323 lines
54 KiB
XML
323 lines
54 KiB
XML
<document id="536085E69E303172948997D2606B1EE2" ID-DOI="10.11646/zootaxa.5448.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231400" ID-ZooBank="F059B3E1-D86C-47C3-A7FA-58ADD4F7BBB0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716278756060" checkinUser="plazi" docAuthor="Jose, Athira, Caleb, John T. D. & Sudhikumar, Ambalaparambil Vasu" docDate="2024" docId="8B60CE05DF47FFFEFF3DFA5952C67D44" docLanguage="en" docName="zootaxa.5448.2.3.pdf" docOrigin="Zootaxa 5448 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5448.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habrocestum benjamin Jose & Caleb & Sudhikumar 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="217" masterDocId="7759B67DDF46FFFBFFAAFFCF50377971" masterDocTitle="On three species of the genus Habrocestum Simon, 1876 (Araneae: Salticidae: Hasariini) from India" masterLastPageNumber="224" masterPageNumber="212" pageNumber="213" updateTime="1716279297164" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0F77BED7C2F3717F5AED084F6416C820">
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<mods:title id="D56D462025FAA3450E362BD3692DC8AD">On three species of the genus Habrocestum Simon, 1876 (Araneae: Salticidae: Hasariini) from India</mods:title>
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</mods:titleInfo>
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<mods:name id="D531255C60331A4FB86B2593F40DFEE2" type="personal">
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<mods:role id="5B6DF6C294145AF8112518CDAE106F55">
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<mods:roleTerm id="CDC6CC7661ABC8CE82C1C1D12B499D6A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CEC1C3F75BE8ED569DE2DB1A6E80FD67">Jose, Athira</mods:namePart>
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<mods:nameIdentifier id="967C8B3AAD5E0015A39DC6453BB441C5" type="ORCID">0000-0001-9884-0718</mods:nameIdentifier>
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<mods:affiliation id="73B2B77CE6F4F1C0A656F93A214BB2BA">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, India (Affiliated to University of Calicut)</mods:affiliation>
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<mods:nameIdentifier id="ABA0F0310842C8DCF896B5F44939CF60" type="email">athirajose112@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="5AD11AC34E8CF230806B21C134C24FFC" type="personal">
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<mods:roleTerm id="74A94DDD9B27D518FBF32A03A13258B7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AD99F48BE5AF7CC95156C28E8548D22F">Caleb, John T. D.</mods:namePart>
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<mods:nameIdentifier id="41ED73CBB744B542E82D546754351F98" type="ORCID">0000-0002-9471-9467</mods:nameIdentifier>
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<mods:affiliation id="AB28211B4CE2EA1383DE1E419AE0733F">Department of Anatomy, Saveetha Medical College & Hospital, Saveetha Institute of Medical and Technical Sciences, Saveetha University, Chennai 602105, Tamil Nadu, India</mods:affiliation>
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<mods:nameIdentifier id="2517153722E2B0AF5E110F5C51DBB601" type="email">araneae.in@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="4B17ABE206C83D3141C9FCE9C532CAA1" type="personal">
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<mods:roleTerm id="49502E823BA271A3F39B4ACCDDFF32CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DBCFBE5AF500A3EDC1D15B2495322C85">Sudhikumar, Ambalaparambil Vasu</mods:namePart>
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<mods:nameIdentifier id="063FDB11C93740A18B1A648075A173B1" type="ORCID">0000-0002-4479-4995</mods:nameIdentifier>
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<mods:affiliation id="B4235973E24B241B146B88A17322767F">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Kerala, India (Affiliated to University of Calicut)</mods:affiliation>
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<mods:nameIdentifier id="80E3DD07AAAB6366C76B79F9C9002E71" type="email">avsudhi@rediffmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="895628F327B03F287FA5DA69574B89B3">text</mods:typeOfResource>
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<mods:title id="1569E8E3EA6E44D75A03986D6806470A">Zootaxa</mods:title>
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<mods:part id="DE9395F3F03B01D6FA9F2BE68FC1F5D5">
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<mods:date id="CE503FAB91D10A2ED136BBE6C402EAF7">2024</mods:date>
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<mods:number id="53CFC4BF29856ADF32B76B47BF865FFD">2024-05-10</mods:number>
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<mods:detail id="1073B2EC87B6D9BFB4BBDF8A7810F45E" type="volume">
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<mods:number id="C27098F1384F7517952D04853164DE38">5448</mods:number>
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<mods:number id="6FD69BDCE92C2DC68F825262CE5EA03D">2</mods:number>
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<mods:start id="FD7ED61149BBF489F468D081525C6E50">212</mods:start>
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<mods:url id="5305058E3A2FAA25FD89FEFDC099F732">http://dx.doi.org/10.11646/zootaxa.5448.2.3</mods:url>
