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<mods:title id="6A7DC53E3496FCFA61EE55B95E53A508">Molecular evidence and differences in gonopod morphology lead to the recognition of a new species of the freshwater crab genus Candidiopotamon Bott, 1967 (Crustacea, Brachyura, Potamidae) from eastern Taiwan</mods:title>
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<mods:namePart id="84B8524F47AE0C489209DE6E7C0A2EAB">Shih, Hsi-Te</mods:namePart>
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<mods:affiliation id="3CF94F1F6ACC49C3177BAB645F574FF5">Department of Life Science and Global Change Biology Research Center, National Chung Hsing University, 250, Kuo Kuang Road, Taichung 402, Taiwan</mods:affiliation>
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<mods:namePart id="B597612120B9F7730AAD57E55A9A3019">Naruse, Tohru</mods:namePart>
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<mods:affiliation id="AF8A887272E517D3806F0BAB779A144D">Tropical Biosphere Research Center, Iriomote Station, University of the Ryukyus, 870 Uehara, Taketomi, Okinawa 907 - 1541, Japan</mods:affiliation>
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<mods:namePart id="5D687E5E21AAD3BDFAB16EF414D885AA">Schubart, Christoph D.</mods:namePart>
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<subSection id="4243BF311585072BF31DA0E9B5810E57" pageId="0" pageNumber="169" type="results">
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<taxonomicName id="F20D7956C0BB00481688AB9709DF4868" LSID="https://zoobank.org/3AC24730-7610-4C20-8F55-963FE8C56D6A" authority="Shih &amp; Naruse &amp; Schubart, 2023" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon penglai" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="penglai" status="sp. nov.">Candidiopotamon penglai</taxonomicName>
<taxonomicNameLabel id="E0F977EFE16C920E944C523096CE85B4" pageId="0" pageNumber="169">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="B5775338F4264EE218455CCE4EA93EC6" pageId="0" pageNumber="169">
<figureCitation id="56DF736766F46BACB8B1EFB002218F2F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A, B) and Candidiopotamon penglai sp. nov. (C-F). A, C, E overall dorsal view B, D, F overall ventral view A, B NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) C, D NCHUZOOL 15204 (male, 39.7 x 34.4 mm, Nanrenshan, Pingtung, southern Taiwan); NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure4" httpUri="https://binary.pensoft.net/fig/904110" pageId="0" pageNumber="169">Figs 4C-F</figureCitation>
<figureCitation id="418B0CAB528E7AF30D16CA63AAF377E4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A-D) and Candidiopotamon penglai sp. nov. (E-H). A, B NCHUZOOL 15194 (male, 35.4 x 31. 2 mm, Tai-an, Miaoli, northwestern Taiwan) C NCHUZOOL 15211 (male, 24.8 x 22.1 mm, Dongshih, Taichung, western Taiwan) D NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) E, F NCHUZOOL 15204 (2 males, 39.7 x 34.4 mm, 40.6 x 34.3 mm, Nanrenshan, Pingtung, southern Taiwan) G NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan) H NCHUZOOL 15229 (male, 33.1 x 29.7 mm, Jili Lake, Guangfu, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure5" httpUri="https://binary.pensoft.net/fig/904111" pageId="0" pageNumber="169">, 5E-H</figureCitation>
<figureCitation id="B5AEB8FBE91A29DF792DF7DFF529B317" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Candidiopotamon penglai sp. nov., holotype male (38.4 x 33.4 mm) from Ji-an, Hualien (NCHUZOOL 12951), G 1 in bent form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal segment, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure6" httpUri="https://binary.pensoft.net/fig/904112" pageId="0" pageNumber="169">, 6</figureCitation>
<figureCitation id="3A2B16E8988CEB79112225E01C3088CF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Candidiopotamon penglai sp. nov., male (42.9 x 36.2 mm) from Shihzih, Pingtung (NCHUZOOL 15207), G 1 in straight form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal joint, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure7" httpUri="https://binary.pensoft.net/fig/904113" pageId="0" pageNumber="169">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A56694C62771A7C4BF19BC1FBD3CA717" pageId="0" pageNumber="169" type="reference_group">
<paragraph id="11734705C47A4D5972263AA3E94B9B2D" pageId="0" pageNumber="169">
<taxonomicName id="D758CEB58F39D7B9047AB2AA0E4077DE" authority="Shih &amp; Naruse &amp; Schubart, 2023" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamon rathbuni" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="rathbuni" status="sp. nov.">Potamon rathbuni</taxonomicName>
-
<bibRefCitation id="3F23AA58C69D8DD4DED151E9EE478AA0" author="Chiu, JK" journalOrPublisher="Bulletin of the Institute of Zoology, Academia Sinica" pageId="0" pageNumber="169" pagination="63 - 73" refId="B10" refString="Chiu, JK, 1964. Prevalence of Paragonimus infection in crab hosts in Taiwan. Bulletin of the Institute of Zoology, Academia Sinica 3: 63 - 73" title="Prevalence of Paragonimus infection in crab hosts in Taiwan." volume="3" year="1964">Chiu 1964</bibRefCitation>
: 67 (part).
</paragraph>
<paragraph id="0AFB7083EC0C9CF5AC171E8161EA1664" pageId="0" pageNumber="169">
<taxonomicName id="61D72BB1DBDC518ABE4E0C5B916FA8F8" authority="Shih &amp; Naruse &amp; Schubart, 2023" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon rathbuni" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="rathbuni" status="sp. nov.">Candidiopotamon rathbuni</taxonomicName>
-
<bibRefCitation id="8246A1ED0B8A92255927083F834CA5B0" DOI="https://doi.org/10.5109/22860" author="Minei, H" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="0" pageNumber="169" pagination="239 - 251" refId="B48" refString="Minei, H, 1974. Potamoid crabs of Taiwan, with description of one new species (Crustacea, Decapoda). Journal of the Faculty of Agriculture, Kyushu University 18 (4): 239 - 251, DOI: https://doi.org/10.5109/22860" title="Potamoid crabs of Taiwan, with description of one new species (Crustacea, Decapoda)." url="https://doi.org/10.5109/22860" volume="18" year="1974">Minei 1974</bibRefCitation>
: 246, figs 7-9 (part); CH
<bibRefCitation id="DD7DDC3F775D417DC9F565129666A1E0" author="Wang, CH" journalOrPublisher="Annual of National Taiwan Museum" pageId="0" pageNumber="169" pagination="39 - 44" refId="B99" refString="Wang, CH, 1984. Primary studies on the crabs in Kenting National Park in Taiwan. Annual of National Taiwan Museum 27: 39 - 44" title="Primary studies on the crabs in Kenting National Park in Taiwan." volume="27" year="1984">Wang 1984</bibRefCitation>
: 41;
<bibRefCitation id="190D02CA1897D8DC33D6E77DDC40F0FB" author="Yu, HP" journalOrPublisher="Crustaceana" pageId="0" pageNumber="169" refId="B103" refString="Yu, HP, Jeng, MS, Chan, TY, Ho, PH, Shy, JY, 1996. Studies on the Land and Aquatic Decapod Crustacean Fauna of the Kenting National Park. Kenting National Park, Pingtung, Taiwan, [2+]79 pp. [In Chinese]" title="Studies on the Land and Aquatic Decapod Crustacean Fauna of the Kenting National Park. Kenting National Park, Pingtung, Taiwan, [2 +] 79 pp. [In Chinese]" year="1996">Yu et al. 1996</bibRefCitation>
: 9, 29, fig. 10; MS
<bibRefCitation id="CA53681361A1346172AD36CB20DCA5E5" author="Jeng, MS" journalOrPublisher="Kenting National Park, Pingtung, Taiwan" pageId="0" pageNumber="169" refId="B28" refString="Jeng, MS, 1998. The Prawns and Crabs of Kenting National Park. Kenting National Park, Pingtung, Taiwan" title="The Prawns and Crabs of Kenting National Park." year="1998">Jeng 1998</bibRefCitation>
: 86, 1 unnumbered fig.;
<bibRefCitation id="D6A591E048790D24C9ADBF0DCB9746A7" author="Shy, JY" journalOrPublisher="National Museum of Marine Biology and Aquarium, Pingtung, Taiwan" pageId="0" pageNumber="169" refId="B90" refString="Shy, JY, Yu, HP, 1999. Freshwater crabs of Taiwan. National Museum of Marine Biology and Aquarium, Pingtung, Taiwan" title="Freshwater crabs of Taiwan." year="1999">Shy and Yu 1999</bibRefCitation>
: 94, 3 unnumbered figs, &quot;fig. 31&quot; on p. 106 (part);
<bibRefCitation id="EBE2CF9246495E4F032901B0F9A12087" author="Ng, PKL" journalOrPublisher="Crustaceana" pageId="0" pageNumber="169" pagination="249 - 251" refId="B56" refString="Ng, PKL, 2000. Book review: A.-Y. Dai. 1999. Fauna Sinica. Crustacea: Decapoda: Parathelphusidae, Potamidae: 1-501., pls. 1-30. (Science Press, 16 Donghuangchenggen, Bei Jei, Beijing 100717, People's Republic of China). [In Chinese with English abstract.] [ISBN 7-03-006407-0] Hard cover. Crustaceana 73: 249 - 251" title="Book review: A. - Y. Dai. 1999. Fauna Sinica. Crustacea: Decapoda: Parathelphusidae, Potamidae: 1 - 501., pls. 1 - 30. (Science Press, 16 Donghuangchenggen, Bei Jei, Beijing 100717, People's Republic of China). [In Chinese with English abstract.] [ISBN 7 - 03 - 006407 - 0] Hard cover." volume="73" year="2000">Ng 2000</bibRefCitation>
: 249 (part);
<bibRefCitation id="6A606B0D615B96B48A9954C3676F60DE" DOI="https://doi.org/10.6532/JNTM.201712_70(3;4).01" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="169" refId="B61" refString="Ng, PKL, Shih, HT, Ho, PH, Wang, CH, 2017. An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum 70(3/4): 1-208. https://doi.org/10.6532/JNTM.201712_70(3;4).01" title="An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum 70 (3 / 4): 1 - 208." url="https://doi.org/10.6532/JNTM.201712_70(3;4).01" year="2017">Ng et al. 2017</bibRefCitation>
: 76;
<bibRefCitation id="62B8A4E9500B2A6582064BB6477FED48" author="Shy, JY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="169" refId="B94" refString="Shy, JY, Shih, HT, Ng, PKL, 2020. Crustacean Fauna of Taiwan: Brachyuran Crabs, Vol. III - Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, Taiwan, [26+]232 pp." title="Crustacean Fauna of Taiwan: Brachyuran Crabs, Vol. III - Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, Taiwan, [26 +] 232 pp." year="2020">Shy et al. 2020</bibRefCitation>
: 2, figs 11A, 13, 14, 16, 17 (part);
<bibRefCitation id="514642F03C8338280F40BEC6F1C1DA3E" author="Li, JJ" journalOrPublisher="National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" pageId="0" pageNumber="169" refId="B41" refString="Li, JJ, Chiu, YW, 2019a. An Atlas of Land Crabs on Hengchun Peninsula 2.0. National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" title="An Atlas of Land Crabs on Hengchun Peninsula 2.0." year="2019 a">Li and Chiu 2019a</bibRefCitation>
: 103, 3 unnumbered figs;
<bibRefCitation id="F05E9630126EDED3808328798C7E8B2D" author="Li, JJ" journalOrPublisher="National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" pageId="0" pageNumber="169" refId="B42" refString="Li, JJ, Chiu, YW, 2019b. A Field Guide of Land Crabs on Hengchun Peninsula 2.0. National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" title="A Field Guide of Land Crabs on Hengchun Peninsula 2.0." year="2019 b">Li and Chiu 2019b</bibRefCitation>
: 2 unnumbered figs on p. 56;
<bibRefCitation id="D188842188EE8EE9738CDABA6CA9FB97" author="Shy, JY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="169" refId="B94" refString="Shy, JY, Shih, HT, Ng, PKL, 2020. Crustacean Fauna of Taiwan: Brachyuran Crabs, Vol. III - Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, Taiwan, [26+]232 pp." title="Crustacean Fauna of Taiwan: Brachyuran Crabs, Vol. III - Freshwater Crabs - Potamidae, Gecarcinucidae. National Penghu University of Science and Technology, Penghu, Taiwan, [26 +] 232 pp." year="2020">Shy et al. 2020</bibRefCitation>
: 2, figs 2E, 14E (part); SW
<bibRefCitation id="6D434C984C751D02DD235F89F0D16CDC" author="Wang, SW" journalOrPublisher="Collection and Research" pageId="0" pageNumber="169" pagination="29 - 40" refId="B100" refString="Wang, SW, Chiu, YW, Huang, HD, Lee, KS, Wang, WL, Liu, SL, Yang, SH, 2021. Notes on freshwater coated grains of the Hengchun Peninsula. Collection and Research 33: 29 - 40" title="Notes on freshwater coated grains of the Hengchun Peninsula." volume="33" year="2021">Wang et al. 2021</bibRefCitation>
: 32, pl. 3E.
