA8D512988B72CBE4D7DBF5905F8D" type="volume">
<mods:number id="277380C7FA1ECB4DAA51FE9AF2BFB1B4">193</mods:number>
</mods:detail>
<mods:extent id="9019D6BABE9D3DA61A5AEF2C9462094E" unit="page">
<mods:start id="61F94A4F2E5053BC52958C73617D44C0">1</mods:start>
<mods:end id="C0CC75F3A05685492E3596F937D6D072">12</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7434512EDA25B1A1E6280361D057E4B2">journal article</mods:classification>
<mods:identifier id="C59B44153A81E04834F2E5FE50EDFE86" type="DOI">10.5281/zenodo.156319</mods:identifier>
<mods:identifier id="5823350E0DDAA7998436401BE48F623A" type="GBIF-Dataset">4f0aa4a5-9be8-4f8d-aba0-88b4940a9ec6</mods:identifier>
<mods:identifier id="4F3EF29DC33AF25FE83257931913CCB5" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="51C93AEFB42F578D3AA2E5BA31F556C5" type="Zenodo-Dep">156319</mods:identifier>
</mods:mods>
<treatment id="03D5C216FFD8FF9FFEDD07E1B2EEFCC7" ID-DOI="http://doi.org/10.5281/zenodo.6277022" ID-GBIF-Taxon="119331443" ID-Zenodo-Dep="6277022" LSID="urn:lsid:plazi:treatment:03D5C216FFD8FF9FFEDD07E1B2EEFCC7" httpUri="http://treatment.plazi.org/id/03D5C216FFD8FF9FFEDD07E1B2EEFCC7" lastPageId="10" lastPageNumber="11" pageId="1" pageNumber="2">
<subSubSection id="C366208BFFD8FF94FEDD07E1B145FC88" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BC37300FFD8FF94FEDD07E1B28DFCA4" blockId="1.[264,695,747,809]" box="[264,695,747,774]" pageId="1" pageNumber="2">
<heading id="D08BC46CFFD8FF94FEDD07E1B28DFCA4" bold="true" box="[264,695,747,774]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B908AF12FFD8FF94FEDD07E1B28DFCA4" bold="true" box="[264,695,747,774]" pageId="1" pageNumber="2">
<taxonomicName id="4C7C0883FFD8FF94FEDD07E1B257FCA7" ID-CoL="4CXVC" box="[264,621,748,774]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFD8FF94FEDD07E1B257FCA7" bold="true" box="[264,621,748,774]" italics="true" pageId="1" pageNumber="2">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFD8FF94FDA007E6B28DFCA4" box="[629,695,747,773]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC37300FFD8FF94FEDD0602B145FC88" blockId="1.[264,695,747,809]" box="[264,383,783,809]" pageId="1" pageNumber="2">
<figureCitation id="13476F85FFD8FF94FEDD0602B145FC88" box="[264,383,783,809]" captionStart-0="FIGURES 1 ­ 3" captionStart-1="FIGURES 4 ­ 7" captionStart-2="FIGURES 8 ­ 15" captionStart-3="FIGURES 16 ­ 23" captionStart-4="FIGURES 24 ­ 31" captionStartId-0="2.[264,383,1719,1743]" captionStartId-1="3.[264,383,1466,1490]" captionStartId-2="3.[264,383,1655,1679]" captionStartId-3="5.[264,383,1684,1708]" captionStartId-4="7.[264,383,1684,1708]" captionTargetBox-0="[278,1308,332,1688]" captionTargetBox-1="[292,1289,354,1441]" captionTargetId-0="figure@2.[273,1314,329,1693]" captionTargetId-1="figure@3.[264,1323,329,1455]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." captionText-1="FIGURES 4 ­ 7. Paralamyctes wellingtonensis n. sp. MCZ 27836, male. 4, leg 12; 5, leg 13; 6, leg 14 (tibial projection atypically non­spinose); 7, leg 15. Scale 500 µm." captionText-2="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." captionText-3="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." captionText-4="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." httpUri-0="https://zenodo.org/record/156320/files/figure.png" httpUri-1="https://zenodo.org/record/156321/files/figure.png" pageId="1" pageNumber="2">Figs. 1­31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD8FF94FEDD0652B18AFC37" pageId="1" pageNumber="2" type="description">
<paragraph id="8BC37300FFD8FF94FEDD0652B18AFC37" blockId="1.[264,1321,863,918]" pageId="1" pageNumber="2">
<taxonomicName id="4C7C0883FFD8FF94FEDD0652B1A1FCD7" box="[264,411,863,886]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B908AF12FFD8FF94FEDD0652B1A1FCD7" box="[264,411,863,886]" italics="true" pageId="1" pageNumber="2">Paralamyctes</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C7C0883FFD8FF94FE1C0652B279FCD7" box="[457,579,863,886]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B908AF12FFD8FF94FE1C0652B279FCD7" box="[457,579,863,886]" italics="true" pageId="1" pageNumber="2">P. chilensis</emphasis>
</taxonomicName>
(Gervais in
<bibRefCitation id="EFED0EF1FFD8FF94FD050652B42BFCD7" author="Walckenaer" box="[720,1041,863,886]" pageId="1" pageNumber="12" refString="Walckenaer, M. LeB. &amp; Gervais, M. (1847) Histoire naturelle des Insectes. Apteres, 4. Libraire Encyclopedique de Roret, Paris." type="book chapter" year="1847">Walckenaer &amp; Gervais, 1847</bibRefCitation>
): Edgecombe et al. 2003, figs. 3A­C.
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD8FF94FEDD06CAB464FBF8" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BC37300FFD8FF94FEDD06CAB464FBF8" blockId="1.[264,1325,967,1832]" pageId="1" pageNumber="2">
<emphasis id="B908AF12FFD8FF94FEDD06CAB145FC41" box="[264,383,967,992]" italics="true" pageId="1" pageNumber="2">Diagnosis</emphasis>
:
<taxonomicName id="4C7C0883FFD8FF94FE4506CAB20BFC41" box="[400,561,967,992]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B908AF12FFD8FF94FE4506CAB20BFC41" box="[400,561,967,992]" italics="true" pageId="1" pageNumber="2">Paralamyctes</emphasis>
</taxonomicName>
with 17­20 (usually 19) antennal articles. Female gonopod with long, slender process on first article, bearing two relatively stout, blunt spurs. Females with 2,2,2,2/2,2,2,2 to 2,3,3,3/2,3,3,3 coxal pores, inner pore tiny when three present; males mostly with 2,2,2,2/2,2,2,2 coxal pores, maximum 2,3,3,2/2,3,3,2.