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<mods:classification id="F508C2D6A39A8E70112366B9764BEBBB">journal article</mods:classification>
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<mods:identifier id="18E2A60F7B0AC0D4D56D93FF96689092" type="DOI">10.11646/zootaxa.5448.2.3</mods:identifier>
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<mods:identifier id="2D501F862A1FDAD7B5D1EEF3114921A8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D8BCF9FE6AEA2C8CD7541A31AC51BB16" type="Zenodo-Dep">11231400</mods:identifier>
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<mods:identifier id="39D78A590F21BE543639E5CB601E2392" type="ZooBank">F059B3E1-D86C-47C3-A7FA-58ADD4F7BBB0</mods:identifier>
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</mods:mods>
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<treatment id="8B60CE05DF47FFFEFF3DFA5952C67D44" LSID="urn:lsid:plazi:treatment:8B60CE05DF47FFFEFF3DFA5952C67D44" httpUri="http://treatment.plazi.org/id/8B60CE05DF47FFFEFF3DFA5952C67D44" lastPageId="5" lastPageNumber="217" pageId="1" pageNumber="213">
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<subSubSection id="4BD32C98DF47FFFAFF3DFA59522F7CC0" box="[151,536,1430,1457]" pageId="1" pageNumber="213" type="nomenclature">
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<paragraph id="03767F13DF47FFFAFF3DFA59522F7CC0" blockId="1.[151,536,1430,1493]" box="[151,536,1430,1457]" pageId="1" pageNumber="213">
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<heading id="583EC87FDF47FFFAFF3DFA59522F7CC0" bold="true" box="[151,536,1430,1457]" fontSize="11" level="1" pageId="1" pageNumber="213" reason="1">
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<emphasis id="31BDA301DF47FFFAFF3DFA59522F7CC0" bold="true" box="[151,536,1430,1457]" pageId="1" pageNumber="213">
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<taxonomicName id="C4C90490DF47FFFAFF3DFA5951847CC0" ID-CoL="3J4V5" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[151,435,1430,1457]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
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<emphasis id="31BDA301DF47FFFAFF3DFA5951847CC0" bold="true" box="[151,435,1430,1457]" italics="true" pageId="1" pageNumber="213">Habrocestum benjamin</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2A8E1E7ADF47FFFAFE11FA58522F7CC0" box="[443,536,1431,1457]" pageId="1" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BD32C98DF47FFFAFF3DFA7452347CA4" box="[151,515,1467,1493]" pageId="1" pageNumber="213" type="description">
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<paragraph id="03767F13DF47FFFAFF3DFA7452347CA4" blockId="1.[151,536,1430,1493]" box="[151,515,1467,1493]" pageId="1" pageNumber="213">
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<figureCitation id="9BF26396DF47FFFAFF3DFA7451397CA4" box="[151,270,1467,1493]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[243,1344,181,1919]" captionTargetId="figure-18@2.[243,1344,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. General habitus of Habrocestum swaminathan sp. nov. (A–F). A. Holotype male, dorsal view; B. Same, lateral view; C. Same, frontal view; D. Paratype female, dorsal view; E. Same lateral view; F. Same, frontal view. General habitus of Habrocestum benjamin sp. nov. (G–H). G. Holotype male, dorsal view; H. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231402" httpUri="https://zenodo.org/record/11231402/files/figure.png" pageId="1" pageNumber="213">Figs 1A–F</figureCitation>
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,
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<figureCitation id="9BF26396DF47FFFAFEB0FA7451567CA4" box="[282,353,1467,1493]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[174,1413,181,1877]" captionTargetId="figure-22@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Somatic morphology of Habrocestum benjamin sp. nov. A–D. Holotype male; A. Dorsal view; B. Lateral view; C. Frontal view; D. Leg I retrolateral view. E–H. Female paratype; E. Dorsal view; F. Lateral view; G. Frontal view; H. Leg 1 retrolateral view. Scale bars: 1 mm (A, D, E); 0.5 mm (B–C, F–H)." figureDoi="http://doi.org/10.5281/zenodo.11231405" httpUri="https://zenodo.org/record/11231405/files/figure.png" pageId="1" pageNumber="213">2A–H</figureCitation>
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,
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<figureCitation id="9BF26396DF47FFFAFEC6FA7451847CA4" box="[364,435,1467,1493]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="1" pageNumber="213">3A–D</figureCitation>
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,
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<figureCitation id="9BF26396DF47FFFAFE15FA7452347CA4" box="[447,515,1467,1493]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="1" pageNumber="213">6A–D</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BD32C98DF47FFFAFF3DF9CC51287F14" pageId="1" pageNumber="213" type="materials_examined">
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<paragraph id="03767F13DF47FFFAFF3DF9CC51287F14" blockId="1.[151,1437,1538,2033]" pageId="1" pageNumber="213">
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<emphasis id="31BDA301DF47FFFAFF3DF9CC517F7F6D" bold="true" box="[151,328,1538,1565]" pageId="1" pageNumber="213">Type material.</emphasis>
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||