</paragraph>
<paragraph id="00277C9AC55599D6AAAD335A073CB8AC" pageId="0" pageNumber="169">
<taxonomicName id="DB62BC41F92BFCFCBA4AE98664D6D446" authority="Shih &amp; Naruse &amp; Schubart, 2023" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon rathbunae" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="rathbunae" status="sp. nov.">Candidiopotamon rathbunae</taxonomicName>
-
<bibRefCitation id="181D85869EB80E50C4A6D52B59B4E99C" author="Ng, PKL" journalOrPublisher="Crustaceana" pageId="0" pageNumber="169" pagination="249 - 251" refId="B56" refString="Ng, PKL, 2000. Book review: A.-Y. Dai. 1999. Fauna Sinica. Crustacea: Decapoda: Parathelphusidae, Potamidae: 1-501., pls. 1-30. (Science Press, 16 Donghuangchenggen, Bei Jei, Beijing 100717, People's Republic of China). [In Chinese with English abstract.] [ISBN 7-03-006407-0] Hard cover. Crustaceana 73: 249 - 251" title="Book review: A. - Y. Dai. 1999. Fauna Sinica. Crustacea: Decapoda: Parathelphusidae, Potamidae: 1 - 501., pls. 1 - 30. (Science Press, 16 Donghuangchenggen, Bei Jei, Beijing 100717, People's Republic of China). [In Chinese with English abstract.] [ISBN 7 - 03 - 006407 - 0] Hard cover." volume="73" year="2000">Ng 2000</bibRefCitation>
: 249 (part);
<bibRefCitation id="03BFCA106ECA3F256E5665657BECC6EF" author="Chen, WJ" journalOrPublisher="Nature Conservation Quarterly" pageId="0" pageNumber="169" pagination="42 - 47" refId="B6" refString="Chen, WJ, Shy, JY, Cheng, JH, Hsu, MJ, 2001. Freshwater crabs in the water systems of Kaohsiung and Pingtung areas. Nature Conservation Quarterly 36: 42 - 47" title="Freshwater crabs in the water systems of Kaohsiung and Pingtung areas." volume="36" year="2001">Chen et al. 2001</bibRefCitation>
: 43, 2 unnumbered figs on p. 43, 46 (part);
<bibRefCitation id="DB71314D9C22AB9F7E8304F2C05284E1" author="Lee, JH" journalOrPublisher="Bigtrees Co., Taipei" pageId="0" pageNumber="169" refId="B38" refString="Lee, JH, 2001. A Field Guide to Crabs in Taiwan. Bigtrees Co., Taipei" title="A Field Guide to Crabs in Taiwan." year="2001">Lee 2001</bibRefCitation>
: 145, 2 unnumbered figs (part);
<bibRefCitation id="2F5052A834204B2D041C38CD21C53082" author="Ng, PKL" journalOrPublisher="National Taiwan Museum Special Publication Series" pageId="0" pageNumber="169" pagination="1 - 86" refId="B62" refString="Ng, PKL, Wang, CH, Ho, PH, Shih, HT, 2001. An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum Special Publication Series 11: 1 - 86" title="An annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda)." volume="11" year="2001">Ng et al. 2001</bibRefCitation>
: 49 (part);
<bibRefCitation id="23DF26C911A7B14C03699E75B9C85D5D" author="Chen, WJ" journalOrPublisher="Kaohsiung County Government, Kaohsiung, Taiwan" pageId="0" pageNumber="169" refId="B7" refString="Chen, WJ, Tsai, SH, Hsu, MJ, Cheng, JH, 2003. Freshwater Crab Fauna in Southern Taiwan. Kaohsiung County Government, Kaohsiung, Taiwan" title="Freshwater Crab Fauna in Southern Taiwan." year="2003">Chen et al. 2003</bibRefCitation>
: 29, 5 unnumbered figs (part);
<bibRefCitation id="6CF5AD8DF6D6D268C1040CBB526E745F" DOI="https://doi.org/10.1111/j.1365-2699.2006.01472.x" author="Shih, HT" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="169" pagination="980 - 989" refId="B86" refString="Shih, HT, Hung, HC, Schubart, CD, Chen, CA, Chang, HW, 2006. Intraspecific genetic diversity of the endemic freshwater crab Candidiopotamon rathbunae (Decapoda, Brachyura, Potamidae) reflects five million years of geological history of Taiwan. Journal of Biogeography 33 (6): 980 - 989, DOI: https://doi.org/10.1111/j.1365-2699.2006.01472.x" title="Intraspecific genetic diversity of the endemic freshwater crab Candidiopotamon rathbunae (Decapoda, Brachyura, Potamidae) reflects five million years of geological history of Taiwan." url="https://doi.org/10.1111/j.1365-2699.2006.01472.x" volume="33" year="2006">Shih et al. 2006</bibRefCitation>
: 983 (part);
<bibRefCitation id="C8A328A0F0C36C4E5FAE9F4260ECF09F" author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="169" refId="B60" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology (Supplement 17): 1-296." title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology (Supplement 17): 1 - 296." year="2008">Ng et al. 2008</bibRefCitation>
: 161 (part);
<bibRefCitation id="D0A33F8A67D069C446274BB21B0088A0" author="Shy, JY" journalOrPublisher="Morning Star Publishing, Taipei" pageId="0" pageNumber="169" refId="B89" refString="Shy, JY, Lee, PW, 2009. Guide to the Taiwanese Freshwater Crabs. Morning Star Publishing, Taipei" title="Guide to the Taiwanese Freshwater Crabs." year="2009">Shy and Lee 2009</bibRefCitation>
: 207, 10 unnumbered figs (part);
<bibRefCitation id="800F7826386F39237D6DD9ACA6C0A717" author="Li, JJ" journalOrPublisher="National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" pageId="0" pageNumber="169" refId="B40" refString="Li, JJ, Chiu, YW, 2013. Land Crabs of Hengchun Peninsula. National Museum of Marine Biology &amp; Aquarium, Pingtung, Taiwan" title="Land Crabs of Hengchun Peninsula." year="2013">Li and Chiu 2013</bibRefCitation>
: 70, 3 unnumbered figs;
<bibRefCitation id="FA790CD0117875F01B1758E8196D425B" author="Chiu, YW" journalOrPublisher="Forestry Bureau, Council of Agriculture, Executive Yuan, Taipei" pageId="0" pageNumber="169" refId="B11" refString="Chiu, YW, Liang, SH, Hsieh, BS, Huang, DJ, Gan, YC, 2015. 50 selected springs of Taiwan. Forestry Bureau, Council of Agriculture, Executive Yuan, Taipei" title="50 selected springs of Taiwan." year="2015">Chiu et al. 2015</bibRefCitation>
: 202, 227, 1 unnumbered fig. on p. 202 (part);
<bibRefCitation id="F89F4FBC1E6675DCD6AD7B21175915F1" author="Ho, PH" editor="Shao, KT" journalOrPublisher="Fisheries Agency, Council of Agriculture, R. O. C., Taipei" pageId="0" pageNumber="169" pagination="136 - 159" refId="B21" refString="Ho, PH, 2015. Arthropoda: crabs. In: Shao, KT, Chang, JS, Jeng, MS, Tu, TH, Ho, CW, Chiu, YW, Chan, TY, Ho, PH, Chao, SM, Joung, SJ, Lin, PL, Eds., A Guide Book of Common Economic Aquatic Animals and Plants in Taiwan. Fisheries Agency, Council of Agriculture, R.O.C., Taipei: 136 - 159" title="Arthropoda: crabs." volumeTitle="A Guide Book of Common Economic Aquatic Animals and Plants in Taiwan." year="2015">Ho 2015</bibRefCitation>
: 141, 2 unnumbered figs (part);
<bibRefCitation id="BE50794BB51041AA1C2889AC6C127DE9" DOI="https://doi.org/10.13140/RG.2.2.22353.89446" author="Sasaki, J" journalOrPublisher="Local Independent Administrative Agency Hokkaido Research Organization, Resources Management and Enhancement Division, Abashiri Fisheries Research Institute, Fisheries Research Department, Hokkaido, Japan" pageId="0" pageNumber="169" refId="B75" refString="Sasaki, J, 2019. The Species List of Decapoda, Euphausiacea, and Stomatopoda, all of the World, version 03-3.1. Local Independent Administrative Agency Hokkaido Research Organization, Resources Management and Enhancement Division, Abashiri Fisheries Research Institute, Fisheries Research Department, Hokkaido, Japan, DOI: https://doi.org/10.13140/RG.2.2.22353.89446" title="The Species List of Decapoda, Euphausiacea, and Stomatopoda, all of the World, version 03 - 3.1." url="https://doi.org/10.13140/RG.2.2.22353.89446" year="2019">Sasaki 2019</bibRefCitation>
: 11552 (part).