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD8FF94FEED016AB222FA38" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BC37300FFD8FF94FEED016AB415FB08" blockId="1.[264,1325,967,1832]" pageId="1" pageNumber="2">
<typeStatus id="54C7CDA2FFD8FF94FEED016AB199FB21" box="[312,419,1127,1152]" pageId="1" pageNumber="2" type="holotype">
<emphasis id="B908AF12FFD8FF94FEED016AB199FB21" box="[312,419,1127,1152]" italics="true" pageId="1" pageNumber="2">Holotype</emphasis>
</typeStatus>
: Male
<collectionCode id="ED6DEBC5FFD8FF94FE2B016AB204FB20" LSID="urn:lsid:biocol.org:col:33791" box="[510,574,1127,1153]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
52636 (
<figureCitation id="13476F85FFD8FF94FD73016AB2D4FB20" box="[678,750,1127,1153]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
), from Puerto Edén,
<collectingCountry id="F36B3390FFD8FF94FC23016AB41AFB20" box="[1014,1056,1127,1153]" name="Iceland" pageId="1" pageNumber="2">Isla</collectingCountry>
Wellington,
<collectingCountry id="F36B3390FFD8FF94FB15016AB539FB20" box="[1216,1283,1127,1153]" name="Chile" pageId="1" pageNumber="2">Chile</collectingCountry>
, ca 49ºS [
<geoCoordinate id="EE4815C7FFD8FF94FE980182B196FB08" box="[333,428,1167,1193]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-49.133335">49°08'S</geoCoordinate>
<geoCoordinate id="EE4815C7FFD8FF94FE660182B21AFB08" box="[435,544,1167,1193]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="-74.45">74°27'W</geoCoordinate>
], leg. P.J. Darlington,
<date id="FFC255C0FFD8FF94FCF70182B411FB08" box="[802,1067,1167,1193]" pageId="1" pageNumber="2" value="1962-12-10" valueMax="1962-12-13" valueMin="1962-12-10">10­13 December 1962</date>
.
</paragraph>
<paragraph id="8BC37300FFD8FF94FEED01BAB2BAFA80" blockId="1.[264,1325,967,1832]" pageId="1" pageNumber="2">
<typeStatus id="54C7CDA2FFD8FF94FEED01BAB195FB71" box="[312,431,1207,1232]" pageId="1" pageNumber="2" type="paratype">
<emphasis id="B908AF12FFD8FF94FEED01BAB195FB71" box="[312,431,1207,1232]" italics="true" pageId="1" pageNumber="2">Paratypes</emphasis>
</typeStatus>
: All from
<typeStatus id="54C7CDA2FFD8FF94FDED01BAB250FB70" box="[568,618,1207,1233]" pageId="1" pageNumber="2">type</typeStatus>
locality, leg. P.J. Darlington,
<date id="FFC255C0FFD8FF94FC0C01BAB4E6FB70" box="[985,1244,1207,1233]" pageId="1" pageNumber="2" value="1962-12-07" valueMax="1962-12-15" valueMin="1962-12-07">7­15 December 1962</date>
.
<collectionCode id="ED6DEBC5FFD8FF94FB3901BAB516FB70" LSID="urn:lsid:biocol.org:col:33791" box="[1260,1324,1207,1233]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
27838,
<specimenCount id="9D7AB889FFD8FF94FEB101D2B1E8FB58" box="[356,466,1247,1273]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
,
<specimenCount id="9D7AB889FFD8FF94FE3401D2B279FB58" box="[481,579,1247,1273]" pageId="1" pageNumber="2" type="male">3 males</specimenCount>
, from
<typeStatus id="54C7CDA2FFD8FF94FD4D01D2B2F0FB58" box="[664,714,1247,1273]" pageId="1" pageNumber="2">type</typeStatus>
collection;
<collectionCode id="ED6DEBC5FFD8FF94FC8801D2B3A7FB58" LSID="urn:lsid:biocol.org:col:33791" box="[861,925,1247,1273]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
27836,
<specimenCount id="9D7AB889FFD8FF94FBD101D2B45DFB58" box="[1028,1127,1247,1273]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
;
<collectionCode id="ED6DEBC5FFD8FF94FBA201D2B48DFB58" LSID="urn:lsid:biocol.org:col:33791" box="[1143,1207,1247,1273]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
47468,
<specimenCount id="9D7AB889FFD8FF94FACB01D2B166FA80" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
;
<collectionCode id="ED6DEBC5FFD8FF94FEB2000AB19DFA80" LSID="urn:lsid:biocol.org:col:33791" box="[359,423,1287,1313]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
47470,
<specimenCount id="9D7AB889FFD8FF94FDD2000AB246FA80" box="[519,636,1287,1313]" pageId="1" pageNumber="2" type="female">4 females</specimenCount>
.
</paragraph>
<paragraph id="8BC37300FFD8FF94FEED0022B222FA38" blockId="1.[264,1325,967,1832]" pageId="1" pageNumber="2">
<emphasis id="B908AF12FFD8FF94FEED0022B1D2FAE9" box="[312,488,1327,1352]" italics="true" pageId="1" pageNumber="2">Other material</emphasis>
:
<collectionCode id="ED6DEBC5FFD8FF94FE2F0022B200FAE8" LSID="urn:lsid:biocol.org:col:33791" box="[506,570,1327,1353]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="2">MCZ</collectionCode>
<collectionCode id="ED6DEBC5FFD8FF94FD970022B2BFFAE8" LSID="urn:lsid:biocol.org:col:15536" box="[578,645,1327,1353]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="1" pageNumber="2">DNA</collectionCode>
<date id="FFC255C0FFD8FF94FD500022B2E7FAE8" box="[645,733,1327,1353]" pageId="1" pageNumber="2" value="08-10-04">100408</date>
,
<specimenCount id="9D7AB889FFD8FF94FD3E0022B2C3FAE8" box="[747,761,1327,1353]" pageId="1" pageNumber="2" type="male">2</specimenCount>
males, near Refugio Chileno, Parque Nacional Torres del Paine, Provincia Esperanza,
<geoCoordinate id="EE4815C7FFD8FF94FD02005AB360FAD0" box="[727,858,1367,1393]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="15" value="-50.945835">50º56'45&quot;S</geoCoordinate>
<geoCoordinate id="EE4815C7FFD8FF94FCB7005AB3D2FAD0" box="[866,1000,1367,1393]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="15" value="72.916664">72º55'00&quot;E</geoCoordinate>
, leg. J. Miller &amp; I. Agnarsson,
<date id="FFC255C0FFD8FF94FE950072B22EFA38" box="[320,532,1407,1433]" pageId="1" pageNumber="2" value="2000-12-08">8 December 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD8FF94FEED00AAB2FFFA60" box="[312,709,1447,1473]" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8BC37300FFD8FF94FEED00AAB2FFFA60" blockId="1.[264,1325,967,1832]" box="[312,709,1447,1473]" pageId="1" pageNumber="2">
<emphasis id="B908AF12FFD8FF94FEED00AAB18FFA61" box="[312,437,1447,1472]" italics="true" pageId="1" pageNumber="2">Etymology</emphasis>
: After
<collectingCountry id="F36B3390FFD8FF94FDDF00AAB20EFA60" box="[522,564,1447,1473]" name="Iceland" pageId="1" pageNumber="2">Isla</collectingCountry>
Wellington.