<materialsCitation id="B3A1754EDF47FFFAFEF8F9CC53A17F30" collectingDate="2023-05-04" collectorName="A. Jose" country="India" elevation="420" latitude="11.472221" location="Thusharagiri" longLatPrecision="21" longitude="76.052505" municipality="Kozhikode" pageId="1" pageNumber="213" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
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<emphasis id="31BDA301DF47FFFAFEF8F9CC51FF7F6C" bold="true" box="[338,456,1539,1565]" pageId="1" pageNumber="213">
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||
<typeStatus id="DC72C1B1DF47FFFAFEF8F9CC51F47F6C" box="[338,451,1539,1565]" pageId="1" pageNumber="213" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="15CFB49ADF47FFFAFE78F9CC51D07F6C" box="[466,487,1539,1565]" count="1" pageId="1" pageNumber="213" type="male">♂</specimenCount>
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(CATE 10321807a) from
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<emphasis id="31BDA301DF47FFFAFCB1F9CD53EF7F6C" bold="true" box="[795,984,1538,1565]" pageId="1" pageNumber="213">
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||
<collectingCountry id="7BDE3F83DF47FFFAFCB1F9CD53457F6D" box="[795,882,1538,1564]" name="India" pageId="1" pageNumber="213">INDIA</collectingCountry>
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||
:
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<emphasis id="31BDA301DF47FFFAFC2BF9CD53F87F6C" bold="true" box="[897,975,1538,1565]" italics="true" pageId="1" pageNumber="213">
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<collectingRegion id="C10DB1F1DF47FFFAFC2BF9CD53F87F6C" box="[897,975,1538,1565]" country="India" name="Kerala" pageId="1" pageNumber="213">Kerala</collectingRegion>
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</emphasis>
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:
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</emphasis>
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<collectingMunicipality id="E312E569DF47FFFAFC48F9CC54567F6C" box="[994,1121,1539,1565]" pageId="1" pageNumber="213">Kozhikode</collectingMunicipality>
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:
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<location id="061629C8DF47FFFAFBC5F9CC54C97F6C" LSID="urn:lsid:plazi:treatment:8B60CE05DF47FFFEFF3DFA5952C67D44:061629C8DF47FFFAFBC5F9CC54C97F6C" box="[1135,1278,1539,1565]" country="India" latitude="11.472221" longLatPrecision="21" longitude="76.052505" municipality="Kozhikode" name="Thusharagiri" pageId="1" pageNumber="213" stateProvince="Kerala">Thusharagiri</location>
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(
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<geoCoordinate id="66FD19D4DF47FFFAFABAF9CB55A07F6E" box="[1296,1431,1540,1567]" degrees="11" direction="north" minutes="28" orientation="latitude" pageId="1" pageNumber="213" precision="15" seconds="20" value="11.472221">11°28′20″N</geoCoordinate>
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,
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<geoCoordinate id="66FD19D4DF47FFFAFF3DF9E751377F32" box="[151,256,1576,1603]" degrees="76" direction="east" minutes="3" orientation="longitude" pageId="1" pageNumber="213" precision="15" seconds="9" value="76.052505">76°3′9″E</geoCoordinate>
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||
,
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<quantity id="C431D2F6DF47FFFAFEA0F9E851677F30" box="[266,336,1575,1601]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="213" unit="m" value="420.0">
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<elevation id="88E49820DF47FFFAFEA0F9E851677F30" box="[266,336,1575,1601]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="213" unit="m" value="420.0">420 m</elevation>
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</quantity>
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alt.),
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<date id="777759D3DF47FFFAFE3AF9E852117F30" box="[400,550,1575,1601]" pageId="1" pageNumber="213" value="2023-05-04">
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||
<collectingDate id="6733A03BDF47FFFAFE3AF9E852117F30" box="[400,550,1575,1601]" pageId="1" pageNumber="213" value="2023-05-04">04 May 2023</collectingDate>
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||
</date>
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||
,
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<collectorName id="AE3C1AC5DF47FFFAFD84F9E852B77F30" box="[558,640,1575,1601]" pageId="1" pageNumber="213">A. Jose</collectorName>
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leg., from litter, by hand
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</materialsCitation>
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||
.
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<materialsCitation id="B3A1754EDF47FFFAFC35F9E8512B7F14" collectingDate="2023-05-04" collectorName="A. Jose" country="India" elevation="420" latitude="11.472221" location="Thusharagiri" longLatPrecision="21" longitude="76.052505" municipality="Kozhikode" pageId="1" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Paratype" typeStatus="paratype">
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<emphasis id="31BDA301DF47FFFAFC35F9E854227F30" bold="true" box="[927,1045,1575,1601]" pageId="1" pageNumber="213">