</paragraph>
</subSubSection>
<subSubSection id="A446F55257C442E2490708DDDF4FB7FD" pageId="0" pageNumber="169" type="type material">
<paragraph id="10010199DDD3774844A77DA8E0561AA5" pageId="0" pageNumber="169">Type material.</paragraph>
<paragraph id="2AEBB615FDEF4B884FB634025ED3E691" pageId="0" pageNumber="169">
Taiwan -
<emphasis id="C9595DA2C6F0568BBA0857104D63B33A" bold="true" italics="true" pageId="0" pageNumber="169">Holotype</emphasis>
: male, 38.4
<normalizedToken id="67E205BB0DFC59E382251D02F4F96C05" originalValue="×">x</normalizedToken>
33.4 mm, NCHUZOOL 12951, Ji-an, Hualien, coll. H.-C. Liu, 4 May 2000. -
<emphasis id="1ECDBDD6F1583FB23D0993B247ED3B6E" bold="true" italics="true" pageId="0" pageNumber="169">Paratypes</emphasis>
: 1 male, 35.6
<normalizedToken id="5A06CC007E5470573AD60A930D020240" originalValue="×">x</normalizedToken>
30.5 mm, RUMF-ZC-8164, 1 female, 18.0
<normalizedToken id="D513EE2F706F14EA499EDB0C84B18ED3" originalValue="×">x</normalizedToken>
15.8 mm, NCHUZOOL 12932, collection data same as for holotype; 1 female, 34.3
<normalizedToken id="546D3DACF07B1463C8714D7289E8AC4A" originalValue="×">x</normalizedToken>
29.6 mm, RUMF-ZC-8165, Shueiyuandi, Shoufeng, Hualien, coll. N.-H. Jang-Liaw, 5 Mar. 2002; 1 male, 23.4
<normalizedToken id="0AEEAEEAF3F3DB6EB9CE268360DF1887" originalValue="×">x</normalizedToken>
20.0.5 mm, NCHUZOOL 12952, Liyu Lake, Shoufeng, Hualien, coll. H.-T. Shih, 26 Jan. 2007; 1 male, 27.5
<normalizedToken id="FD7311CFDB7839163F1757B12710AF16" originalValue="×">x</normalizedToken>
24.2 mm, NCHUZOOL 15213, Liyu Lake, Shoufeng, Hualien, coll. H.-T. Shih, 28 Jan. 2007; 1 male, 36.4
<normalizedToken id="976F9D9BDD7A103B79E8235D78BDA957" originalValue="×">x</normalizedToken>
30.9 mm, NCHUZOOL 15224, 1 male, 27.0
<normalizedToken id="0E3CE65D049192DDAFCCF26A377A5A89" originalValue="×">x</normalizedToken>
23.8, NCHUZOOL 15226, 1 female, 36.5
<normalizedToken id="F746D0272D996A450B04319A4591A9CE" originalValue="×">x</normalizedToken>
31.4, NCHUZOOL 15225, Liyu Lake, Shoufeng, Hualien, coll.
<normalizedToken id="2C439A941AEBBF683E47374C071DE0AB" originalValue="HTSs">HTS's</normalizedToken>
lab students, 13 Apr. 2023; 1 male, 28.3
<normalizedToken id="78212D5443536F0C649EE47DDF2D00AE" originalValue="×">x</normalizedToken>
24.8 mm, 1 female, 35.2
<normalizedToken id="8E5564D17C99D58BB1CF6E31AC5AB729" originalValue="×">x</normalizedToken>
31.6 mm, ZRC 2023.0309, Liyu Lake, Shoufeng, Hualien, coll.
<normalizedToken id="7DAA3C1702B5863EEB027EAA7D8E3246" originalValue="HTSs">HTS's</normalizedToken>
lab students, 13 Apr. 2023.
</paragraph>
</subSubSection>
<subSubSection id="D7D01410065D6CD165C487E49C11EC77" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="FB37D42910943148D6C1BCC3924D346E" pageId="0" pageNumber="169">Other material.</paragraph>
<paragraph id="9FF2991FE427F2724FBBBC603A4628CC" pageId="0" pageNumber="169">
<materialsCitation id="A04A41AB85E126568A1FB6DA4EBDE761" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Taiwan" location="Taiwan" specimenCount="1" specimenCount-male="1">
<collectingCountry id="39972BEFAB21BE0964E64A941019CCE2" name="Taiwan">Taiwan</collectingCountry>
-
<specimenCount id="059662F94DF49F29F0AF3EE88494B753" type="male">1 male</specimenCount>
, 23.9
<normalizedToken id="26CE5799A60AF2E52A7FA80554FF8ECC" originalValue="×">x</normalizedToken>
<quantity id="BB103497DA85DEE6DDEEF76DDA68C595" metricMagnitude="-2" metricUnit="m" metricValue="2.01" unit="mm" value="20.1">20.1 mm</quantity>
, NCHUZOOL 15227
</materialsCitation>
,
<specimenCount id="5BFB05D4520783E19827BB1A9FEF5207" type="male">1 male</specimenCount>
, 29.3
<normalizedToken id="3E34A2861C467FA024B8F42AD485525A" originalValue="×">x</normalizedToken>
<quantity id="35AD70A5B282C36EC978F75AB9D0C6A3" metricMagnitude="-2" metricUnit="m" metricValue="2.76" unit="mm" value="27.6">27.6 mm</quantity>
, NCHUZOOL 15228,
<materialsCitation id="07326D6294BC142FBB9BB563B59E9EF3" collectingDate="2023-04-12" country="Taiwan" location="Guangfu" municipality="Jili Lake" specimenCount="1" specimenCount-male="1" stateProvince="Hualien">
<specimenCount id="26606D7EABCD7ACF1FA410E250D9BFD7" type="male">1 male</specimenCount>
, 33.1
<normalizedToken id="080166B688BCC048000B38251BDE6A04" originalValue="×">x</normalizedToken>
<quantity id="65BC210C858E7974516982E3F5326EFF" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" unit="mm" value="29.7">29.7 mm</quantity>
, NCHUZOOL 15229,
<collectingMunicipality id="877CF62AA053129E2FDDA03C68A79C64">Jili Lake</collectingMunicipality>
,
<location id="42D363A3CAB4778CE81AB3F126F861BB" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:42D363A3CAB4778CE81AB3F126F861BB" country="Taiwan" municipality="Jili Lake" name="Guangfu" stateProvince="Hualien">Guangfu</location>
,
<collectingRegion id="B32CC18FAAA1373AB699B98BB9DD81DB" country="Taiwan" name="Hualien">Hualien</collectingRegion>
, coll.
<normalizedToken id="4C24856F524EA43AD3DAB355F67C7184" originalValue="HTSs">HTS's</normalizedToken>
lab students,
<collectingDate id="F5A1E78524B35EA07254436F8C3A1004" value="2023-04-12">12 Apr. 2023</collectingDate>
</materialsCitation>
;
<materialsCitation id="AB6764B9B5AC47A36C5DA9CB63D67880" collectingDate="2023-04-11" country="Taiwan" location="Fuli" municipality="Luoshan" specimenCount="1" specimenCount-female="1" stateProvince="Hualien">
<specimenCount id="7596C6FFD3116E45485D97AB3F98C8E2" type="female">1 female</specimenCount>
, 37.7
<normalizedToken id="F7D33D83465025EB0EF36D11B3F99B6D" originalValue="×">x</normalizedToken>
<quantity id="F92357149817CB0F5F530770838FF6F6" metricMagnitude="-2" metricUnit="m" metricValue="3.36" unit="mm" value="33.6">33.6 mm</quantity>
, NCHUZOOL 15230,
<collectingMunicipality id="47EA37F765AE98FD1B13A00C510B81CD">Luoshan</collectingMunicipality>
,
<location id="F279CD912E3B3A18CAD12B833254A406" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:F279CD912E3B3A18CAD12B833254A406" country="Taiwan" municipality="Luoshan" name="Fuli" stateProvince="Hualien">Fuli</location>
,
<collectingRegion id="51FE4FA721AB5EC546D3BF0A6DF291A1" country="Taiwan" name="Hualien">Hualien</collectingRegion>
, coll.
<normalizedToken id="81033990D8ECABCB5862B599AB4A6F1E" originalValue="HTSs">HTS's</normalizedToken>
lab students,
<collectingDate id="6E5CC32C145D431D6B4BBF037B1189CB" value="2023-04-11">11 Apr. 2023</collectingDate>
</materialsCitation>
;
<specimenCount id="2B5F9C5BC4AD5CF8EAA418C50FA5A9E9" type="male">2 males</specimenCount>
, 35.9
<normalizedToken id="679377570CEDA85BF926298A7643B558" originalValue="×">x</normalizedToken>
30.6, 37.0
<normalizedToken id="1F50F7782E062F7780689928681D58B3" originalValue="×">x</normalizedToken>
<quantity id="01B2C355136BA43ED3FDC3D9E48332BC" metricMagnitude="-2" metricUnit="m" metricValue="3.19" unit="mm" value="31.9">31.9 mm</quantity>
,
<materialsCitation id="D405A391F0871058BFAA737DE8A56594" collectingDate="2001-06-18" collectorName="M. - Y. Liu" country="Taiwan" location="Jingpu" specimenCount="2" specimenCount-female="2" stateProvince="Hualien">
<specimenCount id="12A67221DB48C1EBB866A1D649AB5EF0" type="female">2 females</specimenCount>
, 27.6
<normalizedToken id="44FC0B1211F227311F223F199EF3B360" originalValue="×">x</normalizedToken>
23.5, 36.7
<normalizedToken id="3FFD8305060572C72B5EC491557ABFEC" originalValue="×">x</normalizedToken>
<quantity id="0AD682D7F1F5A21F8D1562F33FB2E41F" metricMagnitude="-2" metricUnit="m" metricValue="3.19" unit="mm" value="31.9">31.9 mm</quantity>
, RUMF-ZC-56,
<location id="CA19BCA39E5DC4C487B4F73A6563EAD4" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:CA19BCA39E5DC4C487B4F73A6563EAD4" country="Taiwan" name="Jingpu" stateProvince="Hualien">Jingpu</location>
,
<collectingRegion id="E4D1026F1291E4887048CCFE46ECA801" country="Taiwan" name="Hualien">Hualien</collectingRegion>
, coll.
<collectorName id="D4983A1E9C47D83999B8644277EFDE88">M.-Y. Liu</collectorName>
,
<collectingDate id="93E00F7D5003ACA380BB3ADFFD8010AE" value="2001-06-18">18 Jun. 2001</collectingDate>
</materialsCitation>
;
<materialsCitation id="B2B0E82E8BF09E09AC031E998BEFE33C" collectingDate="2001-01" collectorName="M. - Y. Liu" country="Taiwan" location="Jingpu" specimenCount="2" specimenCount-male="2" stateProvince="Hualien">
<specimenCount id="4E4BB5746355F1F46F7467336FB0D8F9" type="male">2 males</specimenCount>
, 33.0
<normalizedToken id="5970FAD8E805F68E5681AAEBC76AE6F8" originalValue="×">x</normalizedToken>
28.9, 30.4
<normalizedToken id="E0A6F78F2A709BDF6535D532104DB00A" originalValue="×">x</normalizedToken>
<quantity id="C3C26E41211B8BBD2569608C7DAF1C5C" metricMagnitude="-2" metricUnit="m" metricValue="2.65" unit="mm" value="26.5">26.5 mm</quantity>
, NSMT-Cr 15170,
<location id="A42879FC1E979821D540C3A2561C49FE" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:A42879FC1E979821D540C3A2561C49FE" country="Taiwan" name="Jingpu" stateProvince="Hualien">Jingpu</location>
,
<collectingRegion id="02742052BEDAF1509C21D1D2EF2BF7FC" country="Taiwan" name="Hualien">Hualien</collectingRegion>
, coll.
