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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="001178297187810C9CCCBE7793A178FF" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole typhlos Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="99FD33D4-9BC1-4595-9231-1A85822A7B35" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>905</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>235</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161776994" LSID="urn:lsid:zoobank.org:act:99FD33D4-9BC1-4595-9231-1A85822A7B35" httpUri="http://treatment.plazi.org/id/001178297187810C9CCCBE7793A178FF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/99FD33D4-9BC1-4595-9231-1A85822A7B35" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos" status="sp. nov.">Pheidole typhlos</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">Figs 20A-F</figureCitation>
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85X</figureCitation>
<figureCitation captionStart="Figure 88" captionStartId="F88" captionText="Figure 88. Distribution. Pheidole petax Forel (A). P. podargea sp. nov. (B). P. praegrandis sp. nov. (C). P. ranohirensis sp. nov. (D). P. rugocephala sp. nov. (E). P. rugofitarata sp. nov. (F). P. scabrata Forel (G). P. typhlos sp. nov. (H). P. vatovavensis sp. nov. (I). P. voasara sp. nov. (J). P. vohemarensis sp. nov. (K). P. zavamanira sp. nov. (L)." figureDoi="10.3897/zookeys.905.39592.figure88" httpUri="https://binary.pensoft.net/fig/373135" pageId="0" pageNumber="1">, 88H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Antsiranana; Galoko chain, Mont Galoko;
<geoCoordinate degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
,
<geoCoordinate degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
; alt. 520 m; 19 Feb 2013; B.L. Fisher et al. leg.; BLF30851, CASENT0302953 (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •1 w.; same data as for holotype; CASENT0923217 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antananarivo</emphasis>
: •6s.; Galoko chain, Mont Galoko;
<geoCoordinate degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
,
<geoCoordinate degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
; alt. 520 m; 16 Feb 2013; B.L. Fisher et al. leg.; CASENT0305030 (CASC). •1w.; Galoko chain, Mont Galoko;
<geoCoordinate degrees="13.5888" direction="south" orientation="latitude" precision="5" value="-13.5888">-13.5888</geoCoordinate>
,
<geoCoordinate degrees="48.72864" direction="east" orientation="longitude" precision="1" value="48.72864">48.72864</geoCoordinate>
; alt. 980 m; 22 Feb 2013; B.L. Fisher et al. leg.; CASENT0304748 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Eyes small and reduced; body yellow; head rectangular, anterior of eyes relatively straight, posterior of eyes slightly convex; sides of the head with sparse, short, erect pilosity; occipital lobes, genae, and posterior part of frons shiny and smooth, sometimes with indistinct, very sparse, longitudinal rugulae; anterior part of frons and malar area shiny, with thick, very sparse, short, and interrupted rugae, interspaces smooth; propodeal spines short, triangular, with acute top; pronotum smooth and shiny, sometimes on lateral sides with indistinct and sparse rugulae; mesonotum and propodeum shiny, with fine, sparse to moderately sparse rugoreticulation; katepisternum indistinctly foveolate; anepisternum with thick rugoreticulation; inner hypostomal teeth distinct, moderately high, narrow, triangular, with rounded apex; outer hypostomal teeth distinct, triangular, and with rounded tops directed outward; outer hypostomal teeth are slightly lower than inner hypostomal teeth.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Eyes small and reduced; body yellow; head and mesosoma shiny and smooth, lateral sides of frons with short, indistinct, longitudinal rugulae; propodeal spines short and triangular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 8): HL: 1.2-1.34 (1.28); HW: 0.96-1.07 (1.01); SL: 0.49-0.52 (0.5); EL: 0.08-0.12 (0.1); WL: 0.79-0.91 (0.84); PSL: 0.15-0.19 (0.16); MTL: 0.48-0.52 (0.51); PNW: 0.48-0.55 (0.51); PTW: 0.14-0.16 (0.15); PPW: 0.37-0.43 (0.4); CI: 75.1-82.0 (79.5); SI: 48.0-52.3 (49.8); PSLI: 11.9-14.3 (12.7); PPI: 36.3-39.9 (37.9); PNI: 47.4-55.3 (50.6); MTI: 46.9-52.9 (49.8).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view elongate, anterior of eyes relatively straight, posterior of eyes slightly convex (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces relatively flat; inner hypostomal teeth invisible. Antennal scrobes absent. Sides of the head with sparse, short, erect pilosity; whole head with dense, short, suberect to erect pilosity. Occipital lobes, genae, and posterior part of frons shiny and smooth, sometimes with indistinct, very sparse, longitudinal rugulae; anterior part of frons and malar area shiny, with thick, very sparse, short, and interrupted rugae, interspaces smooth. Clypeus shiny and smooth, with a few indistinct, longitudinal rugae on the lateral sides; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20B, D</figureCitation>
). Inner hypostomal teeth distinct, moderately high, narrow, triangular, with rounded apex; outer hypostomal teeth distinct, triangular, and with rounded tops directed outward; outer hypostomal teeth are slightly lower than inner hypostomal teeth (Fig.
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85X</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum short, angular, and moderately high, posterior mesonotum convex, with indistinct tubercle-like projection; promesonotal groove absent; metanotal groove absent or very indistinct; propodeal spines short, triangular, with acute top; humeral area with wide and flat tubercles (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D</figureCitation>
). Pronotum smooth and shiny, sometimes on lateral sides with indistinct and sparse rugulae; mesonotum and propodeum shiny, with fine, sparse to moderately sparse rugoreticulation; katepisternum indistinctly foveolate; anepisternum with thick rugoreticulation. Pilosity moderately dense, long, and erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny; peduncle finely foveolate, without horizontal lobes on its basal part; node smooth, moderately high, with convex apex, in rear view node dorsoventrally concave; pilosity moderately dense and erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Finely foveolate and shiny; in dorsal view sides with moderately long, acute, and triangular projections; pilosity moderately long, moderately dense, and erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
.
</emphasis>
Smooth and shiny; pilosity dense, short, and erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Yellow, malar area and lower part of frons dark yellow to bright brown (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923217) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0302953).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 0.55; HW: 0.46; SL: 0.45; EL: 0.05; WL: 0.62; PSL: 0.09; MTL: 0.35; PNW: 0.32; PTW: 0.09; PPW: 0.14; CI: 83.9; SI: 97.4; PSLI: 16.4; PPI: 61.2; PNI: 69.8; MTI: 76.7.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view rectangular, posterior and anterior of eyes slightly convex, occipital margin straight or indistinctly concave; occipital carina absent (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20A</figureCitation>
). Pilosity moderately dense, short, subdecumbent to suberect. Whole head shiny and smooth, only lateral sides of frons with short, indistinct, longitudinal rugulae; antennal sockets with sparse carinae curved outward. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, reaching posterior head margin; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum box-like; promesonotal groove absent; metanotal groove distinct; propodeal spines short and triangular, apex acute (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C</figureCitation>
). Sculpture smooth and shiny, sometimes lateral sides with indistinct, irregular, and sparse rugae. Pilosity moderately sparse, short, and erect (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle short with ventral face slightly convex; node globular; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gaster</emphasis>
.
</emphasis>
Pilosity moderately dense, erect, and short (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellow (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pheidole typhlos sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923217) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0302953)." figureDoi="10.3897/zookeys.905.39592.figure20" httpUri="https://binary.pensoft.net/fig/373067" pageId="0" pageNumber="1">20C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Greek for blind [τυφλός], in reference to the reduced eyes.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at elevation between 520-980 m, in rainforest, in montane forest. Nesting preferences unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is most similar to
<taxonomicName lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
<emphasis italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
<emphasis italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
</taxonomicName>
sp. nov. by sparse and short pilosity on sides of the head, presence of distinct, triangular inner hypostomal teeth and presence of short, triangular propodeal spines.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole typhlos" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlos">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole typhlos</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. parviocula" pageId="0" pageNumber="1" rank="species" species="parviocula">
<emphasis italics="true" pageId="0" pageNumber="1">P. parviocula</emphasis>
</taxonomicName>
sp. nov. by box-like promesonotum, absence of promesonotal groove and presence of short and triangular propodeal spines.
</paragraph>
</subSubSection>
</treatment>
</document>
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