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<typeStatus id="DC72C1B1DF47FFFAFC35F9E854277F30" box="[927,1040,1575,1601]" pageId="1" pageNumber="213" type="paratype">Paratype</typeStatus>
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||
:
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</emphasis>
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||
<specimenCount id="15CFB49ADF47FFFAFBB1F9E854097F30" box="[1051,1086,1575,1601]" count="1" pageId="1" pageNumber="213" type="female">1♀</specimenCount>
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(CATE 10321807b), same data as holotype
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4BD32C98DF47FFFAFF6DF9A055377FDC" pageId="1" pageNumber="213" type="etymology">
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||
<paragraph id="03767F13DF47FFFAFF6DF9A055377FDC" blockId="1.[151,1437,1538,2033]" pageId="1" pageNumber="213">
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||
<emphasis id="31BDA301DF47FFFAFF6DF9A051797FF8" bold="true" box="[199,334,1647,1673]" pageId="1" pageNumber="213">Etymology.</emphasis>
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The specific epithet is a patronym in honor of Prof. Suresh P. Benjamin for his remarkable dedication and significant contributions to the field of arachnology. The name is treated as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="4BD32C98DF47FFFAFF6DF97954B27E80" pageId="1" pageNumber="213" type="diagnosis">
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||
<paragraph id="03767F13DF47FFFAFF6DF97954B27E80" blockId="1.[151,1437,1538,2033]" pageId="1" pageNumber="213">
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||
<emphasis id="31BDA301DF47FFFAFF6DF97951757FA1" bold="true" box="[199,322,1718,1744]" pageId="1" pageNumber="213">Diagnosis.</emphasis>
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The males of
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<taxonomicName id="C4C90490DF47FFFAFE4DF97852DA7FA0" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[487,749,1719,1745]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
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<emphasis id="31BDA301DF47FFFAFE4DF97852DA7FA0" box="[487,749,1719,1745]" italics="true" pageId="1" pageNumber="213">Habrocestum benjamin</emphasis>
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</taxonomicName>
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<emphasis id="31BDA301DF47FFFAFD5FF97853677FA0" bold="true" box="[757,848,1719,1745]" pageId="1" pageNumber="213">
|
||
<taxonomicNameLabel id="2A8E1E7ADF47FFFAFD5FF97853677FA0" box="[757,848,1719,1745]" pageId="1" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
||
can be distinguished from the known congeners by the presence of compound terminal apophysis on the embolus. The palp is similar to
|
||
<taxonomicName id="C4C90490DF47FFFAFBDEF91452107E68" authority="Sankaran, Malamel, Joseph & Sebastian, 2019" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFBDEF91455147F84" box="[1140,1315,1755,1781]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
<bibRefCitation id="675802E2DF47FFFAFA87F91452107E68" author="Sankaran, P. M. & Malamel, J. J. & Joseph, M. M. & Sebastian, P. A." pageId="1" pageNumber="213" pagination="1 - 15" refId="ref6798" refString="Sankaran, P. M., Malamel, J. J., Joseph, M. M. & Sebastian, P. A. (2019) New species of Habrocestum Simon, 1876 and a redescription of Curubis tetrica Simon, 1902 (Araneae: Salticidae: Salticinae: Hasariini) from India. Journal of Natural History, 53 (1 - 2), 1 - 15. https: // doi. org / 10.1080 / 00222933.2019.1567856" type="journal article" year="2019">
|
||
Sankaran, Malamel, Joseph & Sebastian,
|
||
<quantity id="C431D2F6DF47FFFAFE45F93052107E68" box="[495,551,1791,1817]" metricMagnitude="1" metricUnit="m" metricValue="5.12826" pageId="1" pageNumber="213" unit="in" value="2019.0">2019</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in the absence of proximal lobe of tegulum and the presence of anterior median tegular lobe. However, the small, roundish, forwardly bent RTA (vs. broad, flat with finger-like projection in
|
||
<taxonomicName id="C4C90490DF47FFFAFA2BF8EC51167E10" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFA2BF8EC51167E10" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
) and the long, elongated cymbium (vs. short and triangular in
|
||
<taxonomicName id="C4C90490DF47FFFAFC4FF88854A57E10" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" box="[997,1170,1863,1889]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFC4FF88854A57E10" box="[997,1170,1863,1889]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
) with an oval, compact tegulum (roundish and saggy in
|
||
<taxonomicName id="C4C90490DF47FFFAFDABF8A452997EF4" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" box="[513,686,1899,1925]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFDABF8A452997EF4" box="[513,686,1899,1925]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
) distinguishes the two species (
|
||
<figureCitation id="9BF26396DF47FFFAFBB8F8A454A67EF4" box="[1042,1169,1899,1925]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="1" pageNumber="213">Figs 3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF47FFFAFB36F8A454E87EF4" box="[1180,1247,1899,1925]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="1" pageNumber="213">6A–B</figureCitation>