<collectorName id="CD213EBEB2B05406AF0B052B9C8127B6">M.-Y. Liu</collectorName>
,
<collectingDate id="D7CC17835727829CB7468976BDC9DD1A" value="2001-01">Jan. 2001</collectingDate>
</materialsCitation>
;
<materialsCitation id="8D7B4FE7A2E72A5E870C99DD5A9AFD5F" collectingDate="2002-03-06" collectorName="N. - H. Jang-Liaw" country="Taiwan" location="Chenggong" specimenCount="1" specimenCount-male="1" stateProvince="Taitung">
<specimenCount id="A5A475E1B17390CA291F63691A220FF0" type="male">1 male</specimenCount>
, 31.7
<normalizedToken id="2C6898E3AD738C58A2FC1622E9B47005" originalValue="×">x</normalizedToken>
<quantity id="AF05085899C44894A75BA830D31B3139" metricMagnitude="-2" metricUnit="m" metricValue="2.67" unit="mm" value="26.7">26.7 mm</quantity>
, NCHUZOOL 15221,
<location id="66A541A5FF520D3F15C4CB1E0DE5F539" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:66A541A5FF520D3F15C4CB1E0DE5F539" country="Taiwan" name="Chenggong" stateProvince="Taitung">Chenggong</location>
,
<collectingRegion id="57C40246EE403367ED7ED961101E5DCE" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="558E498D9F3F62213ED6AD1EB6770F48">N.-H. Jang-Liaw</collectorName>
,
<collectingDate id="AB905D6AF4CD53F2331259DE67DB5771" value="2002-03-06">6 Mar. 2002</collectingDate>
</materialsCitation>
;
<specimenCount id="4CE5CC6B8B85B011C039EBA259B9164A" type="male">3 males</specimenCount>
, 28.2
<normalizedToken id="277629AE674FB405DE78D995130D2BDB" originalValue="×">x</normalizedToken>
23.3-34.0
<normalizedToken id="9E17F281CFDE3A3F76E53A7B7A904A7D" originalValue="×">x</normalizedToken>
<quantity id="9FF0CF4F79DF98FEF46EA755D3E23CDD" metricMagnitude="-2" metricUnit="m" metricValue="2.88" unit="mm" value="28.8">28.8 mm</quantity>
,
<materialsCitation id="A1CC36B30134FB4387DA2D7B2BB0EB27" collectingDate="1997-08-22" collectorName="H. - C. Liu" country="Taiwan" location="Taimali" specimenCount="2" specimenCount-female="2" stateProvince="Taitung">
<specimenCount id="870CBFA0A61662D9B97D02CB5F7ACFFF" type="female">2 females</specimenCount>
, 16.4
<normalizedToken id="B1C54FABE46E095EE66418B0F3FA5D3C" originalValue="×">x</normalizedToken>
13.6, 38.3
<normalizedToken id="AF3B32346247AD7F400E0C67671C39FF" originalValue="×">x</normalizedToken>
31.0 mm, NCHUZOOL 12931,
<location id="3E17FB00BC060E870F05DB487C152471" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:3E17FB00BC060E870F05DB487C152471" country="Taiwan" name="Taimali" stateProvince="Taitung">Taimali</location>
,
<collectingRegion id="6008027A1B6153A1B0DE516E36203110" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="01F96D83BB1AC9F89C24EB1CF2615173">H.-C. Liu</collectorName>
,
<collectingDate id="2C6A95C04ACE875F23DC0AE79FAC92E0" value="1997-08-22">22 Aug. 1997</collectingDate>
</materialsCitation>
;
<specimenCount id="E611EAB9D0ED9075B5E9CF9A9945B2E0" type="male">4 males</specimenCount>
, 28.6
<normalizedToken id="026AE18E8767CAFE4BE76703E569DF16" originalValue="×">x</normalizedToken>
24.8-33.7
<normalizedToken id="2B2E09F742672D70462DCB7903081C7E" originalValue="×">x</normalizedToken>
<quantity id="CD41E2AC5A954D465824DEDE29EEDC1C" metricMagnitude="-2" metricUnit="m" metricValue="2.91" unit="mm" value="29.1">29.1 mm</quantity>
,
<materialsCitation id="5D18FC6175BB4E213B3E707400598B03" collectingDate="1997-09-05" collectorName="H. - C. Liu, H. - L. Hsu" country="Taiwan" location="Chenggong" municipality="Sansiantai" specimenCount="1" specimenCount-female="1" stateProvince="Taitung">
<specimenCount id="FD6B2BFA2EE699B1399B3BD006AA0265" type="female">1 female</specimenCount>
, 32.5
<normalizedToken id="64BDE1DA3B8BF830CA9E4AA82522CAE8" originalValue="×">x</normalizedToken>
27.0 mm, NCHUZOOL 15206,
<collectingMunicipality id="1FFC8B739297C5484890DD80B600DD4C">Sansiantai</collectingMunicipality>
,
<location id="C2B6DAA4E7D8C53360D28CF14D441491" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:C2B6DAA4E7D8C53360D28CF14D441491" country="Taiwan" municipality="Sansiantai" name="Chenggong" stateProvince="Taitung">Chenggong</location>
,
<collectingRegion id="DB4B659C5862E3FF3C6AFED96AFA3AA7" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="CDF807B762D854498ACC41691368FF69">H.-C. Liu</collectorName>
and
<collectorName id="B2962F80DBA546C3C4FE31EE0526CB77">H.-L. Hsu</collectorName>
,
<collectingDate id="8C722C37EEFA1468598B0095BE4E56D9" value="1997-09-05">5 Sep. 1997</collectingDate>
</materialsCitation>
;
<materialsCitation id="5AC0E1B6D03A17205640EAEFDA323DAE" collectingDate="2001-08-22" collectorName="S. - P. Wu" country="Taiwan" elevation="1250" location="Beinan" municipality="Lijia" specimenCount="2" specimenCount-male="2" stateProvince="Taitung">
<specimenCount id="122624E57E68F3BF7D2C6D6487064D7C" type="male">2 males</specimenCount>
, 31.4
<normalizedToken id="292176A58A63D1D21FAB5A4334606CE2" originalValue="×">x</normalizedToken>
25.9, 27.0
<normalizedToken id="D94EFFE13C9BFC08E8BC6F3447DC72CF" originalValue="×">x</normalizedToken>
<quantity id="E7E2A5C837F0F80A978BDA0277B1E324" metricMagnitude="-2" metricUnit="m" metricValue="2.33" unit="mm" value="23.3">23.3 mm</quantity>
, NCHUZOOL 12933,
<collectingMunicipality id="A39B245F404C84566C3104C0F86D8759">Lijia</collectingMunicipality>
,
<location id="D9B07C6BE16916CF9A7ABF5EF722E4A4" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:D9B07C6BE16916CF9A7ABF5EF722E4A4" country="Taiwan" municipality="Lijia" name="Beinan" stateProvince="Taitung">Beinan</location>
,
<collectingRegion id="993553960E37DF7F76360281F93A8D3B" country="Taiwan" name="Taitung">Taitung</collectingRegion>
,
<quantity id="F3D34523401C2118D22A530CC44C0CBD" metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">
<elevation id="69218F1CBF9AFDFE67CEBAB0D5035BCA" metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">1250 m</elevation>
</quantity>
a.s.l, coll.
<collectorName id="8134495BED982D73A2E21F884022D2D0">S.-P. Wu</collectorName>
,
<collectingDate id="6BF6D2FF5B4459EBAA80F3EC4AD52EB6" value="2001-08-22">22 Aug. 2001</collectingDate>
</materialsCitation>
;
<materialsCitation id="376E1F427F188A91E0DAD98EDF40536C" collectingDate="2001-01-10" collectorName="H. - T. Shih" country="Taiwan" location="Yanping" municipality="Luanshan" specimenCount="1" specimenCount-male="1" stateProvince="Taitung">
<specimenCount id="B32C64F20916346B12B07C74472190F6" type="male">1 male</specimenCount>
, 28.4
<normalizedToken id="998D6D840BC629DD41EE7E9169F4FEBA" originalValue="×">x</normalizedToken>
24.6, NCHUZOOL 12938, 32.9
<normalizedToken id="E15644C6C1518E386AF62A8F8DC5F112" originalValue="×">x</normalizedToken>
<quantity id="9875CB11C993CE14B434388E0C4C3C8E" metricMagnitude="-2" metricUnit="m" metricValue="2.87" unit="mm" value="28.7">28.7 mm</quantity>
, ZRC 2003.0036,
<collectingMunicipality id="990655FBBB18210C6FDA1B14AE47E08C">Luanshan</collectingMunicipality>
,
<location id="981533265ABCC70BF78C528A738081BD" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:981533265ABCC70BF78C528A738081BD" country="Taiwan" municipality="Luanshan" name="Yanping" stateProvince="Taitung">Yanping</location>
,
<collectingRegion id="B6390ABCB25C72D8F0F786E817F0632B" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="3EA9351B1157A8FEAAC9B31658653D86">H.-T. Shih</collectorName>
,
<collectingDate id="CA47B4748C7D738CE98311FCEA7918BE" value="2001-01-10">10 Jan. 2001</collectingDate>
</materialsCitation>
;
<specimenCount id="2B6DADDE4A912B6762B6831641BF4597" type="male">4 males</specimenCount>
, 29.0
<normalizedToken id="54DC44F8D0E10D1D7215EB0D29423BE6" originalValue="×">x</normalizedToken>
24.6-35.6
<normalizedToken id="3DD43C1FA2C8B225C34B13A0AD1CF634" originalValue="×">x</normalizedToken>
<quantity id="3CD3BE8B80D1CE6ACD2E37E6372EC91D" metricMagnitude="-2" metricUnit="m" metricValue="2.93" unit="mm" value="29.3">29.3 mm</quantity>
,
<materialsCitation id="D7D1AD1A2CEBCD60C9E858BE109CF8C8" collectingDate="1995-04-22" collectorName="H. - C. Liu" country="Taiwan" location="Daren" municipality="Taiban" specimenCount="1" specimenCount-female="1" stateProvince="Taitung">
<specimenCount id="6F11DEC1BF6E8CC4690AE67EFBE7CCEB" type="female">1 female</specimenCount>
, 21.9
<normalizedToken id="6043E956E16C8F081B24333F41036D4F" originalValue="×">x</normalizedToken>
<quantity id="8B9B1D129776987259736C3FBF396676" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" unit="mm" value="18.4">18.4 mm</quantity>
, NCHUZOOL 15218,
<collectingMunicipality id="0225BEDF65467FF4A2A134AA3CAD1A07">Taiban</collectingMunicipality>
,
<location id="CE166B1E89A5B6B2DEECFB09B174C5D4" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:CE166B1E89A5B6B2DEECFB09B174C5D4" country="Taiwan" municipality="Taiban" name="Daren" stateProvince="Taitung">Daren</location>
,
<collectingRegion id="9818169E70112E5072C9F200F373D12E" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="BE51BBE04C493E1F61E038F6DC73CE79">H.-C. Liu</collectorName>
,
<collectingDate id="16A36F181ECD2F1AC9024E90540690A5" value="1995-04-22">22 Apr. 1995</collectingDate>
</materialsCitation>
;
<materialsCitation id="2E5C07708E67FCFC778A0A3746466B1D" collectingDate="1997-08-22" collectorName="H. - C. Liu, C. - H. Wang" country="Taiwan" location="Dawu" specimenCount="1" specimenCount-female="1" stateProvince="Taitung">
<specimenCount id="7BDBB97E96A860E5A40F830342D2245F" type="female">1 female</specimenCount>
, 44.7
<normalizedToken id="F760C938DDA8ED4B1FD9EBE9BFE87E8B" originalValue="×">x</normalizedToken>
<quantity id="E9F5DF065FD6ADCB7039D368CE4BB6B1" metricMagnitude="-2" metricUnit="m" metricValue="3.59" unit="mm" value="35.9">35.9 mm</quantity>
, NCHUZOOL 15210,
<location id="194B32ACB941FB2FCEE27035777355B4" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:194B32ACB941FB2FCEE27035777355B4" country="Taiwan" name="Dawu" stateProvince="Taitung">Dawu</location>
,
<collectingRegion id="CE077A1A8B90F0E1C4F7C0C6A536110C" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="A76C3B5B9A159BE88F62443FF85FDB74">H.-C. Liu</collectorName>
and
<collectorName id="6D79411D8645AC642ECB8809C0365177">C.-H. Wang</collectorName>
,
<collectingDate id="BF63CDF3BAE5050AEDE1B854A7FFA135" value="1997-08-22">22 Aug. 1997</collectingDate>
</materialsCitation>
;
<specimenCount id="2466E4463140E685DCEA36FEB6C88245" type="male">1 male</specimenCount>
, 28.9
<normalizedToken id="5DCD2ED8C7914E60CEAA0AFBED48D79B" originalValue="×">x</normalizedToken>
<quantity id="03581BC6496D6D49530934286FAD39C4" metricMagnitude="-2" metricUnit="m" metricValue="2.41" unit="mm" value="24.1">24.1 mm</quantity>
,
<materialsCitation id="5C1636BEE7B8A2F7E93DE02517C5F940" collectingDate="2001-01-09" collectorName="H. - T. Shih" country="Taiwan" location="Dawu" specimenCount="1" specimenCount-female="1" stateProvince="Taitung">
<specimenCount id="E4F73F2C74CED39C49D6228F5B5384BE" type="female">1 female</specimenCount>
, 29.8
<normalizedToken id="43B71A000A360DD5FE935D0041B9585A" originalValue="×">x</normalizedToken>
25.0 mm, NCHUZOOL 12943,
<location id="F7EBFCDD44B2E2DE8B245A3682F0ECA2" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:F7EBFCDD44B2E2DE8B245A3682F0ECA2" country="Taiwan" name="Dawu" stateProvince="Taitung">Dawu</location>
,
<collectingRegion id="78036A5BF6B1D55DDEEE848A31E76431" country="Taiwan" name="Taitung">Taitung</collectingRegion>
, coll.