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD8FF9CFEED00C2B34FFACF" lastPageId="9" lastPageNumber="10" pageId="1" pageNumber="2" type="description">
<paragraph id="8BC37300FFD8FF94FEED00C2B231F889" blockId="1.[264,1325,967,1832]" pageId="1" pageNumber="2">
<emphasis id="B908AF12FFD8FF94FEED00C2B1F8FA49" box="[312,450,1487,1512]" italics="true" pageId="1" pageNumber="2">Description</emphasis>
: Length (anterior margin of head shield to end of telson) up to 16.6 mm; length of head shield up to 1.6 mm. Colour (based on specimens in absolute ethanol): head shield and tergites pale yellow to pale orange with purple mottling; dark purple patch around ocellus and on cephalic pleurite beneath ocellus; antenna yellow­brown to moderate orange; maxillipedes yellow to pale orange, deeper orange on distal part of telopodite; tergites with purple longitudinal median band; anterior sternites pale yellow­brown, posterior sternites variably orange; legs yellow with purple mottling on femur and tibia, also on prefemur of legs 14 and 15; tarsus pale orange, deeper on posterior legs; genital sternite and gonopods orange.
</paragraph>
<caption id="DF032388FFDBFF97FEDD03BAB4A8F8B5" httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="2" pageNumber="3" targetBox="[278,1308,332,1688]" targetPageId="2">
<paragraph id="8BC37300FFDBFF97FEDD03BAB4A8F8B5" blockId="2.[264,1323,1719,1812]" pageId="2" pageNumber="3">
<emphasis id="B908AF12FFDBFF97FEDD03BAB190F96E" bold="true" box="[264,426,1719,1743]" pageId="2" pageNumber="3">FIGURES 1­3</emphasis>
.
<taxonomicName id="4C7C0883FFDBFF97FE6F03B5B2C1F96E" box="[442,763,1720,1743]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFDBFF97FE6F03B5B2C1F96E" box="[442,763,1720,1743]" italics="true" pageId="2" pageNumber="3">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFDBFF97FCD603B5B304F96E" box="[771,830,1720,1743]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm.
</paragraph>
</caption>
<caption id="DF032388FFDAFF96FEDD00B7B3D4FA55" httpUri="https://zenodo.org/record/156321/files/figure.png" pageId="3" pageNumber="4" targetBox="[292,1289,354,1441]" targetPageId="3">
<paragraph id="8BC37300FFDAFF96FEDD00B7B3D4FA55" blockId="3.[264,1323,1466,1524]" pageId="3" pageNumber="4">
<emphasis id="B908AF12FFDAFF96FEDD00B7B193FA73" bold="true" box="[264,425,1466,1490]" pageId="3" pageNumber="4">FIGURES 4­7</emphasis>
.
<taxonomicName id="4C7C0883FFDAFF96FE6D00B7B2C2FA70" box="[440,760,1466,1489]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFDAFF96FE6D00B7B2C2FA70" box="[440,760,1466,1489]" italics="true" pageId="3" pageNumber="4">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFDAFF96FD2A00B7B303FA70" box="[767,825,1466,1489]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
MCZ 27836, male. 4, leg 12; 5, leg 13; 6, leg 14 (tibial projection atypically non­spinose); 7, leg 15. Scale 500 µm.
</paragraph>
</caption>
<caption id="DF032388FFDAFF96FEDD037AB296F89D" pageId="3" pageNumber="4">
<paragraph id="8BC37300FFDAFF96FEDD037AB296F89D" blockId="3.[264,1323,1655,1852]" pageId="3" pageNumber="4">
<emphasis id="B908AF12FFDAFF96FEDD037AB18CF92E" bold="true" box="[264,438,1655,1679]" pageId="3" pageNumber="4">FIGURES 8­15</emphasis>
.
<taxonomicName id="4C7C0883FFDAFF96FE160375B338F92E" box="[451,770,1656,1679]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFDAFF96FE160375B338F92E" box="[451,770,1656,1679]" italics="true" pageId="3" pageNumber="4">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFDAFF96FCDD0375B37BF92E" box="[776,833,1656,1679]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9­11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm.