|
||
vs. figs 3A–E in
|
||
<bibRefCitation id="675802E2DF47FFFAFF3DF84051B57ED8" author="Sankaran, P. M. & Malamel, J. J. & Joseph, M. M. & Sebastian, P. A." box="[151,386,1935,1961]" pageId="1" pageNumber="213" pagination="1 - 15" refId="ref6798" refString="Sankaran, P. M., Malamel, J. J., Joseph, M. M. & Sebastian, P. A. (2019) New species of Habrocestum Simon, 1876 and a redescription of Curubis tetrica Simon, 1902 (Araneae: Salticidae: Salticinae: Hasariini) from India. Journal of Natural History, 53 (1 - 2), 1 - 15. https: // doi. org / 10.1080 / 00222933.2019.1567856" type="journal article" year="2019">
|
||
Sankaran
|
||
<emphasis id="31BDA301DF47FFFAFEACF84051087ED8" box="[262,319,1935,1961]" italics="true" pageId="1" pageNumber="213">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
). Females can be differentiated by the narrow epigynal pockets (vs. broad in
|
||
<taxonomicName id="C4C90490DF47FFFAFB75F84055BC7ED8" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" box="[1247,1419,1935,1961]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFB75F84055BC7ED8" box="[1247,1419,1935,1961]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
), small accessory glands (vs. large in
|
||
<taxonomicName id="C4C90490DF47FFFAFD94F87C52D97EBC" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" box="[574,750,1971,1997]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFD94F87C52D97EBC" box="[574,750,1971,1997]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
) and large multi chambered spermatheca (vs. small, single chambered in
|
||
<taxonomicName id="C4C90490DF47FFFAFE9CF81851D57E80" authorityName="Sankaran, Malamel, Joseph & Sebastian" authorityYear="2019" box="[310,482,2007,2033]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="213" phylum="Arthropoda" rank="species" species="longispinum">
|
||
<emphasis id="31BDA301DF47FFFAFE9CF81851D57E80" box="[310,482,2007,2033]" italics="true" pageId="1" pageNumber="213">H. longispinum</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="9BF26396DF47FFFAFE51F818524F7E80" box="[507,632,2007,2033]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="1" pageNumber="213">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF47FFFAFD2EF81852F07E80" box="[644,711,2007,2033]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="1" pageNumber="213">6C–D</figureCitation>
|
||
vs. figs 3G–H in
|
||
<bibRefCitation id="675802E2DF47FFFAFC24F818544E7E80" author="Sankaran, P. M. & Malamel, J. J. & Joseph, M. M. & Sebastian, P. A." box="[910,1145,2007,2033]" pageId="1" pageNumber="213" pagination="1 - 15" refId="ref6798" refString="Sankaran, P. M., Malamel, J. J., Joseph, M. M. & Sebastian, P. A. (2019) New species of Habrocestum Simon, 1876 and a redescription of Curubis tetrica Simon, 1902 (Araneae: Salticidae: Salticinae: Hasariini) from India. Journal of Natural History, 53 (1 - 2), 1 - 15. https: // doi. org / 10.1080 / 00222933.2019.1567856" type="journal article" year="2019">
|
||
Sankaran
|
||
<emphasis id="31BDA301DF47FFFAFC57F81854017E80" box="[1021,1078,2007,2033]" italics="true" pageId="1" pageNumber="213">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="57B62F9BDF44FFF9FF3DF85854887E86" ID-DOI="http://doi.org/10.5281/zenodo.11231402" ID-Zenodo-Dep="11231402" httpUri="https://zenodo.org/record/11231402/files/figure.png" pageId="2" pageNumber="214" startId="2.[151,250,1943,1967]" targetBox="[243,1344,181,1919]" targetPageId="2" targetType="figure">
|
||
<paragraph id="03767F13DF44FFF9FF3DF85854887E86" blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="214">
|
||
<emphasis id="31BDA301DF44FFF9FF3DF85851227EDE" bold="true" box="[151,277,1943,1967]" pageId="2" pageNumber="214">FIGURE 1.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="C4C90490DF44FFF9FE4EF85852CC7EDE" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[484,763,1943,1967]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="214" phylum="Arthropoda" rank="species" species="swaminathan" status="sp. nov.">
|
||
<emphasis id="31BDA301DF44FFF9FE4EF85852CC7EDE" box="[484,763,1943,1967]" italics="true" pageId="2" pageNumber="214">Habrocestum swaminathan</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31BDA301DF44FFF9FCA9F85753627EC1" bold="true" box="[771,853,1944,1968]" pageId="2" pageNumber="214">
|
||
<taxonomicNameLabel id="2A8E1E7ADF44FFF9FCA9F85753627EC1" box="[771,853,1944,1968]" pageId="2" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A–F). A. Holotype male, dorsal view; B. Same, lateral view; C. Same, frontal view; D. Paratype female, dorsal view; E. Same lateral view; F. Same, frontal view. General habitus of
|
||
<taxonomicName id="C4C90490DF44FFF9FF3DF81051B47E86" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[151,387,2015,2039]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="214" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
|
||
<emphasis id="31BDA301DF44FFF9FF3DF81051B47E86" box="[151,387,2015,2039]" italics="true" pageId="2" pageNumber="214">Habrocestum benjamin</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31BDA301DF44FFF9FE20F82F51ED7E89" bold="true" box="[394,474,2016,2040]" pageId="2" pageNumber="214">
|
||