<collectorName id="61E3221012418483A7B13B449A2F66A6">H.-T. Shih</collectorName>
,
<collectingDate id="58B47E00173A9D51559F55090C8389A2" value="2001-01-09">9 Jan. 2001</collectingDate>
</materialsCitation>
;
<specimenCount id="AFE5E41DF72DEA348F63D4B73A95E6B3" type="male">2 males</specimenCount>
, 40.1
<normalizedToken id="5CA3D1C19232995274D7722E7A27B4EB" originalValue="×">x</normalizedToken>
33.6, 42.9
<normalizedToken id="1DF66896282B1A896AF64AED4A88A36C" originalValue="×">x</normalizedToken>
<quantity id="CC0A5190A3BDF0F68ED333EEBC185256" metricMagnitude="-2" metricUnit="m" metricValue="3.62" unit="mm" value="36.2">36.2 mm</quantity>
,
<materialsCitation id="DFC73D9FBC113307743CE70D79DDC3F0" collectingDate="1999-03-12" collectorName="H. - T. Shih" country="Taiwan" location="Shihzih" municipality="Danlu" specimenCount="1" specimenCount-female="1" stateProvince="Pingtung">
<specimenCount id="96FDEDA5F294A6895071886966B1435D" type="female">1 female</specimenCount>
, 38.4
<normalizedToken id="7E7DF59651991F5FF10AAE0931FE549A" originalValue="×">x</normalizedToken>
<quantity id="43004AF0E67A1711B9FE7408926E5985" metricMagnitude="-2" metricUnit="m" metricValue="3.15" unit="mm" value="31.5">31.5 mm</quantity>
, NCHUZOOL 15207,
<collectingMunicipality id="8E41C884B9D81C307BEFE449839622FC">Danlu</collectingMunicipality>
,
<location id="FAA7063BC17B08257DE932F19E358725" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:FAA7063BC17B08257DE932F19E358725" country="Taiwan" municipality="Danlu" name="Shihzih" stateProvince="Pingtung">Shihzih</location>
,
<collectingRegion id="001B6ED027F72A3ED9C3D40B924411F8" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="4E40F294E5E27CD5721AF565B05BFE06">H.-T. Shih</collectorName>
,
<collectingDate id="9361CF4A21EE823D07177EB3910EDF36" value="1999-03-12">12 Mar. 1999</collectingDate>
</materialsCitation>
;
<materialsCitation id="5910EAAF8CAF7688979F7D5DD6D48AD5" collectingDate="2000-03-05" collectorName="H. - T. Shih" country="Taiwan" location="Hengchun" municipality="Wangsha" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="D64C6F5CC2BA63FD5026866A497C4CFA" type="male">1 male</specimenCount>
, 28.0
<normalizedToken id="A6887C7B5B6C44EA4AACBC6F739DB337" originalValue="×">x</normalizedToken>
<quantity id="CB329A277F1A1D09DE2DB0AF84D947CA" metricMagnitude="-2" metricUnit="m" metricValue="2.33" unit="mm" value="23.3">23.3 mm</quantity>
, NCHUZOOL 15208,
<collectingMunicipality id="CBAC2C3C53AE807E42D3887EA37C020A">Wangsha</collectingMunicipality>
,
<location id="8DA2F77DA4E2A2C1FBE3BDC20324691A" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:8DA2F77DA4E2A2C1FBE3BDC20324691A" country="Taiwan" municipality="Wangsha" name="Hengchun" stateProvince="Pingtung">Hengchun</location>
,
<collectingRegion id="BFBC785FC60C05619CF164B72076D1ED" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="9509626551D540BA832FEC4BAEED2D08">H.-T. Shih</collectorName>
,
<collectingDate id="0384E6CE21408313003D34DCEBB1B848" value="2000-03-05">5 Mar. 2000</collectingDate>
</materialsCitation>
;
<materialsCitation id="62ABBDA9077510D3B8DA337A2C893EE2" collectingDate="1999-03-31" collectorName="H. - T. Shih" country="Taiwan" location="Hengchun" municipality="Wangsha" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="0C2EE0E864DBB6B4A438BB2C3FFEFFB6" type="male">1 male</specimenCount>
, 31.0
<normalizedToken id="275134844B702A073FCC309DA450E9FF" originalValue="×">x</normalizedToken>
<quantity id="2BA6A8F44D2943CE079C069E1F534D7E" metricMagnitude="-2" metricUnit="m" metricValue="2.52" unit="mm" value="25.2">25.2 mm</quantity>
, RUMF-ZC-57,
<collectingMunicipality id="099B08129D63AF6D982C59922C08392E">Wangsha</collectingMunicipality>
,
<location id="958B40C191D097D3D8CA3F5A3F6876EC" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:958B40C191D097D3D8CA3F5A3F6876EC" country="Taiwan" municipality="Wangsha" name="Hengchun" stateProvince="Pingtung">Hengchun</location>
,
<collectingRegion id="A246C84B230D54D1E04DEAD180DB067C" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="E54E5F72AACB6C7DEA3172652BBB6141">H.-T. Shih</collectorName>
,
<collectingDate id="2614A459B98DF084767C5753966659AB" value="1999-03-31">31 Mar. 1999</collectingDate>
</materialsCitation>
;
<specimenCount id="59AA9AEB15C0DFEE19429204F470C5E6" type="male">2 males</specimenCount>
, 41.7
<normalizedToken id="9FCE1C697687F6AB5E289A231399FC82" originalValue="×">x</normalizedToken>
35.0, 41.9
<normalizedToken id="AC2A8FEC630E95355347CFD39229D336" originalValue="×">x</normalizedToken>
35.0 mm,
<materialsCitation id="B7068B1C86836016CB257EF2F320F882" collectingDate="1996-10-19" collectorName="H. - T. Shih" country="Taiwan" location="Manjhou" municipality="Nanrenshan" specimenCount="1" specimenCount-female="1" stateProvince="Pingtung">
<specimenCount id="BD55EB45B212336AB60A3159C6BA371C" type="female">1 female</specimenCount>
, 35.7
<normalizedToken id="9C73E1CB70115359DA2EC58EBE83F78E" originalValue="×">x</normalizedToken>
<quantity id="24C6E98F07EECD0B073BB61291C7D19F" metricMagnitude="-2" metricUnit="m" metricValue="2.98" unit="mm" value="29.8">29.8 mm</quantity>
, NCHUZOOL 15217,
<collectingMunicipality id="5276531B02DD8CCA331E3B62FD6C42CE">Nanrenshan</collectingMunicipality>
,
<location id="D6229566C3F16DB495D7407F7082BF87" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:D6229566C3F16DB495D7407F7082BF87" country="Taiwan" municipality="Nanrenshan" name="Manjhou" stateProvince="Pingtung">Manjhou</location>
,
<collectingRegion id="F5784DA0D4446659F23AD13E427A88AD" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="B69B82CD1A089C6B4274089B3773805B">H.-T. Shih</collectorName>
,
<collectingDate id="2BAEE5EDD1574D5ACAA94A023081140F" value="1996-10-19">19 Oct. 1996</collectingDate>
</materialsCitation>
;
<materialsCitation id="7D094CE4234D8E0B2CF835949CA8B142" collectingDate="2003-02-08" collectorName="R. - H. Lee" country="Taiwan" location="Manjhou" municipality="Nanrenshan" specimenCount="3" specimenCount-male="3" stateProvince="Pingtung">
<specimenCount id="3A62B48BEEADFC6C3DDB45B2D8A617B5" type="male">3 males</specimenCount>
, 37.0
<normalizedToken id="B645D308731D00743329ECC30D2660ED" originalValue="×">x</normalizedToken>
30.8-40.6
<normalizedToken id="B47F1EDC1FD1C756BF5C490B36FB1FC3" originalValue="×">x</normalizedToken>
<quantity id="10BDCBD0C14941162FF2F44A1551B1CD" metricMagnitude="-2" metricUnit="m" metricValue="3.47" unit="mm" value="34.7">34.7 mm</quantity>
, NCHUZOOL 15204,
<collectingMunicipality id="9252CB8A291E02F8A7CA8BC99E015E76">Nanrenshan</collectingMunicipality>
,
<location id="EC4FC07A030F8AE0E615D6579F7789B1" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:EC4FC07A030F8AE0E615D6579F7789B1" country="Taiwan" municipality="Nanrenshan" name="Manjhou" stateProvince="Pingtung">Manjhou</location>
,
<collectingRegion id="5F797615EC19BF92ACAA542F00A969F4" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="CB13D9C70153AEAFDAD4EA8BE3602309">R.-H. Lee</collectorName>
,
<collectingDate id="CE1EC0663CD145D704AA13C7CD7A7B00" value="2003-02-08">8 Feb. 2003</collectingDate>
</materialsCitation>
;
<materialsCitation id="AD25FF9840550DEAB1E0F5F265B96177" collectingDate="1999-02-24" collectorName="H. - T. Shih" country="Taiwan" location="Manjhou" municipality="Nanrenshan" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="44858293D7B6F132AD66A3C1004182D7" type="male">1 male</specimenCount>
, 44.3
<normalizedToken id="CF4D38C2958CF1E425066C7D96036830" originalValue="×">x</normalizedToken>
37.0 mm, ZRC 2003.0037,
<collectingMunicipality id="AF69065C577167917470015F13B68D73">Nanrenshan</collectingMunicipality>
,
<location id="1C1E627ABC398A63836D421C939CBE3E" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:1C1E627ABC398A63836D421C939CBE3E" country="Taiwan" municipality="Nanrenshan" name="Manjhou" stateProvince="Pingtung">Manjhou</location>
,
<collectingRegion id="61894DB47D72C89826740C07C48EFD34" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="ACC05C1D6F3470EFD790763908CBB3FC">H.-T. Shih</collectorName>
,
<collectingDate id="DD63BB6BF3E0C0B34646A46A0780E379" value="1999-02-24">24 Feb. 1999</collectingDate>
</materialsCitation>
;
<materialsCitation id="2D15C1637F393C343C978DAAB6CC37A8" collectingDate="2000-01-06" collectorName="H. - T. Shih" country="Taiwan" location="Hengchun" municipality="Kenting" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
<specimenCount id="78DC7C7A46978BEFF0D32BD6F82BB516" type="male">1 male</specimenCount>
, 43.8
<normalizedToken id="D60EC86DE0C17D0D549AA1A23D7279E4" originalValue="×">x</normalizedToken>
<quantity id="33F5B7C69AE055ADBAC79EF4194A4ABD" metricMagnitude="-2" metricUnit="m" metricValue="3.67" unit="mm" value="36.7">36.7 mm</quantity>
, RUMF-ZC-58,
<collectingMunicipality id="DF3A0FB1CE6F9B58E288012734923B02">Kenting</collectingMunicipality>
,
<location id="C8EBDF37F242B6FF9D281F2F1DE7A15D" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:C8EBDF37F242B6FF9D281F2F1DE7A15D" country="Taiwan" municipality="Kenting" name="Hengchun" stateProvince="Pingtung">Hengchun</location>
,
<collectingRegion id="E5A92EC418BD3BA1FA0A8017C27CAC78" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="5B13A05D608432AED3A91017DEE77BE0">H.-T. Shih</collectorName>