</paragraph>
</caption>
<paragraph id="8BC37300FFDCFF90FEED0411B2A7FD4F" blockId="5.[264,1324,284,1630]" pageId="5" pageNumber="6">
Head shield smooth, 90% width of widest tergites (TT8 and 10). Anterior margin with deep median notch; longitudinal median furrow deeply impressed to transverse suture (
<figureCitation id="13476F85FFDCFF90FEC40461B158FE27" box="[273,354,364,390]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 8</figureCitation>
), variably impressed behind suture, in some specimens extending back to midlength of head shield (
<figureCitation id="13476F85FFDCFF90FDEB0499B2BEFE0F" box="[574,644,404,430]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
). Posterior margin of head shield gently concave. Border slightly wider posterolaterally than posteromedially. Antenna up to 3.9 times length of head shield; 19 articles on both antennae in most specimens, single specimens with 19/17, 18/19 and 19/20 articles (15 articles on one antenna in
<typeStatus id="54C7CDA2FFDCFF90FC530701B3D7FD87" box="[902,1005,524,550]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
possibly teratological:
<figureCitation id="13476F85FFDCFF90FB280701B12FFDEF" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
); antenna usually extends back to anterior part of T5; proximal two articles slightly larger than third and fourth; most articles substantially longer than wide, with many trichoid sensilla (
<figureCitation id="13476F85FFDCFF90FE690789B22FFD3F" box="[444,533,644,670]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 12</figureCitation>
); terminal article nearly twice length of penultimate. Ocellus relatively large, domed. Tömösváry organ small, elliptical, its outer margin just inside lateral edge of cephalic pleurite (
<figureCitation id="13476F85FFDCFF90FDEC07D9B2B5FD4F" box="[569,655,724,750]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BC37300FFDCFF90FEED07F1B4ADFC2F" blockId="5.[264,1324,284,1630]" pageId="5" pageNumber="6">
Clypeus bearing transverse band of four setae posteromedially, just in front of labrum. Labrum with transverse seta on sidepiece pit that projects towards midpiece (
<figureCitation id="13476F85FFDCFF90FB5E0629B4E9FC9F" box="[1163,1235,804,830]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 9</figureCitation>
); labral margin with weak break in curvature where fringe of branching bristles projects; bristles with many slender, spine­like marginal branches along their lengths (
<figureCitation id="13476F85FFDCFF90FBE70679B4B0FC2F" box="[1074,1162,884,910]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BC37300FFDCFF90FEED0691B53DFB07" blockId="5.[264,1324,284,1630]" pageId="5" pageNumber="6">
Maxillipede: Dental margin with 4+5, 5+4, 5+5, 5+6 or 6+5 pointed teeth; inner tooth smaller when more than four teeth present; each half of dental margin gently (
<figureCitation id="13476F85FFDCFF90FB7906C9B538FC7F" box="[1196,1282,964,990]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 15</figureCitation>
) to moderately convex with rounded shoulder laterally; median notch shallow. Setation on coxosternite slightly concentrated anteriorly, with numerous mostly short setae, though setae are scattered over posterior part of coxosternite as well (
<figureCitation id="13476F85FFDCFF90FC020131B417FBF7" box="[983,1069,1084,1110]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 14</figureCitation>
). Pretarsal part of tarsungulum slender, up to 1.4 times length of tarsal part (
<figureCitation id="13476F85FFDCFF90FC4E0169B3C9FBDF" box="[923,1011,1124,1150]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="5" pageNumber="6">Fig. 13</figureCitation>
); setation on inner margin of tarsal part of tarsungulum, tibia and femur of similar density to that on outer margin.
</paragraph>
<paragraph id="8BC37300FFDCFF90FEED01B9B2DEF9FF" blockId="5.[264,1324,284,1630]" pageId="5" pageNumber="6">
Mandible: Four paired teeth (
<figureCitation id="13476F85FFDCFF90FD4301B9B2D5FB6F" box="[662,751,1204,1230]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 16</figureCitation>
). 9­13 aciculae; up to 27 blunt barbs along both margins of all aciculae (
<figureCitation id="13476F85FFDCFF90FDCA01D1B24CFB57" box="[543,630,1244,1270]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 17</figureCitation>
). Fringe of branching bristles skirts aciculae (
<figureCitation id="13476F85FFDCFF90FB5F01D1B4DAFB57" box="[1162,1248,1244,1270]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 19</figureCitation>
), bristles in ventral part of mandible narrow­based, evenly branching to their bases, splaying into two or more thicker, elongate spines distally; bristles gradually shortening dorsally, grading into multiramous scales that form a dense fringe of two largely intermixed scale rows; fringe extending to furry pad (
<figureCitation id="13476F85FFDCFF90FD640071B332FA37" box="[689,776,1404,1430]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 18</figureCitation>
). Accessory denticle field with strong grooved ridge on three dorsal teeth, ridge bearing numerous angular denticles (
<figureCitation id="13476F85FFDCFF90FBAC00A9B4ECFA1F" box="[1145,1238,1444,1470]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 20</figureCitation>
); most accessory denticles triangular, with pointed apices, grading into rod­shaped denticles then small elongate scales near fringe of branching bristles. Furry pad strongly differentiated from accessory denticle field, composed of abundant mostly simple bristles, mostly curled back towards dorsalmost tooth (
<figureCitation id="13476F85FFDCFF90FDAA0349B2ECF9FF" box="[639,726,1604,1630]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="5" pageNumber="6">Fig. 18</figureCitation>
).
</paragraph>
<caption id="DF032388FFDCFF90FEDD0399B428F8C7" pageId="5" pageNumber="6">
<paragraph id="8BC37300FFDCFF90FEDD0399B428F8C7" blockId="5.[264,1323,1684,1894]" pageId="5" pageNumber="6">
<emphasis id="B908AF12FFDCFF90FEDD0399B1F3F90D" bold="true" box="[264,457,1684,1708]" pageId="5" pageNumber="6">FIGURES 16­23</emphasis>
.
<taxonomicName id="4C7C0883FFDCFF90FE090399B31BF90A" box="[476,801,1684,1707]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFDCFF90FE090399B31BF90A" box="[476,801,1684,1707]" italics="true" pageId="5" pageNumber="6">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFDCFF90FCF80399B356F90A" box="[813,876,1684,1707]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
All from Puerto Edén, Isla Wellington except 19­20, from Parque Nacional Torres del Paine. 16­18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19­20, MCZ DNA100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21­23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm.
</paragraph>
</caption>
<paragraph id="8BC37300FFDEFF92FEED0411B207FDEF" blockId="7.[264,1324,284,1470]" pageId="7" pageNumber="8">
First maxilla: Bell­shaped sternite of typical size for genus, anterolateral margins set in arthrodial membrane (
<figureCitation id="13476F85FFDEFF92FDC00449B24AFEFF" box="[533,624,324,350]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="7" pageNumber="8">Fig. 21</figureCitation>
). Coxal process with blunt apex bearing about 13 simple setae (
<figureCitation id="13476F85FFDEFF92FE800461B195FE27" box="[341,431,364,390]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="7" pageNumber="8">Fig. 22</figureCitation>
), this cluster separated from a few setae on medial edge of coxal process. Cluster of six sensilla microtrichoidea between coxal process and telopodite. Distal article of telopodite with two rows of long, plumose setae along inner margin; slender branches developed along distal half of these setae (
<figureCitation id="13476F85FFDEFF92FCD604E9B366FE5F" box="[771,860,484,510]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="7" pageNumber="8">Fig. 23</figureCitation>
); short, simple setae near bases of plumose setae on their ventral side; main, more sclerotised field of distal article of telopodite with a few scattered setae.
</paragraph>
<paragraph id="8BC37300FFDEFF92FEED0751B32CFC9F" blockId="7.[264,1324,284,1470]" pageId="7" pageNumber="8">
Second maxilla: Sternite small, triangular, fused with coxae, sutures distinct. Band of about six short setae along anterior part of coxa. Trochanter­prefemur joint defined by incision on inner margin; tarsus bearing numerous simple setae on outer surface, about 11 plumose setae on inner surface; branches on plumose setae confined to distal half (
<figureCitation id="13476F85FFDEFF92FB2907D9B113FCB7" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="7" pageNumber="8">Fig. 24</figureCitation>
). Claw composed of five digits, the middle the thickest and longest (
<figureCitation id="13476F85FFDEFF92FBBF07F1B4F9FCB7" box="[1130,1219,764,790]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="7" pageNumber="8">Fig. 25</figureCitation>
), pair of slender digits between it and stout outer pair.
</paragraph>
<paragraph id="8BC37300FFDEFF92FEED0641B2DCFABF" blockId="7.[264,1324,284,1470]" pageId="7" pageNumber="8">
Tergites gently wrinkled and turned up laterally. T1 about 86% width of head shield, weakly trapezoidal, with transverse or faintly concave posterior margin; TT3 and 5 with subparallel lateral margins, posterior angles rounded, posterior margins gently concave with most curvature medially (
<figureCitation id="13476F85FFDEFF92FDA506C9B28FFC7F" box="[624,693,964,990]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
); TT6­14 bordered laterally only; posterior margin of T7 moderately, evenly concave, posterior angle slightly projected; posterior margins of TT8, 10 and 12 concave, increasingly so more posteriorly, posterior angles form obtuse, blunt corners; posterior margins of TT9, 11 and 13 with largely transverse median extent or T13 weakly convex; posterior angle of T9 with blunt projection, T11 and, more so, T13, with strong projections but not toothed; posterior margin of T14 gently concave, with sharp posterior angles lacking projection; posterior margin of tergite of intermediate segment concave in both sexes, with rounded posterior angle. Tergites of first genital segment and telson well sclerotised in both sexes.
</paragraph>
<paragraph id="8BC37300FFDEFF92FEED0021B378FA1F" blockId="7.[264,1324,284,1470]" pageId="7" pageNumber="8">Numerous short, fine setae along lateral borders of tergites; several fine setae along posterior margins of all tergites, numerous from T4; surface of tergites scattered with short setae. Sternites with several setae in two ill­defined transverse bands across anterior third; two or three longer setae at anterolateral margin.</paragraph>
<caption id="DF032388FFDEFF92FEDD0399B350F8E1" pageId="7" pageNumber="8">
<paragraph id="8BC37300FFDEFF92FEDD0399B350F8E1" blockId="7.[264,1323,1684,1856]" pageId="7" pageNumber="8">
<emphasis id="B908AF12FFDEFF92FEDD0399B1FFF90D" bold="true" box="[264,453,1684,1708]" pageId="7" pageNumber="8">FIGURES 24­31</emphasis>
.
<taxonomicName id="4C7C0883FFDEFF92FE010399B32EF90A" box="[468,788,1684,1707]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFDEFF92FE010399B32EF90A" box="[468,788,1684,1707]" italics="true" pageId="7" pageNumber="8">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFDEFF92FCC90399B36CF90A" box="[796,854,1684,1707]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
Puerto Edén, Isla Wellington. 24­27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29­31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31).
</paragraph>
</caption>
<paragraph id="8BC37300FFD0FF9CFEED0411B330FD3F" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Articulations between two tarsomeres strong, continuous on dorsal side even on anterior legs, with condyle on legs 13­15. Setae with similar densities and thicknesses on all legs; setae along inner edge of distitarsus sloping distally on legs 1­14. Sharp distal spinose projection on tibiae of legs 1­13 (
<figureCitation id="13476F85FFD0FF9CFD1F0499B303FE0F" box="[714,825,404,430]" captionStart="FIGURES 4 ­ 7" captionStartId="3.[264,383,1466,1490]" captionTargetBox="[292,1289,354,1441]" captionTargetId="figure@3.[264,1323,329,1455]" captionTargetPageId="3" captionText="FIGURES 4 ­ 7. Paralamyctes wellingtonensis n. sp. MCZ 27836, male. 4, leg 12; 5, leg 13; 6, leg 14 (tibial projection atypically non­spinose); 7, leg 15. Scale 500 µm." httpUri="https://zenodo.org/record/156321/files/figure.png" pageId="9" pageNumber="10">Figs. 4, 5</figureCitation>
), spinose projection usually present on leg 14, exceptionally blunt (
<figureCitation id="13476F85FFD0FF9CFDF104B1B256FE77" box="[548,620,444,470]" captionStart="FIGURES 4 ­ 7" captionStartId="3.[264,383,1466,1490]" captionTargetBox="[292,1289,354,1441]" captionTargetId="figure@3.[264,1323,329,1455]" captionTargetPageId="3" captionText="FIGURES 4 ­ 7. Paralamyctes wellingtonensis n. sp. MCZ 27836, male. 4, leg 12; 5, leg 13; 6, leg 14 (tibial projection atypically non­spinose); 7, leg 15. Scale 500 µm." httpUri="https://zenodo.org/record/156321/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Distitarsus 52­54% length of basitarsus on leg 14, 51­57% on leg 15; leg 15 basitarsus about 11 times longer than wide (
<figureCitation id="13476F85FFD0FF9CFC2504E9B400FE5F" box="[1008,1082,484,510]" captionStart="FIGURES 4 ­ 7" captionStartId="3.[264,383,1466,1490]" captionTargetBox="[292,1289,354,1441]" captionTargetId="figure@3.[264,1323,329,1455]" captionTargetPageId="3" captionText="FIGURES 4 ­ 7. Paralamyctes wellingtonensis n. sp. MCZ 27836, male. 4, leg 12; 5, leg 13; 6, leg 14 (tibial projection atypically non­spinose); 7, leg 15. Scale 500 µm." httpUri="https://zenodo.org/record/156321/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
). Posterior pretarsal accessory claw slightly longer than anterior accessory claw, 35­45% length of main claw (
<figureCitation id="13476F85FFD0FF9CFEC40739B150FDEF" box="[273,362,564,590]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 26</figureCitation>
); scutes weakly defined on accessory claws (
<figureCitation id="13476F85FFD0FF9CFC570739B3E1FDEF" box="[898,987,564,590]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 27</figureCitation>
); elongate scutes defined by sutures along length of main pretarsal claw, including its dorsoproximal part (
<figureCitation id="13476F85FFD0FF9CFB170751B524FDD7" box="[1218,1310,604,630]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 26</figureCitation>
), without pores; posteroventral spine lacking.
</paragraph>
<paragraph id="8BC37300FFD0FF9CFEED07A1B156FD4F" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">Coxal pores on legs 12­15, round, numbers as in Diagnosis; pore rows not set in grooves.</paragraph>
<paragraph id="8BC37300FFD0FF9CFEED07F1B428FBF7" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Female (
<figureCitation id="13476F85FFD0FF9CFE4A07F1B1C9FCB7" box="[415,499,764,790]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13476F85FFD0FF9CFE2A07F1B273FCB7" box="[511,585,764,790]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">29­31</figureCitation>
): Sternite of segment 15 transverse or faintly concave posteromedially, numerous setae on posterolateral margin, a few on posterior margin and scattered over surface. Sternite of first genital segment with up to 48 short to moderately long setae scattered over most of surface, with slight to distinct concentration near posterolateral margin. Basal article of gonopod bearing 13­20 setae, with two blunt spurs of about equal size at distal end of slender, elongate process (
<figureCitation id="13476F85FFD0FF9CFCA006C9B3F7FC7F" box="[885,973,964,990]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 31</figureCitation>
); dorsal surface of spurs concave, with several low tubercles (
<figureCitation id="13476F85FFD0FF9CFD4106E1B2D6FBA7" box="[660,748,1004,1030]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 30</figureCitation>
); second article of gonopod with 5­9 setae; third article with two or three setae. Claw undivided, bearing many strong pores on its dorsodistal surface, each pore housing a sensillum coeloconicum (
<figureCitation id="13476F85FFD0FF9CFC780131B43EFBF7" box="[941,1028,1084,1110]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BC37300FFD0FF9CFEED0169B34FFACF" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Male (
<figureCitation id="13476F85FFD0FF9CFE530169B1E0FBDF" box="[390,474,1124,1150]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
,
<figureCitation id="13476F85FFD0FF9CFE3C0169B231FBDF" box="[489,523,1124,1150]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">28</figureCitation>
): Sternite of segment 15 transverse posteromedially; setae sparsely scattered on its anterior half, about 12 setae fringing posterolateral and posteromedian margins. Sternite of first genital segment undivided, posterior margin bulging backwards between gonopods; up to 26 setae scattered over sternite, a few long setae variably present on posterior margin (
<figureCitation id="13476F85FFD0FF9CFDD70009B261FABF" box="[514,603,1284,1310]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 28</figureCitation>
). Basal two articles of gonopod each bearing a single row of three to seven setae, usually four or five on first article; third article with four to seven scattered setae; terminal process tapering, spine­like.
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD0FF9CFEED0071B4E2F8EF" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BC37300FFD0FF9CFEED0071B4E2F8EF" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<emphasis id="B908AF12FFD0FF9CFEED0071B182FA34" box="[312,440,1404,1429]" italics="true" pageId="9" pageNumber="10">Discussion</emphasis>
:
<taxonomicName id="4C7C0883FFD0FF9CFE1D0071B312FA34" box="[456,808,1404,1429]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="wellingtonensis" status="sp. nov.">
<emphasis id="B908AF12FFD0FF9CFE1D0071B312FA34" box="[456,808,1404,1429]" italics="true" pageId="9" pageNumber="10">Paralamyctes wellingtonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B1269FFD0FF9CFCFA0071B355FA37" box="[815,879,1404,1430]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
is most obviously distinguished from
<taxonomicName id="4C7C0883FFD0FF9CFEDD00A9B1B7FA1C" box="[264,397,1444,1469]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B908AF12FFD0FF9CFEDD00A9B1B7FA1C" box="[264,397,1444,1469]" italics="true" pageId="9" pageNumber="10">P. chilensis</emphasis>
</taxonomicName>
(Gervais in
<bibRefCitation id="EFED0EF1FFD0FF9CFDF600A9B346FA1F" author="Walckenaer" box="[547,892,1444,1470]" pageId="9" pageNumber="12" refString="Walckenaer, M. LeB. &amp; Gervais, M. (1847) Histoire naturelle des Insectes. Apteres, 4. Libraire Encyclopedique de Roret, Paris." type="book chapter" year="1847">Walckenaer &amp; Gervais, 1847</bibRefCitation>
) by its more elongated spur­bearing process on the female gonopod (compare
<figureCitation id="13476F85FFD0FF9CFD2F00C1B377FA47" box="[762,845,1484,1510]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
,
<figureCitation id="13476F85FFD0FF9CFC8E00C1B342FA47" box="[859,888,1484,1510]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">29</figureCitation>
herein with
<figureCitation id="13476F85FFD0FF9CFBDA00C1B45EFA47" box="[1039,1124,1484,1510]" captionStart="FIGURES 24 ­ 31" captionStartId="7.[264,383,1684,1708]" captionText="FIGURES 24 ­ 31. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 24 ­ 27, MCZ 27836, male. 24, plumose setae on tarsus of second maxilla, scale 15 µm; 25, claw of second maxilla, scale 10 µm; 26, 27, dorsal and posterior views of pretarsus, leg 14, scales 20 µm, 15 µm. 28, MCZ 27838, male, first genital sternite and gonopods, scale 50 µm. 29 ­ 31, MCZ 47470, female gonopods, scales 100 µm (Figs. 29, 30), 50 µm (Fig. 31)." pageId="9" pageNumber="10">Fig. 24</figureCitation>
B of
<bibRefCitation id="EFED0EF1FFD0FF9CFB7400C1B17DF9AF" author="Edgecombe" pageId="9" pageNumber="11" refString="Edgecombe, G. D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201 - 241." type="journal article" year="2001">Edgecombe 2001</bibRefCitation>
). Analysis of molecular sequence data and combined molecular and morphological data indicates that the two species are each others' closest relative (
<bibRefCitation id="EFED0EF1FFD0FF9CFBF10311B17DF9FF" author="Edgecombe" pageId="9" pageNumber="11" refString="Edgecombe, G. D. &amp; Giribet, G. (2003) Relationships of Henicopidae (Chilopoda: Lithobiomorpha): new molecular data, classification and biogeography. In: Hamer, M. (Ed) Proceedings of the 12 th International Congress of Myriapodology. African Invertebrates, 44." type="book chapter" year="2003">Edgecombe &amp; Giribet 2003</bibRefCitation>
). Morphologically, they share the usual presence of 19 mostly elongate antennal articles, relatively large, bulging eyes (
<figureCitation id="13476F85FFD0FF9CFD700361B2D6F927" box="[677,748,1644,1670]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="9" pageNumber="10">Fig. 8</figureCitation>
), a small Tömösváry organ (
<figureCitation id="13476F85FFD0FF9CFBEF0361B4B7F927" box="[1082,1165,1644,1670]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="9" pageNumber="10">Fig. 11</figureCitation>
versus
<bibRefCitation id="EFED0EF1FFD0FF9CFB300361B1A0F90F" author="Edgecombe" pageId="9" pageNumber="11" refString="Edgecombe, G. D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201 - 241." type="journal article" year="2001">Edgecombe 2001</bibRefCitation>
, fig. 25M), four or five pointed teeth on each dental margin of the maxillipede in most specimens (
<figureCitation id="13476F85FFD0FF9CFE2003B1B276F977" box="[501,588,1724,1750]" captionStart="FIGURES 8 ­ 15" captionStartId="3.[264,383,1655,1679]" captionText="FIGURES 8 ­ 15. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington. 8, 12, 14, MCZ 27838, male. 8, dorsal view of head shield, scale 500 µm; 12, dorsal view of articles from middle of antenna, scale 100 µm; 14, ventral view of maxillipedes, scale 300 µm. 9 ­ 11, 13, 15, MCZ 27836, male. 9, 10, labrum and detail of marginal bristles, scales 100 µm, 10 µm; 11, cephalic pleurite with Tömösváry organ, scale 100 µm; 13, 15, dorsal view of maxillipedes and detail of dental margin (ventral view), scales 200 µm, 100 µm." pageId="9" pageNumber="10">Fig. 15</figureCitation>
), and strong articulations between tarsomeres. The mandibles are very similar, sharing many small, barb­like pinnules along both margins of the aciculae (
<figureCitation id="13476F85FFD0FF9CFEC40201B15FF887" box="[273,357,1804,1830]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="9" pageNumber="10">Fig. 17</figureCitation>
versus
<bibRefCitation id="EFED0EF1FFD0FF9CFE6A0201B2B7F887" author="Edgecombe" box="[447,653,1804,1830]" pageId="9" pageNumber="11" refString="Edgecombe, G. D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201 - 241." type="journal article" year="2001">Edgecombe 2001</bibRefCitation>
, fig. 25K) and abundant bristles in the furry pad that curl back towards the dorsalmost tooth (
<figureCitation id="13476F85FFD0FF9CFD7F0239B2C4F8EF" box="[682,766,1844,1870]" captionStart="FIGURES 16 ­ 23" captionStartId="5.[264,383,1684,1708]" captionText="FIGURES 16 ­ 23. Paralamyctes wellingtonensis n. sp. All from Puerto Edén, Isla Wellington except 19 ­ 20, from Parque Nacional Torres del Paine. 16 ­ 18, MCZ 27838, male. 16, gnathal edge of right mandible, scale 50 µm; 17, aciculae, scale 20 µm; 18, furry pad, scale 20 µm. 19 ­ 20, MCZ DNA 100408, male, ventral and dorsal parts of right mandible, scales 30 µm, 50 µm. 21 ­ 23, MCZ 27836, male. 21, first maxillae, scale 100 µm; 22, coxal process and telopodite of first maxilla, scale 50 µm; 23, inner margin of telopodite of first maxilla, scale 20 µm." pageId="9" pageNumber="10">Fig. 18</figureCitation>
versus Edgecombe et al. 2002, fig. 7D).
</paragraph>
</subSubSection>
<subSubSection id="C366208BFFD3FF9FFEED0411B2EEFCC7" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BC37300FFD3FF9FFEED0411B2EEFCC7" blockId="10.[264,1324,284,870]" pageId="10" pageNumber="11">
<taxonomicName id="4C7C0883FFD3FF9FFEED0411B270FE94" ID-CoL="4CXV9" box="[312,586,284,309]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B908AF12FFD3FF9FFEED0411B270FE94" box="[312,586,284,309]" italics="true" pageId="10" pageNumber="11">Paralamyctes chilensis</emphasis>
</taxonomicName>
was assigned by
<bibRefCitation id="EFED0EF1FFD3FF9FFCCA0411B43AFE97" author="Edgecombe" box="[799,1024,284,310]" pageId="10" pageNumber="11" refString="Edgecombe, G. D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201 - 241." type="journal article" year="2001">Edgecombe (2001)</bibRefCitation>
to the otherwise Australian subgenus
<emphasis id="B908AF12FFD3FF9FFE6A0449B1F4FEFC" box="[447,462,324,349]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<taxonomicName id="4C7C0883FFD3FF9FFE380449B2A0FEFC" box="[493,666,324,349]" class="Myriapoda" family="Curculionidae" genus="Nothofagobius" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B908AF12FFD3FF9FFE380449B2A0FEFC" box="[493,666,324,349]" italics="true" pageId="10" pageNumber="11">Nothofagobius</emphasis>
</taxonomicName>
). Morphological support for this assignment is also shared by
<taxonomicName id="4C7C0883FFD3FF9FFE520461B267FE24" ID-CoL="4CXVC" box="[391,605,364,389]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="wellingtonensis">
<emphasis id="B908AF12FFD3FF9FFE520461B267FE24" box="[391,605,364,389]" italics="true" pageId="10" pageNumber="11">P. wellingtonensis</emphasis>
</taxonomicName>
, including the extended first article of the female gonopod and bulging posterior margin of the male first genital sternite (
<figureCitation id="13476F85FFD3FF9FFC350499B41DFE0F" box="[992,1063,404,430]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). However, molecular data and their combination with morphological data (
<bibRefCitation id="EFED0EF1FFD3FF9FFCA904B1B4FDFE77" author="Edgecombe" box="[892,1223,444,470]" pageId="10" pageNumber="11" refString="Edgecombe, G. D. &amp; Giribet, G. (2003) Relationships of Henicopidae (Chilopoda: Lithobiomorpha): new molecular data, classification and biogeography. In: Hamer, M. (Ed) Proceedings of the 12 th International Congress of Myriapodology. African Invertebrates, 44." type="book chapter" year="2003">Edgecombe &amp; Giribet 2003</bibRefCitation>
) instead favour a closer relationship between the two Chilean species and the Australasian subgenera
<emphasis id="B908AF12FFD3FF9FFEE10701B179FD84" box="[308,323,524,549]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<emphasis id="B908AF12FFD3FF9FFE890701B1C0FD84" box="[348,506,524,549]" italics="true" pageId="10" pageNumber="11">Thingathinga</emphasis>
)
<bibRefCitation id="EFED0EF1FFD3FF9FFDD90701B2DEFD87" author="Edgecombe" box="[524,740,524,550]" pageId="10" pageNumber="11" refString="Edgecombe, G. D. (2001) Revision of Paralamyctes (Chilopoda: Lithobiomorpha: Henicopidae), with six new species from eastern Australia. Records of the Australian Museum, 53, 201 - 241." type="journal article" year="2001">Edgecombe, 2001</bibRefCitation>
, and
<emphasis id="B908AF12FFD3FF9FFCF60701B308FD84" box="[803,818,524,549]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<taxonomicName id="4C7C0883FFD3FF9FFC9E0701B387FD84" ID-CoL="4S7M" box="[843,957,524,549]" class="Agaricomycetes" family="Tricholomataceae" genus="Haasiella" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="11" phylum="Basidiomycota" rank="genus">
<emphasis id="B908AF12FFD3FF9FFC9E0701B387FD84" box="[843,957,524,549]" italics="true" pageId="10" pageNumber="11">Haasiella</emphasis>
</taxonomicName>
)
<bibRefCitation id="EFED0EF1FFD3FF9FFC1A0701B449FD87" author="Pocock" box="[975,1139,524,550]" pageId="10" pageNumber="12" refString="Pocock, R. I. (1901) Some new Genera and Species of Lithobiomorphous Chilopoda. Annals and Magazine of Natural History, 7, 448 - 451." type="journal article" year="1901">Pocock, 1901</bibRefCitation>
. The most congruent parameter set for combined data resolves the Chilean species with
<emphasis id="B908AF12FFD3FF9FFBBC0739B442FDEC" box="[1129,1144,564,589]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<taxonomicName id="4C7C0883FFD3FF9FFB450739B538FDEC" ID-CoL="4S7M" box="[1168,1282,564,589]" class="Agaricomycetes" family="Tricholomataceae" genus="Haasiella" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="11" phylum="Basidiomycota" rank="genus">
<emphasis id="B908AF12FFD3FF9FFB450739B538FDEC" box="[1168,1282,564,589]" italics="true" pageId="10" pageNumber="11">Haasiella</emphasis>
</taxonomicName>
) in particular. That relationship accounts for some shared morphological characters, notably bifid aciculae on the mandible, the only cases of this character state within
<taxonomicName id="4C7C0883FFD3FF9FFB570789B519FD3C" ID-CoL="6F5F" box="[1154,1315,644,669]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B908AF12FFD3FF9FFB570789B519FD3C" box="[1154,1315,644,669]" italics="true" pageId="10" pageNumber="11">Paralamyctes</emphasis>
</taxonomicName>
. In
<emphasis id="B908AF12FFD3FF9FFEF207A1B10CFD64" box="[295,310,684,709]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<taxonomicName id="4C7C0883FFD3FF9FFE9B07A1B1FAFD64" ID-CoL="4S7M" box="[334,448,684,709]" class="Agaricomycetes" family="Tricholomataceae" genus="Haasiella" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="11" phylum="Basidiomycota" rank="genus">
<emphasis id="B908AF12FFD3FF9FFE9B07A1B1FAFD64" box="[334,448,684,709]" italics="true" pageId="10" pageNumber="11">Haasiella</emphasis>
</taxonomicName>
) and in
<taxonomicName id="4C7C0883FFD3FF9FFDCA07A1B2CBFD64" ID-CoL="4CXVC" box="[543,753,684,709]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="wellingtonensis">
<emphasis id="B908AF12FFD3FF9FFDCA07A1B2CBFD64" box="[543,753,684,709]" italics="true" pageId="10" pageNumber="11">P. wellingtonensis</emphasis>
</taxonomicName>
the median furrow on the head shield is variably incised behind the transverse suture (
<figureCitation id="13476F85FFD3FF9FFD0707D9B321FD4F" box="[722,795,724,750]" captionStart="FIGURES 1 ­ 3" captionStartId="2.[264,383,1719,1743]" captionTargetBox="[278,1308,332,1688]" captionTargetId="figure@2.[273,1314,329,1693]" captionTargetPageId="2" captionText="FIGURES 1 ­ 3. Paralamyctes wellingtonensis n. sp. Puerto Edén, Isla Wellington, Chile. 1, holotype MCZ 52636, male, scale 1 mm; 2, MCZ 47468, female, terminal segments and gonopods, scale 100 µm; 3, MCZ 27836, male, terminal segments and gonopods, scale 100 µm." httpUri="https://zenodo.org/record/156320/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
). However, most parameter sets resolve
<emphasis id="B908AF12FFD3FF9FFAC007D9B51EFD4C" box="[1301,1316,724,749]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<taxonomicName id="4C7C0883FFD3FF9FFEC707F1B1BEFCB4" ID-CoL="4S7M" box="[274,388,764,789]" class="Agaricomycetes" family="Tricholomataceae" genus="Haasiella" kingdom="Fungi" order="Agaricales" pageId="10" pageNumber="11" phylum="Basidiomycota" rank="genus">
<emphasis id="B908AF12FFD3FF9FFEC707F1B1BEFCB4" box="[274,388,764,789]" italics="true" pageId="10" pageNumber="11">Haasiella</emphasis>
</taxonomicName>
) closer to
<emphasis id="B908AF12FFD3FF9FFDD707F1B22BFCB4" box="[514,529,764,789]" italics="true" pageId="10" pageNumber="11">P</emphasis>
. (
<emphasis id="B908AF12FFD3FF9FFDFC07F1B2FDFCB4" box="[553,711,764,789]" italics="true" pageId="10" pageNumber="11">Thingathinga</emphasis>
) than to the Chilean species (
<bibRefCitation id="EFED0EF1FFD3FF9FFBF007F1B17DFC9F" author="Edgecombe" pageId="10" pageNumber="11" refString="Edgecombe, G. D. &amp; Giribet, G. (2003) Relationships of Henicopidae (Chilopoda: Lithobiomorpha): new molecular data, classification and biogeography. In: Hamer, M. (Ed) Proceedings of the 12 th International Congress of Myriapodology. African Invertebrates, 44." type="book chapter" year="2003">Edgecombe &amp; Giribet 2003</bibRefCitation>
). Because the relationships of the Chilean clade are unstable, a subgeneric assignment within
<taxonomicName id="4C7C0883FFD3FF9FFE490641B207FCC4" ID-CoL="6F5F" box="[412,573,844,869]" class="Chilopoda" family="Henicopidae" genus="Paralamyctes" kingdom="Animalia" order="Lithobiomorpha" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B908AF12FFD3FF9FFE490641B207FCC4" box="[412,573,844,869]" italics="true" pageId="10" pageNumber="11">Paralamyctes</emphasis>
</taxonomicName>
is not made.
</paragraph>
</subSubSection>
</treatment>
</document>
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