<taxonomicNameLabel id="2A8E1E7ADF44FFF9FE20F82F51ED7E89" box="[394,474,2016,2040]" pageId="2" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(G–H). G. Holotype male, dorsal view; H. Paratype female, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="57B62F9BDF45FFF8FF3DF85853097E86" ID-DOI="http://doi.org/10.5281/zenodo.11231405" ID-Zenodo-Dep="11231405" httpUri="https://zenodo.org/record/11231405/files/figure.png" pageId="3" pageNumber="215" startId="3.[151,250,1943,1967]" targetBox="[174,1413,181,1877]" targetPageId="3" targetType="figure">
|
||
<paragraph id="03767F13DF45FFF8FF3DF85853097E86" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="215">
|
||
<emphasis id="31BDA301DF45FFF8FF3DF85851237EDE" bold="true" box="[151,276,1943,1967]" pageId="3" pageNumber="215">FIGURE 2.</emphasis>
|
||
Somatic morphology of
|
||
<taxonomicName id="C4C90490DF45FFF8FDBDF85853337EDE" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[535,772,1943,1967]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="215" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
|
||
<emphasis id="31BDA301DF45FFF8FDBDF85853337EDE" box="[535,772,1943,1967]" italics="true" pageId="3" pageNumber="215">Habrocestum benjamin</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31BDA301DF45FFF8FCA6F85853697EDE" bold="true" box="[780,862,1943,1967]" pageId="3" pageNumber="215">
|
||
<taxonomicNameLabel id="2A8E1E7ADF45FFF8FCA6F85853697EDE" box="[780,862,1943,1967]" pageId="3" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–D. Holotype male; A. Dorsal view; B. Lateral view; C. Frontal view; D. Leg I retrolateral view. E–H. Female paratype; E. Dorsal view; F. Lateral view; G. Frontal view; H. Leg 1 retrolateral view. Scale bars: 1 mm (A, D, E); 0.5 mm (B–C, F–H).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="57B62F9BDF42FFFFFF3DFB8051157D86" ID-DOI="http://doi.org/10.5281/zenodo.11231409" ID-Zenodo-Dep="11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="4" pageNumber="216" startId="4.[151,250,1103,1127]" targetBox="[158,1436,181,1079]" targetPageId="4" targetType="figure">
|
||
<paragraph id="03767F13DF42FFFFFF3DFB8051157D86" blockId="4.[151,1437,1103,1271]" pageId="4" pageNumber="216">
|
||
<emphasis id="31BDA301DF42FFFFFF3DFB8051217D16" bold="true" box="[151,278,1103,1127]" pageId="4" pageNumber="216">FIGURE 3.</emphasis>
|
||
Genital morphology of
|
||
<taxonomicName id="C4C90490DF42FFFFFDBFFB8053337D16" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[533,772,1103,1127]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="216" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
|
||
<emphasis id="31BDA301DF42FFFFFDBFFB8053337D16" box="[533,772,1103,1127]" italics="true" pageId="4" pageNumber="216">Habrocestum benjamin</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31BDA301DF42FFFFFCA7FB8053567D16" bold="true" box="[781,865,1103,1127]" pageId="4" pageNumber="216">
|
||
<taxonomicNameLabel id="2A8E1E7ADF42FFFFFCA7FB8053567D16" box="[781,865,1103,1127]" pageId="4" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4BD32C98DF42FFFEFF6DFAEA52D17D60" lastPageId="5" lastPageNumber="217" pageId="4" pageNumber="216" type="description">
|
||
<paragraph id="03767F13DF42FFFEFF6DFAEA51AB78F8" blockId="4.[151,1437,1317,2028]" lastBlockId="5.[151,1437,151,1077]" lastPageId="5" lastPageNumber="217" pageId="4" pageNumber="216">
|
||
<emphasis id="31BDA301DF42FFFFFF6DFAEA51967C31" bold="true" box="[199,417,1317,1344]" pageId="4" pageNumber="216">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="DC72C1B1DF42FFFFFE19FAEA522E7C4E" box="[435,537,1317,1343]" pageId="4" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
, CATE 10321807a) (
|
||
<figureCitation id="9BF26396DF42FFFFFCB8FAEA53A37C31" box="[786,916,1317,1344]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[243,1344,181,1919]" captionTargetId="figure-18@2.[243,1344,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. General habitus of Habrocestum swaminathan sp. nov. (A–F). A. Holotype male, dorsal view; B. Same, lateral view; C. Same, frontal view; D. Paratype female, dorsal view; E. Same lateral view; F. Same, frontal view. General habitus of Habrocestum benjamin sp. nov. (G–H). G. Holotype male, dorsal view; H. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231402" httpUri="https://zenodo.org/record/11231402/files/figure.png" pageId="4" pageNumber="216">Figs 1A–C</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF42FFFFFC0EFAE953DB7C31" box="[932,1004,1318,1344]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[174,1413,181,1877]" captionTargetId="figure-22@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Somatic morphology of Habrocestum benjamin sp. nov. A–D. Holotype male; A. Dorsal view; B. Lateral view; C. Frontal view; D. Leg I retrolateral view. E–H. Female paratype; E. Dorsal view; F. Lateral view; G. Frontal view; H. Leg 1 retrolateral view. Scale bars: 1 mm (A, D, E); 0.5 mm (B–C, F–H)." figureDoi="http://doi.org/10.5281/zenodo.11231405" httpUri="https://zenodo.org/record/11231405/files/figure.png" pageId="4" pageNumber="216">2A–D</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF42FFFFFC51FAE954767C31" box="[1019,1089,1318,1344]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="4" pageNumber="216">3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF42FFFFFBFAFAEA54A07C4E" box="[1104,1175,1317,1343]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="4" pageNumber="216">6A–B</figureCitation>
|
||