,
<collectingDate id="404E1CE4569AA4AB67C0901241A22439" value="2000-01-06">6 Jan. 2000</collectingDate>
</materialsCitation>
;
<specimenCount id="E302A286283CB0927CE5834CA2C4C232" type="male">1 male</specimenCount>
, 30.6
<normalizedToken id="CBDC65A396E0E3F6E221A32A47C5273F" originalValue="×">x</normalizedToken>
<quantity id="39EF75418BEDFDD388B6EF276222C480" metricMagnitude="-2" metricUnit="m" metricValue="2.62" unit="mm" value="26.2">26.2 mm</quantity>
,
<materialsCitation id="A7E066B52C2DB8CA2EA2F89E8D1A5059" collectingDate="1999-03-10" collectorName="Sihchong R., H. - T. Shih" country="Taiwan" location="Mudan" specimenCount="1" specimenCount-female="1" stateProvince="Pingtung">
<specimenCount id="ECF154BE7E26DF0408656AE51A849C99" type="female">1 female</specimenCount>
, 35.2
<normalizedToken id="06CD9442F4E2D16E698463BCEA7B49C1" originalValue="×">x</normalizedToken>
<quantity id="DA121613C75323552C799BC7C9465A63" metricMagnitude="-2" metricUnit="m" metricValue="2.93" unit="mm" value="29.3">29.3 mm</quantity>
, NSMT-Cr 15171,
<collectorName id="8CBCCCD9828EE61D24E0808F29DFA3DA">Sihchong R.</collectorName>
,
<location id="CD620CC529DA3E65A0D4EC145B4578EF" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:CD620CC529DA3E65A0D4EC145B4578EF" country="Taiwan" name="Mudan" stateProvince="Pingtung">Mudan</location>
,
<collectingRegion id="14BC31B4E44C841F6D9DEB1C546F9D08" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="9D9C9F62CC59F574C6A7647C2C55E35C">H.-T. Shih</collectorName>
,
<collectingDate id="DBFAD678ADA77E1ADC2C38E3826EE64F" value="1999-03-10">10 Mar. 1999</collectingDate>
</materialsCitation>
;
<specimenCount id="9F6A352DFE3AD18D5E35B32896D939A1" type="male">1 male</specimenCount>
, 38.7
<normalizedToken id="092606ED11D076EA1FB292945C764665" originalValue="×">x</normalizedToken>
<quantity id="C726AF02EACA4AD481A8357B3A7ADA43" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" unit="mm" value="32.4">32.4 mm</quantity>
,
<materialsCitation id="1B5DB160A4F491CAD93282C46B52EB44" collectingDate="1997-07-22" collectorName="H. - C. Liu" country="Taiwan" location="Shihzih" municipality="Nanshih" specimenCount="3" specimenCount-female="3" stateProvince="Pingtung">
<specimenCount id="080117131651CF9223BA1226E26C49B3" type="female">3 females</specimenCount>
, 38.3
<normalizedToken id="2B1DBD26C7E820B545C8BBBD8136E073" originalValue="×">x</normalizedToken>
33.5-43.6
<normalizedToken id="B2DEA7D8F0343F10EE9F1D179873FDD5" originalValue="×">x</normalizedToken>
<quantity id="DB9AE9BBA146EB6699CCE898E07E63AB" metricMagnitude="-2" metricUnit="m" metricValue="3.59" unit="mm" value="35.9">35.9 mm</quantity>
, NCHUZOOL 15209,
<collectingMunicipality id="692A908E2A47919E3B5A257BACDAF9CA">Nanshih</collectingMunicipality>
,
<location id="23D475147B322B807EE89EBD2118556C" LSID="urn:lsid:plazi:treatment:FB5DB50C96815662AFBAECC7F366EA46:23D475147B322B807EE89EBD2118556C" country="Taiwan" municipality="Nanshih" name="Shihzih" stateProvince="Pingtung">Shihzih</location>
,
<collectingRegion id="19ED0311CAE63826642BF59C400AE7CF" country="Taiwan" name="Pingtung">Pingtung</collectingRegion>
, coll.
<collectorName id="B0F297F7B73A33B733DD2771213C7417">H.-C. Liu</collectorName>
,
<collectingDate id="34A4B5DFA51E880A0EA4119FB55D708D" value="1997-07-22">22 Jul. 1997</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5299919EC10886424DAFA838106071D8" pageId="0" pageNumber="169" type="diagnosis">
<paragraph id="059FFFE8F0FD60739AD5AFFC84664081" pageId="0" pageNumber="169">Diagnosis.</paragraph>
<paragraph id="6B4ADE444F6FAC3A67ED5C0E63A8FB4F" pageId="0" pageNumber="169">
Carapace subtrapezoidal, CW 1.14-1.24
<normalizedToken id="27F21D926DB184666A788B86E6C5AC18" originalValue="×">x</normalizedToken>
CL (mean 1.18,
<emphasis id="FBA3073E12639C28E64CBE4B27C33F26" italics="true" pageId="0" pageNumber="169">n</emphasis>
= 45), dorsal surface almost flat, relatively rough, epigastric and postfrontal cristae distinct, both cristae represented by 1-3 oblique lines of granules, lateral end of postfrontal cristae far apart from epibranchial tooth; external orbital and epibranchial teeth distinct, sharp, directed anteriorly. G1 terminal segment proportionally longer and slender, distal opening directed laterally, distal inner edge with keel-like projection in dorsal view, projection proportionally large, directed more distally; G1 subterminal segment curving inwards (in eastern to southeastern Taiwan populations) or straight (in southern Taiwan populations).
</paragraph>
</subSubSection>
<subSubSection id="BEA9B545AEF7C4897577D2BD1021BB42" pageId="0" pageNumber="169" type="etymology">
<paragraph id="F2545B4CE508CCCA2A992C1FF1572C26" pageId="0" pageNumber="169">Etymology.</paragraph>
<paragraph id="4B2B8E9016CF04CA5321E24812E1BD6D" pageId="0" pageNumber="169">
The new species is named after Penglai, the ancient name of Taiwan. It also refers to the fact that the island of Taiwan was formed by the Penglai Orogeny (ca 5 million years ago) (
<bibRefCitation id="DD7CF7AE1290731C97479AF1490666AC" DOI="https://doi.org/10.1016/0040-1951(90)90188-E" author="Teng, LS" journalOrPublisher="Tectonophysics" pageId="0" pageNumber="169" pagination="57 - 76" refId="B97" refString="Teng, LS, 1990. Geotectonic evolution of late Cenozoic arc-continent collision in Taiwan. Tectonophysics 183 (1-4): 57 - 76, DOI: https://doi.org/10.1016/0040-1951(90)90188-E" title="Geotectonic evolution of late Cenozoic arc-continent collision in Taiwan." url="https://doi.org/10.1016/0040-1951(90)90188-E" volume="183" year="1990">Teng 1990</bibRefCitation>
;
<bibRefCitation id="B47E079B5DBDC1868C4D6F6BB5BF6BE2" DOI="https://doi.org/10.1016/S0040-1951(00)00028-7" author="Liu, TK" journalOrPublisher="Tectonophysics" pageId="0" pageNumber="169" pagination="69 - 82" refId="B46" refString="Liu, TK, Chen, YG, Chen, WS, Jiang, SH, 2000. Rates of cooling and denudation of the early Penglai Orogeny, Taiwan, as assessed by fission-track constraints. Tectonophysics 320 (1): 69 - 82, DOI: https://doi.org/10.1016/S0040-1951(00)00028-7" title="Rates of cooling and denudation of the early Penglai Orogeny, Taiwan, as assessed by fission-track constraints." url="https://doi.org/10.1016/S0040-1951(00)00028-7" volume="320" year="2000">Liu et al. 2000</bibRefCitation>
), and the ancestral
<taxonomicName id="E7D78E5496FBC1A3900F2145E5925D16" authorityName="Bott" authorityYear="1967" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB7B2EE2240562D9C2642A40F60612" italics="true" pageId="0" pageNumber="169">Candidiopotamon</emphasis>
</taxonomicName>
most likely colonized Taiwan at that time (
<bibRefCitation id="B97CDF2EED72939FBECD764ED809FBEE" DOI="https://doi.org/10.1111/j.1365-2699.2006.01472.x" author="Shih, HT" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="169" pagination="980 - 989" refId="B86" refString="Shih, HT, Hung, HC, Schubart, CD, Chen, CA, Chang, HW, 2006. Intraspecific genetic diversity of the endemic freshwater crab Candidiopotamon rathbunae (Decapoda, Brachyura, Potamidae) reflects five million years of geological history of Taiwan. Journal of Biogeography 33 (6): 980 - 989, DOI: https://doi.org/10.1111/j.1365-2699.2006.01472.x" title="Intraspecific genetic diversity of the endemic freshwater crab Candidiopotamon rathbunae (Decapoda, Brachyura, Potamidae) reflects five million years of geological history of Taiwan." url="https://doi.org/10.1111/j.1365-2699.2006.01472.x" volume="33" year="2006">Shih et al. 2006</bibRefCitation>
). The species name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="26983C58BF3CDFCDFABAC630BDF653C4" pageId="0" pageNumber="169" type="coloration">
<paragraph id="307C6755FA41DD8B55485BC20E1092D3" pageId="0" pageNumber="169">Coloration</paragraph>
<paragraph id="27DAA3EEBFC34B1C880961A3A1AA6085" pageId="0" pageNumber="169">
<emphasis id="759EDC09139FE0C67A26A869FFCF82CA" bold="true" pageId="0" pageNumber="169">
(Figs
<figureCitation id="718E1969555DCE728511FA9C5E1C7F7D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A, B) and Candidiopotamon penglai sp. nov. (C-F). A, C, E overall dorsal view B, D, F overall ventral view A, B NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) C, D NCHUZOOL 15204 (male, 39.7 x 34.4 mm, Nanrenshan, Pingtung, southern Taiwan); NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure4" httpUri="https://binary.pensoft.net/fig/904110" pageId="0" pageNumber="169">4C-F</figureCitation>
,
<figureCitation id="9A849816510CAA81F2E31BC3FF33AF90" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A-D) and Candidiopotamon penglai sp. nov. (E-H). A, B NCHUZOOL 15194 (male, 35.4 x 31. 2 mm, Tai-an, Miaoli, northwestern Taiwan) C NCHUZOOL 15211 (male, 24.8 x 22.1 mm, Dongshih, Taichung, western Taiwan) D NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) E, F NCHUZOOL 15204 (2 males, 39.7 x 34.4 mm, 40.6 x 34.3 mm, Nanrenshan, Pingtung, southern Taiwan) G NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan) H NCHUZOOL 15229 (male, 33.1 x 29.7 mm, Jili Lake, Guangfu, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure5" httpUri="https://binary.pensoft.net/fig/904111" pageId="0" pageNumber="169">5E-H</figureCitation>
).
</emphasis>
Same as
<taxonomicName id="AE00907741DAAA074542C29A1E1969BA" lsidName="C. rathbuni" pageId="0" pageNumber="169" rank="species" species="rathbuni">
<emphasis id="253BC2BCCCDAB9EDB20B55CECB75DB17" italics="true" pageId="0" pageNumber="169">C. rathbuni</emphasis>
</taxonomicName>
, but young individuals tend to be reddish (e.g. Fig.
<figureCitation id="7610C7DA4BB3292993BE3338B4D20537" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A-D) and Candidiopotamon penglai sp. nov. (E-H). A, B NCHUZOOL 15194 (male, 35.4 x 31. 2 mm, Tai-an, Miaoli, northwestern Taiwan) C NCHUZOOL 15211 (male, 24.8 x 22.1 mm, Dongshih, Taichung, western Taiwan) D NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) E, F NCHUZOOL 15204 (2 males, 39.7 x 34.4 mm, 40.6 x 34.3 mm, Nanrenshan, Pingtung, southern Taiwan) G NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan) H NCHUZOOL 15229 (male, 33.1 x 29.7 mm, Jili Lake, Guangfu, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure5" httpUri="https://binary.pensoft.net/fig/904111" pageId="0" pageNumber="169">5H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4C6AB2C4215F481D76F2C97DE7655F95" pageId="0" pageNumber="169" type="distribution">
<paragraph id="C3EC49F4CFA89A2AE54E67DC903F7C87" pageId="0" pageNumber="169">Distribution</paragraph>
<paragraph id="BAD1BB26351FBCF3623B2E1F8280EFF8" pageId="0" pageNumber="169">
<emphasis id="EE5300DAA4F6B738DB4DF14E0C6388A3" bold="true" pageId="0" pageNumber="169">
(Fig.
<figureCitation id="F3D85802E3A919064918EE4A53DB0D79" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites for Candidiopotamon rathbuni (De Man, 1914) (circles) and Candidiopotamon penglai sp. nov. (squares) in Taiwan, as well as other congeners from the Ryukyus, Japan. Different colors indicate sampling sites of clades. Dotted lines indicate the possible biogeographic boundaries between clades NW, W, SW, S, SE, E 1, and E 2, corresponding to the color in Fig. 2; as well as species. The red star and blue star indicate the type localities of C. rathbuni and C. penglai sp. nov., respectively. Three forms of male left G 1 are shown for clades in C. rathbuni and two forms for clades in C. penglai sp. nov." figureDoi="10.3897/zookeys.1179.106718.figure1" httpUri="https://binary.pensoft.net/fig/904107" pageId="0" pageNumber="169">1</figureCitation>
).
</emphasis>
<taxonomicName id="35702687D4C4840C99C66065B44C9148" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon penglai" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="penglai">
<emphasis id="A1DEABEF5837CBA38CC50874DD7F4DAD" italics="true" pageId="0" pageNumber="169">Candidiopotamon penglai</emphasis>
</taxonomicName>
sp. nov. is distributed along the eastern side of the Central Range of Taiwan (except for the northeastern area), i.e., Hualien and Taitung in eastern Taiwan, as well as Hengchun Peninsula (the southern part of Pingtung and the southern tip of Taitung) in southern Taiwan. The type locality of
<taxonomicName id="3D3C835077E0DC91F5A1F3C6D8732046" lsidName="C. penglai" pageId="0" pageNumber="169" rank="species" species="penglai">
<emphasis id="71C311D3EB4D26C5DA91423F24620F69" italics="true" pageId="0" pageNumber="169">C. penglai</emphasis>
</taxonomicName>
sp. nov. is Ji-an, Hualien (Fig.
<figureCitation id="F8AD708D6E60D378762FBA624A418FFB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites for Candidiopotamon rathbuni (De Man, 1914) (circles) and Candidiopotamon penglai sp. nov. (squares) in Taiwan, as well as other congeners from the Ryukyus, Japan. Different colors indicate sampling sites of clades. Dotted lines indicate the possible biogeographic boundaries between clades NW, W, SW, S, SE, E 1, and E 2, corresponding to the color in Fig. 2; as well as species. The red star and blue star indicate the type localities of C. rathbuni and C. penglai sp. nov., respectively. Three forms of male left G 1 are shown for clades in C. rathbuni and two forms for clades in C. penglai sp. nov." figureDoi="10.3897/zookeys.1179.106718.figure1" httpUri="https://binary.pensoft.net/fig/904107" pageId="0" pageNumber="169">1</figureCitation>
: no. 55).
</paragraph>
</subSubSection>
<subSubSection id="CA62C29AE2008D8C0C7D9173DB1CEBD7" pageId="0" pageNumber="169" type="habitat">
<paragraph id="886CD0DCD121A0246B55DFF7F2776D63" pageId="0" pageNumber="169">Habitat.</paragraph>
<paragraph id="F37C87694E665049D17D1C30A5C69B13" pageId="0" pageNumber="169">
Same as
<taxonomicName id="0485C0CE7E6E4CAA0F8E7D33A84AFD70" lsidName="C. rathbuni" pageId="0" pageNumber="169" rank="species" species="rathbuni">
<emphasis id="A469BBEEE321DC6A79BC3625FF66BB11" italics="true" pageId="0" pageNumber="169">C. rathbuni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1B509BA7455980DBF4635E19608BF6BC" pageId="0" pageNumber="169" type="remarks">
<paragraph id="41CBD8DC5F47B8487037DAC1C290624E" pageId="0" pageNumber="169">Remarks.</paragraph>
<paragraph id="74BD5CD9CDDFF3216AD74B4971390C16" pageId="0" pageNumber="169">
A detailed morphological comparison reveals that specimens of
<taxonomicName id="7909C9535DC0D774E9BEBA36DD4BE15F" lsidName="C. rathbuni" pageId="0" pageNumber="169" rank="species" species="rathbuni">
<emphasis id="731156A163F810DC839F7E36E7460AAF" italics="true" pageId="0" pageNumber="169">C. rathbuni</emphasis>
</taxonomicName>
from western Taiwan and
<taxonomicName id="B858977509D942164E4C6700E3A3F0A4" lsidName="C. penglai" pageId="0" pageNumber="169" rank="species" species="penglai">
<emphasis id="1AC54A36ABE0985A7E3765F6D89992BC" italics="true" pageId="0" pageNumber="169">C. penglai</emphasis>
</taxonomicName>
sp. nov. from eastern Taiwan show differences in G1 structure.
<taxonomicName id="C9E6217978CE32AC80D003A7D06A716A" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon rathbuni" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="rathbuni">
<emphasis id="E9735B45447B9D8617FB41A10A366E85" italics="true" pageId="0" pageNumber="169">Candidiopotamon rathbuni</emphasis>
</taxonomicName>
has a smaller and more laterally directed keel-like projection on the distal inner part of the terminal segment (Fig.
<figureCitation id="47281046CEEE08E8ED358F27F1D9FA53" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Candidiopotamon rathbuni (De Man, 1914): male (44.6 x 36.8 mm) from Chiayi, western Taiwan (RUMF-ZC- 86) (A - E); male (35.4 x 31.2 mm) from Miaoli, northwestern Taiwan (NCHUZOOL 15194) (F 1); male (36.7 x 31.1 mm) from Chiayi, western Taiwan (NCHUZOOL 15205) (F 2); male (36.3 x 30.7 mm) from Taiwu, Pingtung, southwestern Taiwan (NCHUZOOL 15199) (F 3). A carapace, dorsal view B carapace, frontal view C male major chela, outer view D 1 left G 1, ventral view D 2 left G 1, dorsal view D 3 left G 1 terminal segment, dorsal view E left G 2 F 1 - 3 left G 1 s terminal segment, dorsal view. Scale bars: 10 mm (A - C); 2 mm (D - F). Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure3" httpUri="https://binary.pensoft.net/fig/904109" pageId="0" pageNumber="169">3D3</figureCitation>
) [vs larger and more distally directed structure in
<taxonomicName id="7B38CE567A091B3CFDCA8E1114BF1A29" lsidName="C. penglai" pageId="0" pageNumber="169" rank="species" species="penglai">
<emphasis id="D80CA4B9DD43C101C0C25929A4CB68BB" italics="true" pageId="0" pageNumber="169">C. penglai</emphasis>
</taxonomicName>
(Figs
<figureCitation id="379B827B143A918C21B63A7500BDD5BC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A-D) and Candidiopotamon penglai sp. nov. (E-H). A, B NCHUZOOL 15194 (male, 35.4 x 31. 2 mm, Tai-an, Miaoli, northwestern Taiwan) C NCHUZOOL 15211 (male, 24.8 x 22.1 mm, Dongshih, Taichung, western Taiwan) D NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) E, F NCHUZOOL 15204 (2 males, 39.7 x 34.4 mm, 40.6 x 34.3 mm, Nanrenshan, Pingtung, southern Taiwan) G NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan) H NCHUZOOL 15229 (male, 33.1 x 29.7 mm, Jili Lake, Guangfu, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure5" httpUri="https://binary.pensoft.net/fig/904111" pageId="0" pageNumber="169">5C</figureCitation>
,
<figureCitation id="D2ED3B39A6FBFC012FCEA254D9F7EBA0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Candidiopotamon penglai sp. nov., holotype male (38.4 x 33.4 mm) from Ji-an, Hualien (NCHUZOOL 12951), G 1 in bent form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal segment, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure6" httpUri="https://binary.pensoft.net/fig/904112" pageId="0" pageNumber="169">6C</figureCitation>
)]. Furthermore,
<taxonomicName id="0DAA2BADE9DA79EEFB14EA400DF2197F" lsidName="C. rathbuni" pageId="0" pageNumber="169" rank="species" species="rathbuni">
<emphasis id="672292F551EEE9271D19B17E9CBC6529" italics="true" pageId="0" pageNumber="169">C. rathbuni</emphasis>
</taxonomicName>
show almost straight subterminal segment of G1 (Fig.
<figureCitation id="33854349325BA1D2EF8D52F92A6EFA70" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Candidiopotamon rathbuni (De Man, 1914): male (44.6 x 36.8 mm) from Chiayi, western Taiwan (RUMF-ZC- 86) (A - E); male (35.4 x 31.2 mm) from Miaoli, northwestern Taiwan (NCHUZOOL 15194) (F 1); male (36.7 x 31.1 mm) from Chiayi, western Taiwan (NCHUZOOL 15205) (F 2); male (36.3 x 30.7 mm) from Taiwu, Pingtung, southwestern Taiwan (NCHUZOOL 15199) (F 3). A carapace, dorsal view B carapace, frontal view C male major chela, outer view D 1 left G 1, ventral view D 2 left G 1, dorsal view D 3 left G 1 terminal segment, dorsal view E left G 2 F 1 - 3 left G 1 s terminal segment, dorsal view. Scale bars: 10 mm (A - C); 2 mm (D - F). Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure3" httpUri="https://binary.pensoft.net/fig/904109" pageId="0" pageNumber="169">3D2</figureCitation>
), whereas those of
<taxonomicName id="725A315A5D7CEA7ED900EDA9AB1B2293" lsidName="C. penglai" pageId="0" pageNumber="169" rank="species" species="penglai">
<emphasis id="D50211F925568DD828AA9C0F7D5C68DA" italics="true" pageId="0" pageNumber="169">C. penglai</emphasis>
</taxonomicName>
are varied either straight (S clade, Fig.
<figureCitation id="BC89E0846CE566A5FCEC90F66C2BA09D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Candidiopotamon penglai sp. nov., male (42.9 x 36.2 mm) from Shihzih, Pingtung (NCHUZOOL 15207), G 1 in straight form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal joint, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure7" httpUri="https://binary.pensoft.net/fig/904113" pageId="0" pageNumber="169">7</figureCitation>
) or bent (E1, E2 and SE clades, Fig.
<figureCitation id="B8A6C8BFC12B06A3DFC89654E7C582BC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Candidiopotamon penglai sp. nov., holotype male (38.4 x 33.4 mm) from Ji-an, Hualien (NCHUZOOL 12951), G 1 in bent form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal segment, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure6" httpUri="https://binary.pensoft.net/fig/904112" pageId="0" pageNumber="169">6</figureCitation>
) (Figs
<figureCitation id="1D55E7DB8E5AA36D68E278B20D1376E3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites for Candidiopotamon rathbuni (De Man, 1914) (circles) and Candidiopotamon penglai sp. nov. (squares) in Taiwan, as well as other congeners from the Ryukyus, Japan. Different colors indicate sampling sites of clades. Dotted lines indicate the possible biogeographic boundaries between clades NW, W, SW, S, SE, E 1, and E 2, corresponding to the color in Fig. 2; as well as species. The red star and blue star indicate the type localities of C. rathbuni and C. penglai sp. nov., respectively. Three forms of male left G 1 are shown for clades in C. rathbuni and two forms for clades in C. penglai sp. nov." figureDoi="10.3897/zookeys.1179.106718.figure1" httpUri="https://binary.pensoft.net/fig/904107" pageId="0" pageNumber="169">1</figureCitation>
,
<figureCitation id="64BED15D74F38846A955AA9CF5B31A29" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Color in life of Candidiopotamon rathbuni (De Man, 1914) (A-D) and Candidiopotamon penglai sp. nov. (E-H). A, B NCHUZOOL 15194 (male, 35.4 x 31. 2 mm, Tai-an, Miaoli, northwestern Taiwan) C NCHUZOOL 15211 (male, 24.8 x 22.1 mm, Dongshih, Taichung, western Taiwan) D NCHUZOOL 15181 (male, 28.0 x 23.9 mm, Rihyuetan, Nantou, western Taiwan) E, F NCHUZOOL 15204 (2 males, 39.7 x 34.4 mm, 40.6 x 34.3 mm, Nanrenshan, Pingtung, southern Taiwan) G NCHUZOOL 15224 (paratype male, 36.4 x 30.9 mm, Liyu Lake, Shoufeng, Hualien, eastern Taiwan) H NCHUZOOL 15229 (male, 33.1 x 29.7 mm, Jili Lake, Guangfu, Hualien, eastern Taiwan)." figureDoi="10.3897/zookeys.1179.106718.figure5" httpUri="https://binary.pensoft.net/fig/904111" pageId="0" pageNumber="169">5B</figureCitation>
,
<figureCitation id="36F95E4A1702F081547B898C99F261A7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Candidiopotamon penglai sp. nov., holotype male (38.4 x 33.4 mm) from Ji-an, Hualien (NCHUZOOL 12951), G 1 in bent form A left G 1, ventral view B left G 1, dorsal view C left G 1 terminal segment, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment." figureDoi="10.3897/zookeys.1179.106718.figure6" httpUri="https://binary.pensoft.net/fig/904112" pageId="0" pageNumber="169">6B</figureCitation>
; see Discussion).
</paragraph>
<caption id="D4A8B1C2E3DD90267DEB108A920199F3" doi="10.3897/zookeys.1179.106718.figure6" httpUri="https://binary.pensoft.net/fig/904112" pageId="0" pageNumber="169" start="Figure 6" startId="F6">
<paragraph id="C32BEA3E9AD5911DBA4AC79460758421" pageId="0" pageNumber="169">
<emphasis id="693D3308B6E0E1D4D41B5E17DE0A1A64" bold="true" pageId="0" pageNumber="169">Figure 6.</emphasis>
<taxonomicName id="7DEEE3E260FB871EFADF730BEB4F289B" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon penglai" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="penglai">
<emphasis id="64CF98A2C7D433CE2055B8603A2051D9" italics="true" pageId="0" pageNumber="169">Candidiopotamon penglai</emphasis>
</taxonomicName>
sp. nov., holotype male (38.4
<normalizedToken id="86488DF14E54E8D1B7A20DEE6955A986" originalValue="×">x</normalizedToken>
33.4 mm) from Ji-an, Hualien (NCHUZOOL 12951), G1 in bent form
<emphasis id="EBD0A5083CC761C3EC009DE09F361FC7" bold="true" pageId="0" pageNumber="169">A</emphasis>
left G1, ventral view
<emphasis id="D52E10B536908073FD798214851EECCE" bold="true" pageId="0" pageNumber="169">B</emphasis>
left G1, dorsal view
<emphasis id="2E0CEB979F00D47ACDCCCE5DE60C9EF6" bold="true" pageId="0" pageNumber="169">C</emphasis>
left G1 terminal segment, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment.
</paragraph>
</caption>
<caption id="5B996BDBED2991F3C0AE7D360B034769" doi="10.3897/zookeys.1179.106718.figure7" httpUri="https://binary.pensoft.net/fig/904113" pageId="0" pageNumber="169" start="Figure 7" startId="F7">
<paragraph id="79A816071A608110481F0DD9B49C1BF7" pageId="0" pageNumber="169">
<emphasis id="612679BA44D9CD55460D44ED2ED5B113" bold="true" pageId="0" pageNumber="169">Figure 7.</emphasis>
<taxonomicName id="623F59E32830D0E6C622129F9356DE84" authorityName="Shih &amp; Naruse &amp; Schubart" authorityYear="2023" class="Malacostraca" family="Potamidae" genus="Candidiopotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Candidiopotamon penglai" order="Decapoda" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="penglai">
<emphasis id="A1B59F33F6A2E4E281175C267C15A847" italics="true" pageId="0" pageNumber="169">Candidiopotamon penglai</emphasis>
</taxonomicName>
sp. nov., male (42.9
<normalizedToken id="74639203F99D5B9C8A4E25D72BDCF217" originalValue="×">x</normalizedToken>
36.2 mm) from Shihzih, Pingtung (NCHUZOOL 15207), G1 in straight form
<emphasis id="B76ABD657F2A359EC12AD66077493C1A" bold="true" pageId="0" pageNumber="169">A</emphasis>
left G1, ventral view
<emphasis id="9F6FE94CEB67C07DB0BA234F66EE9A71" bold="true" pageId="0" pageNumber="169">B</emphasis>
left G1, dorsal view
<emphasis id="FB51E844156C1FA4FC56F0148EDC79D1" bold="true" pageId="0" pageNumber="169">C</emphasis>
left G1 terminal joint, dorsal view. Scale bars: 1 mm. Arrow indicates the keel-like projection on the distal inner edge of the terminal segment.
</paragraph>
</caption>
<paragraph id="15E86F661E35A6B0A284D31E5E4D4A92" pageId="0" pageNumber="169">
This new species includes clades from eastern Taiwan (E1, E2, SE and S, see Fig.
<figureCitation id="5B85A2E4FC06E023E499C57C5887D1FD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites for Candidiopotamon rathbuni (De Man, 1914) (circles) and Candidiopotamon penglai sp. nov. (squares) in Taiwan, as well as other congeners from the Ryukyus, Japan. Different colors indicate sampling sites of clades. Dotted lines indicate the possible biogeographic boundaries between clades NW, W, SW, S, SE, E 1, and E 2, corresponding to the color in Fig. 2; as well as species. The red star and blue star indicate the type localities of C. rathbuni and C. penglai sp. nov., respectively. Three forms of male left G 1 are shown for clades in C. rathbuni and two forms for clades in C. penglai sp. nov." figureDoi="10.3897/zookeys.1179.106718.figure1" httpUri="https://binary.pensoft.net/fig/904107" pageId="0" pageNumber="169">1</figureCitation>
) that are genetically distinct from
<taxonomicName id="1A7A2BDC68B08452FCA3519A2F12F7C2" lsidName="C. rathbuni" pageId="0" pageNumber="169" rank="species" species="rathbuni">
<emphasis id="FD57A0B56CF57624D84208974774C41A" italics="true" pageId="0" pageNumber="169">C. rathbuni</emphasis>
</taxonomicName>
from western Taiwan (NW, W and SW) (Table
<tableCitation id="A7B7F8F5C7F69EDA9FEDFFD85FA646C6" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Matrix of percentage pairwise nucleotide divergences with K 2 P distance and the number of bp differences based on COI within and between clades of Candidiopotamon species from Taiwan (see Table 1). In the right half, lower-left values are K 2 P distance and upper-right ones are bp differences. Values of the range are given in parentheses." httpUri="http://table.plazi.org/id/75481B5911D43028B059B4B0CA59AE74" pageId="0" pageNumber="169" tableUuid="75481B5911D43028B059B4B0CA59AE74">2</tableCitation>
, Fig.
<figureCitation id="A4F11A6CC2614BBB89F161B889C88620" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A Bayesian inference (BI) tree for Candidiopotamon rathbuni (De Man, 1914) and Candidiopotamon penglai sp. nov., as well as outgroups, based on the combined 16 S rDNA and cytochrome oxidase I. Probability values at the nodes represent support values for BI and maximum likelihood (ML). For haplotype names, see Table 1. The color of clades corresponds to the color of collection sites in Fig. 1, which are also shown as the numbers in parenthesis after the haplotype names." figureDoi="10.3897/zookeys.1179.106718.figure2" httpUri="https://binary.pensoft.net/fig/904108" pageId="0" pageNumber="169">2</figureCitation>
), with a reliable character of G1 also separating the two species (see above). Within
<taxonomicName id="E7E00F0E665A59276735451388A617E6" lsidName="C. penglai" pageId="0" pageNumber="169" rank="species" species="penglai">
<emphasis id="6107A20CD6A9D9AD96DEAEEC2F28B558" italics="true" pageId="0" pageNumber="169">C. penglai</emphasis>
</taxonomicName>
sp. nov., there are two forms of the curvature in G1, with the bent form found in the E1, E2 and SE clades, and the straight form in the S clade (Fig.
<figureCitation id="8347BD5B6FBCB7FEC80D2A0A4CC8B1DD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Collection sites for Candidiopotamon rathbuni (De Man, 1914) (circles) and Candidiopotamon penglai sp. nov. (squares) in Taiwan, as well as other congeners from the Ryukyus, Japan. Different colors indicate sampling sites of clades. Dotted lines indicate the possible biogeographic boundaries between clades NW, W, SW, S, SE, E 1, and E 2, corresponding to the color in Fig. 2; as well as species. The red star and blue star indicate the type localities of C. rathbuni and C. penglai sp. nov., respectively. Three forms of male left G 1 are shown for clades in C. rathbuni and two forms for clades in C. penglai sp. nov." figureDoi="10.3897/zookeys.1179.106718.figure1" httpUri="https://binary.pensoft.net/fig/904107" pageId="0" pageNumber="169">1</figureCitation>
; see Discussion).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>