). Measurements: body length 3.88; carapace 2.12 long, 1.72 wide, height at PME 1.20; abdomen 1.76 long, 1.20 wide. Eye sizes and ocular distance: AME 0.53, ALE 0.31, PME 0.06, PLE 0.18; AME–AME 0.03, AME–ALE 0.03, ALE–ALE 0.08, PME–PME 1.25, PME–PLE 0.36, PLE–PLE 1.08, ALE–PME 0.38, ALE–PLE 0.77; Length of chelicerae 0.66. Clypeus height at AMEs 0.21. Length of pedipalp and legs: pedipalp 2.09 (0.80, 0.36, 0.27, 0.66), I 6.94 (2.26, 1.13, 1.80, 1.09, 0.66), II 4.48 (1.48, 0.75, 1.05, 0.76, 0.44), III 4.94 (1.75, 0.59, 1.13, 1.05, 0.42), IV 4.23 (1.29, 0.47, 0.85, 1.25, 0.37). Leg formula: 1324. Leg spination: femur pld 2, II pld 1 rld 2, III do 1 pld 3, IV do 3 pld 1; patella I–II 0, III pl 1 IV do 1; tibia I plv 4 rlv 3, II pl 1 plv 2 rlv 4, III pl 3 plv 1 rl 2 rld 1, IV pl 2 rl 3 rlv 1; metatarsus I plv 2 rlv 2, II pl 2 plv 2 rlv 2, III pl 2 pld 2 plv 2 rl 1 rld 2 rlv 1, IV pl 1 pld 2 rld 3; tarsus I–IV 0. Colouration (in alcohol): carapace orange-brown, adorned with posterolateral white stripes that extend to the dorsum; seven white patches present, one down each PME, one on inner margin of each PME, one down near each PLE, and one down the fovea. Eye field dark, covered with pale golden yellow hairs, intermixed with long black vertical hairs, purfled above AER. Anterior eyes surrounded by orange hairs; clypeus orange-brown, devoid of hairs. Chelicerae chestnut brown provided with sparse black, long hairs; endites, labium and sternum yellow brown. Abdomen hirsute with thin anterior scutum, slightly reddish, marked with nearly cordiform black patch; to the rear a deep black patch present, venter creamy yellow with olive-gray mottling. Anterior and median spinnerets creamy white, posterior spinnerets pale brown. Carapace broad, wide U-shaped, flattish, approximately twice the width of abdomen, with an abruptly truncated base. Fovea small, linear; centrally placed between PMEs. Chelicerae with two closely spaced teeth on the promargin and single bicuspid tooth on the retromargin, with one ramus reduced and blunt; endites marked by sclerotized, smooth anterior edge with reduced scopulae; absent on labium. Leg I dark with distinct long spines on femur and metatarsus; subsequent legs pale with black annulations near joints. Abdomen elliptical, narrower than prosoma. Anterior and median spinnerets are longer than posterior. Pedipalp (
|
||
<figureCitation id="9BF26396DF43FFFEFB10FF74550C79A4" box="[1210,1339,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">Figs 3A–B</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFAE2FF7455A779A4" box="[1352,1424,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6A–B</figureCitation>
|
||
): femur brown at the distal end, other segments creamy yellow, distoprolateraly adorned with white hairs. Cymbium elongated and tri-oval. RTA small, roundish with a significant forward bent (
|
||
<figureCitation id="9BF26396DF43FFFEFBA1FECC545E786C" box="[1035,1129,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFBDDFECC54AC786C" box="[1143,1179,259,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6B</figureCitation>
|
||
). Bulbus sac-like with an anterior median tegular lobe positioned at tegular cleft; tegular ledge extends conspicuously to embolic base. Embolus short, triangular, emerging at 12 o'clock position; compound terminal apophysis present, broad, twisted dorsally (
|
||
<figureCitation id="9BF26396DF43FFFEFEAAFEA0516A78F8" box="[256,349,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFEC3FEA051B878F8" box="[361,399,367,393]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="03767F13DF43FFFEFF6DFE5C54767AD4" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="217">
|
||
<emphasis id="31BDA301DF43FFFEFF6DFE5C512978DC" bold="true" box="[199,286,403,429]" pageId="5" pageNumber="217">Female</emphasis>
|
||
(
|
||
<typeStatus id="DC72C1B1DF43FFFEFE85FE5C51A478DC" box="[303,403,403,429]" pageId="5" pageNumber="217" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<figureCitation id="9BF26396DF43FFFEFE06FE5C521078DC" box="[428,551,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[243,1344,181,1919]" captionTargetId="figure-18@2.[243,1344,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. General habitus of Habrocestum swaminathan sp. nov. (A–F). A. Holotype male, dorsal view; B. Same, lateral view; C. Same, frontal view; D. Paratype female, dorsal view; E. Same lateral view; F. Same, frontal view. General habitus of Habrocestum benjamin sp. nov. (G–H). G. Holotype male, dorsal view; H. Paratype female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11231402" httpUri="https://zenodo.org/record/11231402/files/figure.png" pageId="5" pageNumber="217">Figs 1D–F</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFD9FFE5C524E78DC" box="[565,633,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[174,1413,181,1877]" captionTargetId="figure-22@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Somatic morphology of Habrocestum benjamin sp. nov. A–D. Holotype male; A. Dorsal view; B. Lateral view; C. Frontal view; D. Leg I retrolateral view. E–H. Female paratype; E. Dorsal view; F. Lateral view; G. Frontal view; H. Leg 1 retrolateral view. Scale bars: 1 mm (A, D, E); 0.5 mm (B–C, F–H)." figureDoi="http://doi.org/10.5281/zenodo.11231405" httpUri="https://zenodo.org/record/11231405/files/figure.png" pageId="5" pageNumber="217">2E–H</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFD2DFE5C52FB78DC" box="[647,716,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFD70FE5C531578DC" box="[730,802,403,429]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6C–D</figureCitation>
|
||
). Measurements: body length 3.64; carapace 1.74 long, 1.34 wide, 1.01 height; abdomen 1.90 long, 1.19 wide. Eye sizes and ocular distance: AME 0.41, ALE 0.27, PME 0.08, PLE 0.17; AME–AME 0.03, AME–ALE 0.03, ALE–ALE 0.08, PME–PME 0.96, PME–PLE 0.26, PLE–PLE 0.87, ALE–PME 0.21, ALE–PLE 0.54. Length of chelicerae 0.46. Clypeus height at AMEs 0.16. Length of palp and legs: palp 1.46 (0.55, 0.22, 0.27, 0.42), I 3.54 (1.16, 0.53, 0.96, 0.47, 0.42), II 2.59 (0.84, 0.45, 0.55, 0.42, 0.33), III 3.44 (1.29, 0.49, 0.60, 0.68, 0.38), IV 3.31 (0.97, 0.52, 0.52, 0.85, 0.45). Leg formula 1342. Spination of legs: femur I 0, II pld 3, III pld 2 rld 1, IV do 2 pld 1; patella I–II 0, III–IV rl 1; tibia I plv 4 rlv 3, II pl 1 plv 2 rlv 3, III pl 1 rl 2 rlv 1, IV 0; metatarsus I plv 2 rlv 2, II plv 4 rlv 2, III pl 1 pld 2 plv 1 rl 1 rld 1 rlv 2, IV plv 2 pl 1 rlv 2; tarsus I–IV 0. Colouration (in alcohol): body form and color same as in male except for the following: carapace not exceeding width of abdomen, lateral margin not apparent in dorsal view, abdomen ovoid with median W-shaped black patch on creamy-white background. Palps pale yellow. Epigyne with a postero-median epigynal pocket. Copulatory openings present at mid-posterior region, positioned above epigynal pockets (
|
||
<figureCitation id="9BF26396DF43FFFEFC39FCD053D97A48" box="[915,1006,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFC51FCD054287A48" box="[1019,1055,799,825]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6C</figureCitation>
|
||
). Copulatory ducts narrow, gently bending and moving anteriorly entering spermathecae at mid-lateral region; spermathecae moderately sclerotized, multi-chambered, anterior portion tubular and bent, with thick walls. Accessory glands present laterally. Fertilization ducts originated from anterior portion of tubular part of spermatheca (
|
||
<figureCitation id="9BF26396DF43FFFEFC0CFC4454347AD4" box="[934,1027,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1103,1127]" captionTargetBox="[158,1436,181,1079]" captionTargetId="figure-18@4.[151,1436,181,1079]" captionTargetPageId="4" captionText="FIGURE 3. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D). Abbreviations: AG—accessory gland; amTL—anterior median tegular lobe; CO—copulatory opening; CTA—compound terminal apophysis; E—embolus; EP—epigynal pocket; FD—fertilization duct; RTA—retrolateral tibial apophysis; S—spermatheca; SM—spermaphor; TL— tegular ledge." figureDoi="http://doi.org/10.5281/zenodo.11231409" httpUri="https://zenodo.org/record/11231409/files/figure.png" pageId="5" pageNumber="217">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="9BF26396DF43FFFEFBA4FC4454037AD4" box="[1038,1076,907,933]" captionStart="FIGURE 6" captionStartId="8.[151,250,1846,1870]" captionTargetBox="[162,1428,1000,1816]" captionTargetId="figure-417@8.[151,1436,993,1822]" captionTargetPageId="8" captionText="FIGURE 6. Genital morphology of Habrocestum benjamin sp. nov. A. Left male palp, ventral view; B. Same, retrolateral view; C. Female epigyne, ventral view; D. Vulva, dorsal view. Scale bars: 0.2 mm (A–D)." figureDoi="http://doi.org/10.5281/zenodo.11231418" httpUri="https://zenodo.org/record/11231418/files/figure.png" pageId="5" pageNumber="217">6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="03767F13DF43FFFEFF6DFC6052D17D60" blockId="5.[151,1437,151,1077]" pageId="5" pageNumber="217">
|
||
<emphasis id="31BDA301DF43FFFEFF6DFC6051B27AB9" bold="true" box="[199,389,942,969]" pageId="5" pageNumber="217">Natural history.</emphasis>
|
||
<taxonomicName id="C4C90490DF43FFFEFE26FC6052A67AB8" authority="Jose & Caleb & Sudhikumar, 2024" authorityName="Jose & Caleb & Sudhikumar" authorityYear="2024" box="[396,657,943,969]" class="Arachnida" family="Salticidae" genus="Habrocestum" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="217" phylum="Arthropoda" rank="species" species="benjamin" status="sp. nov.">
|
||
<emphasis id="31BDA301DF43FFFEFE26FC6052A67AB8" box="[396,657,943,969]" italics="true" pageId="5" pageNumber="217">Habrocestum benjamin</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="31BDA301DF43FFFEFD33FC6052C47AB8" bold="true" box="[665,755,943,969]" pageId="5" pageNumber="217">
|
||
<taxonomicNameLabel id="2A8E1E7ADF43FFFEFD33FC6052C47AB8" box="[665,755,943,969]" pageId="5" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was spotted in the damp leaf litter of tropical rainforest, in a slightly shaded area along the forest edge. Several females and juveniles were observed staying within retreats made on small, low-lying leaves, by folding the leaf blade.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BD32C98DF43FFFEFF6DFBD552C67D44" box="[199,753,1050,1077]" pageId="5" pageNumber="217" type="distribution">
|
||
<paragraph id="03767F13DF43FFFEFF6DFBD552C67D44" blockId="5.[151,1437,151,1077]" box="[199,753,1050,1077]" pageId="5" pageNumber="217">
|
||
<emphasis id="31BDA301DF43FFFEFF6DFBD551567D45" bold="true" box="[199,353,1050,1076]" pageId="5" pageNumber="217">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="DC72C1B1DF43FFFEFDCAFBD452A77D44" box="[608,656,1051,1077]" pageId="5" pageNumber="217">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |