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treatments-xml/data/6C/57/96/6C5796057B7CFFC7FF093F869D692CF9.xml
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<document id="38794EFF2DDB65BCC7EA7F759C3C0230" ID-CLB-Dataset="22256" ID-DOI="10.11646/zootaxa.4772.3.7" ID-GBIF-Dataset="8b277405-a91c-4d0c-b77e-8b1d4820c7cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819764" ID-ZooBank="8DE2F40F-0931-4002-97C4-5603E363B7E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175825495" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel Ángel" docDate="2020" docId="6C5796057B7CFFC7FF093F869D692CF9" docLanguage="en" docName="zootaxa.4772.3.7.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrodactylus eniocanoi Arce-Perez &amp; Moron 2020, new species" docType="treatment" docVersion="7" lastPageNumber="569" masterDocId="906EEE7D7B7EFFC5FF9E3F1F990C2A65" masterDocTitle="Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus" masterLastPageNumber="584" masterPageNumber="567" pageNumber="569" updateTime="1698839127964" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9849951952774B306FDF3B50EE745A15" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="599BA45370F5CEBEF9CC9D8D24C70943">
<mods:title id="05AFE4F6DD86242D00302037EA51920B">Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus</mods:title>
</mods:titleInfo>
<mods:name id="B3C568F642F51913844A818E296E0EA1" type="personal">
<mods:role id="82526C5A32989BD0C0548A25D508AF24">
<mods:roleTerm id="7A948AEE6E565F8FB20F9AE94DB6217D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82070D04194FA0E7A491D841A1FB1A3F">Arce-Pérez, Roberto</mods:namePart>
</mods:name>
<mods:name id="CDE2165A71D7098315261C708CD84115" type="personal">
<mods:role id="F5FA1276A2A35B664C302FAB753DD17B">
<mods:roleTerm id="559012074C5C3EB5684E609BB837D6C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E202ED1DA3FB4733672D7C3836C908C0">Morón, Miguel Ángel</mods:namePart>
</mods:name>
<mods:typeOfResource id="0060E655659D08B6ECDA502A77D3F44F">text</mods:typeOfResource>
<mods:relatedItem id="2C483B3566B6DC5927775734AD7949F8" type="host">
<mods:titleInfo id="73A9866245CCCA2C329BEB4394F2E234">
<mods:title id="E43D1872912BE59ECACE5D2576A545DF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="08DA7473C12F4CF5F6E39E1784014F43">
<mods:date id="7DF1ED738E1DE6788A28BB906F7EC67E">2020</mods:date>
<mods:detail id="75D5FAA77CACE833079B4A0F62A6B246" type="pubDate">
<mods:number id="BEAE37F2F7A3316D7BB6FFC8651B744E">2020-05-11</mods:number>
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<mods:detail id="7D1930ECF04A7F3D576D2C77DF02AECE" type="volume">
<mods:number id="57F304B9CA7F8FD534756FBA20375EF2">4772</mods:number>
</mods:detail>
<mods:detail id="42C4A9354CAA5A70AFFF0BB0BD348E2A" type="issue">
<mods:number id="6DBF145E66F49BFC365975C521A67AA6">3</mods:number>
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<mods:start id="852DE3C5A4032EAD7804E88DA47E8341">567</mods:start>
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<mods:classification id="986B115D7A26811F931937DABAF4F5E8">journal article</mods:classification>
<mods:identifier id="7FEB8C1DA6BA4640E4F3CDCF8A27D44D" type="CLB-Dataset">22256</mods:identifier>
<mods:identifier id="4CFCD8280A7CE4456620C7749AEA125E" type="DOI">10.11646/zootaxa.4772.3.7</mods:identifier>
<mods:identifier id="7A47671320968CF4C3C924698F4A940A" type="GBIF-Dataset">8b277405-a91c-4d0c-b77e-8b1d4820c7cd</mods:identifier>
<mods:identifier id="8956998707642E8436FC010C6F3714FD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1707D483EAEFA436F27A03D62C53446B" type="Zenodo-Dep">3819764</mods:identifier>
<mods:identifier id="E6D1AECBC39440CEA2F9E61D42991A4A" type="ZooBank">8DE2F40F-0931-4002-97C4-5603E363B7E8</mods:identifier>
</mods:mods>
<treatment id="6C5796057B7CFFC7FF093F869D692CF9" ID-DOI="http://doi.org/10.5281/zenodo.3843957" ID-GBIF-Taxon="164035917" ID-Zenodo-Dep="3843957" LSID="urn:lsid:plazi:treatment:6C5796057B7CFFC7FF093F869D692CF9" httpUri="http://treatment.plazi.org/id/6C5796057B7CFFC7FF093F869D692CF9" lastPageNumber="569" pageId="2" pageNumber="569">
<subSubSection id="ACE474987B7CFFC7FF093F86986D2AB2" pageId="2" pageNumber="569" type="nomenclature">
<paragraph id="E44127137B7CFFC7FF093F869A792AD6" blockId="2.[151,885,153,216]" box="[151,885,153,180]" pageId="2" pageNumber="569">
<heading id="BF09907F7B7CFFC7FF093F869A792AD6" bold="true" box="[151,885,153,180]" fontSize="11" level="1" pageId="2" pageNumber="569" reason="1">
<emphasis id="D68AFB017B7CFFC7FF093F869A792AD6" bold="true" box="[151,885,153,180]" pageId="2" pageNumber="569">
<taxonomicName id="23FE5C907B7CFFC7FF093F869BD52AD1" authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" authorityYear="2020" box="[151,729,153,180]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="eniocanoi" status="sp. nov.">
<emphasis id="D68AFB017B7CFFC7FF093F8698C52AD6" bold="true" box="[151,457,153,180]" italics="true" pageId="2" pageNumber="569">Macrodactylus eniocanoi</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
,
<taxonomicNameLabel id="CDB9467A7B7CFFC7FD783F859A792AD6" box="[742,885,153,180]" pageId="2" pageNumber="569" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E44127137B7CFFC7FF093FA2986D2AB2" blockId="2.[151,885,153,216]" box="[151,353,189,216]" pageId="2" pageNumber="569">Figs. 1, 3, 5–8, 84</paragraph>
</subSubSection>
<subSubSection id="ACE474987B7CFFC7FF093E1A98B82BD5" pageId="2" pageNumber="569" type="materials_examined">
<paragraph id="E44127137B7CFFC7FF093E1A98B82BD5" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF093E1A98752B45" bold="true" box="[151,377,261,288]" pageId="2" pageNumber="569">Material examined</emphasis>
(
<specimenCount id="F2F8EC9A7B7CFFC7FE163E1A9B5D2B45" box="[392,593,261,288]" pageId="2" pageNumber="569" type="male">12 specimens 9 ♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B7CFFC7FDC33E1A9B8D2B7A" box="[605,641,261,287]" pageId="2" pageNumber="569" type="female">3♀</specimenCount>
):
<typeStatus id="3B4599B17B7CFFC7FD093E1A9BF32B7A" box="[663,767,261,287]" pageId="2" pageNumber="569" type="holotype">Holotype</typeStatus>
male label:
<materialsCitation id="54962D4E7B7CFFC7FC193E1A9B3F2B21" ID-GBIF-Occurrence="2611565304" collectingDate="1993-05-09" collectionCode="UVGC" collectorName="E. Valdez" country="Guatemala" county="San Lorenzo" location="Minas" municipality="Sierra" pageId="2" pageNumber="569" specimenCount="1" stateProvince="Zacapa" typeStatus="holotype">
<emphasis id="D68AFB017B7CFFC7FC193E1A9D072B45" bold="true" box="[903,1035,261,288]" pageId="2" pageNumber="569">
<collectingCountry id="9CE967837B7CFFC7FC193E1A9D072B45" box="[903,1035,261,288]" name="Guatemala" pageId="2" pageNumber="569">Guatemala</collectingCountry>
</emphasis>
:
<collectingRegion id="263AE9F17B7CFFC7FB843E199D622B45" box="[1050,1134,262,288]" country="Guatemala" name="Zacapa" pageId="2" pageNumber="569">Zacapa</collectingRegion>
,
<collectingCounty id="0D205F9F7B7CFFC7FBE63E1A9C042B45" box="[1144,1288,261,288]" pageId="2" pageNumber="569">San Lorenzo</collectingCounty>
,
<collectingMunicipality id="0425BD697B7CFFC7FA8D3E1A9C5A2B7A" box="[1299,1366,261,287]" pageId="2" pageNumber="569">Sierra</collectingMunicipality>
de las
<location id="E12171C87B7CFFC7FF093E3699EC2B26" LSID="urn:lsid:plazi:treatment:6C5796057B7CFFC7FF093F869D692CF9:E12171C87B7CFFC7FF093E3699EC2B26" box="[151,224,297,323]" country="Guatemala" county="San Lorenzo" municipality="Sierra" name="Minas" pageId="2" pageNumber="569" stateProvince="Zacapa">Minas</location>
,
<date id="904001D37B7CFFC7FF723E3698422B26" box="[236,334,297,323]" pageId="2" pageNumber="569" value="1993-05-09">
<collectingDate id="8004F83B7B7CFFC7FF723E3698422B26" box="[236,334,297,323]" pageId="2" pageNumber="569" value="1993-05-09">9.v.1993</collectingDate>
</date>
,
<collectorName id="490B42C57B7CFFC7FEC43E3598C92B26" box="[346,453,297,324]" pageId="2" pageNumber="569">E. Valdez</collectorName>
[
<collectionCode id="82EFBFD67B7CFFC7FE4A3E369B202B21" box="[468,556,297,324]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="2" pageNumber="569">UVGC</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="54962D4E7B7CFFC7FDA03E359BD02B0D" ID-GBIF-Occurrence="2611565325" collectingDate="1994-04-14" collectionCode="UVGC, IEXA" country="Guatemala" elevation="2190" location="San Lorenzo" pageId="2" pageNumber="569" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Zacapa" typeStatus="paratype">
<typeStatus id="3B4599B17B7CFFC7FDA03E359BA72B21" box="[574,683,298,324]" pageId="2" pageNumber="569" type="paratype">Paratypes</typeStatus>
(12):
<emphasis id="D68AFB017B7CFFC7FD6E3E369A782B21" bold="true" box="[752,884,297,324]" pageId="2" pageNumber="569">
<collectingCountry id="9CE967837B7CFFC7FD6E3E369A782B21" box="[752,884,297,324]" name="Guatemala" pageId="2" pageNumber="569">Guatemala</collectingCountry>
</emphasis>
:
<collectingRegion id="263AE9F17B7CFFC7FC1D3E359ADB2B21" box="[899,983,298,324]" country="Guatemala" name="Zacapa" pageId="2" pageNumber="569">Zacapa</collectingRegion>
,
<location id="E12171C87B7CFFC7FC7D3E369D782B21" LSID="urn:lsid:plazi:treatment:6C5796057B7CFFC7FF093F869D692CF9:E12171C87B7CFFC7FC7D3E369D782B21" box="[995,1140,297,324]" country="Guatemala" name="San Lorenzo" pageId="2" pageNumber="569" stateProvince="Zacapa">San Lorenzo</location>
, cerro de los monos,
<quantity id="23068AF67B7CFFC7FAFA3E3699A32B0D" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="2" pageNumber="569" unit="m" value="2190.0">
<elevation id="6FD3C0207B7CFFC7FAFA3E3699A32B0D" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="2" pageNumber="569" unit="m" value="2190.0">2190 m</elevation>
</quantity>
,
<date id="904001D37B7CFFC7FF243E52983E2B02" box="[186,306,333,359]" pageId="2" pageNumber="569" value="1994-04-14">
<collectingDate id="8004F83B7B7CFFC7FF243E52983E2B02" box="[186,306,333,359]" pageId="2" pageNumber="569" value="1994-04-14">14.iv.1994</collectingDate>
</date>
, (
<specimenCount id="F2F8EC9A7B7CFFC7FED83E5298672B02" box="[326,363,333,359]" pageId="2" pageNumber="569" type="male">2♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B7CFFC7FEE83E52989B2B02" box="[374,407,333,359]" pageId="2" pageNumber="569" type="female">1♀</specimenCount>
) [
<collectionCode id="82EFBFD67B7CFFC7FE313E529B0A2B0D" box="[431,518,333,360]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="2" pageNumber="569">UVGC</collectionCode>
], (
<specimenCount id="F2F8EC9A7B7CFFC7FDBF3E529B4A2B02" box="[545,582,333,359]" pageId="2" pageNumber="569" type="male">1♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B7CFFC7FDCE3E529B7D2B02" box="[592,625,333,359]" pageId="2" pageNumber="569" type="female">1♀</specimenCount>
) [
<collectionCode id="82EFBFD67B7CFFC7FD143E529BD92B0D" box="[650,725,333,360]" pageId="2" pageNumber="569">IEXA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="54962D4E7B7CFFC7FD783E5198F72BE9" ID-GBIF-Occurrence="2611565322" collectingDate="1993-06-07" collectionCode="PKLC" country="Guatemala" county="Sierra" elevation="2100" location="San Lorenzo" municipality="Minas" pageId="2" pageNumber="569" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Zacapa" typeStatus="paratype">
<collectingRegion id="263AE9F17B7CFFC7FD783E519A362B0D" box="[742,826,334,360]" country="Guatemala" name="Zacapa" pageId="2" pageNumber="569">Zacapa</collectingRegion>
,
<collectingCounty id="0D205F9F7B7CFFC7FCDA3E529A8B2B02" box="[836,903,333,359]" pageId="2" pageNumber="569">Sierra</collectingCounty>
de las
<collectingMunicipality id="0425BD697B7CFFC7FC4C3E529D172B02" box="[978,1051,333,359]" pageId="2" pageNumber="569">Minas</collectingMunicipality>
,
<quantity id="23068AF67B7CFFC7FBBB3E529D512B02" box="[1061,1117,333,360]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="569" unit="km" value="5.0">5 km</quantity>
NE from
<location id="E12171C87B7CFFC7FB543E529C562B0D" LSID="urn:lsid:plazi:treatment:6C5796057B7CFFC7FF093F869D692CF9:E12171C87B7CFFC7FB543E529C562B0D" box="[1226,1370,333,360]" country="Guatemala" county="Sierra" municipality="Minas" name="San Lorenzo" pageId="2" pageNumber="569" stateProvince="Zacapa">San Lorenzo</location>
,
<quantity id="23068AF67B7CFFC7FAFA3E5299A32BE9" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="569" unit="m" value="2100.0">
<elevation id="6FD3C0207B7CFFC7FAFA3E5299A32BE9" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="569" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="904001D37B7CFFC7FF243E6E982A2BEE" box="[186,294,369,395]" pageId="2" pageNumber="569" value="1993-06-07">
<collectingDate id="8004F83B7B7CFFC7FF243E6E982A2BEE" box="[186,294,369,395]" pageId="2" pageNumber="569" value="1993-06-07">7.vi.1993</collectingDate>
</date>
, (
<specimenCount id="F2F8EC9A7B7CFFC7FEA43E6E98532BE9" box="[314,351,369,396]" pageId="2" pageNumber="569" type="male">5♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B7CFFC7FEF43E6E98872BEE" box="[362,395,369,395]" pageId="2" pageNumber="569" type="female">1♀</specimenCount>
) [
<collectionCode id="82EFBFD67B7CFFC7FE3D3E6E98FF2BE9" box="[419,499,369,396]" pageId="2" pageNumber="569">PKLC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="54962D4E7B7CFFC7FD9B3E6D98BD2BD5" ID-GBIF-Occurrence="2611565302" collectingDate="1993-06-17" collectionCode="CMNC" collectorName="J. M. Campbell" country="Guatemala" county="Sierra" elevation="2100" location="San Lorenzo" municipality="Minas" pageId="2" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="Zacapa" typeStatus="paratype">
<collectingRegion id="263AE9F17B7CFFC7FD9B3E6D9B552BE9" box="[517,601,370,396]" country="Guatemala" name="Zacapa" pageId="2" pageNumber="569">Zacapa</collectingRegion>
,
<collectingCounty id="0D205F9F7B7CFFC7FDFD3E6E9BAA2BEE" box="[611,678,369,395]" pageId="2" pageNumber="569">Sierra</collectingCounty>
de las
<collectingMunicipality id="0425BD697B7CFFC7FD6F3E6E9A362BEE" box="[753,826,369,395]" pageId="2" pageNumber="569">Minas</collectingMunicipality>
,
<quantity id="23068AF67B7CFFC7FCDA3E6E9A702BEE" box="[836,892,369,396]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="569" unit="km" value="5.0">5 km</quantity>
NE from
<location id="E12171C87B7CFFC7FC743E6E9D762BE9" LSID="urn:lsid:plazi:treatment:6C5796057B7CFFC7FF093F869D692CF9:E12171C87B7CFFC7FC743E6E9D762BE9" box="[1002,1146,369,396]" country="Guatemala" county="Sierra" municipality="Minas" name="San Lorenzo" pageId="2" pageNumber="569" stateProvince="Zacapa">San Lorenzo</location>
,
<quantity id="23068AF67B7CFFC7FB1B3E6E9DD72BE9" box="[1157,1243,369,396]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="569" unit="m" value="2100.0">
<elevation id="6FD3C0207B7CFFC7FB1B3E6E9DD72BE9" box="[1157,1243,369,396]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="569" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="904001D37B7CFFC7FB783E6E9C6C2BEE" box="[1254,1376,369,395]" pageId="2" pageNumber="569" value="1993-06-17">
<collectingDate id="8004F83B7B7CFFC7FB783E6E9C6C2BEE" box="[1254,1376,369,395]" pageId="2" pageNumber="569" value="1993-06-17">17.vi.1993</collectingDate>
</date>
,
<collectorName id="490B42C57B7CFFC7FAF43E6D98082BCA" pageId="2" pageNumber="569">J.M. Campbell</collectorName>
(
<specimenCount id="F2F8EC9A7B7CFFC7FE883E8A983A2BCA" box="[278,310,405,431]" pageId="2" pageNumber="569" type="female">1♀</specimenCount>
) [
<collectionCode id="82EFBFD67B7CFFC7FED03E8A98A52BD5" box="[334,425,405,432]" pageId="2" pageNumber="569">CMNC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ACE474987B7CFFC7FF593EA69AB92806" pageId="2" pageNumber="569" type="diagnosis">
<paragraph id="E44127137B7CFFC7FF593EA69AB92806" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593EA6984E2BB1" bold="true" box="[199,322,441,468]" pageId="2" pageNumber="569">Diagnosis.</emphasis>
Integument of the body glossy black with yellow-green iridescence; elytra and legs reddish yellow; pronotum, elytra, and pygidium covered with pale yellow setae; venter with vestiture long and dense (Figs. 1, 3); profemur with a subtriangular projection near the base, elongate and twisted near the acute apex, followed by an obvious depression; tarsi with rings of long, white setae. Female pronotum wider than long, the posterior angles acute and extended laterally, and with poorly defined glabrous midline.
</paragraph>
</subSubSection>
<subSubSection id="ACE474987B7CFFC7FF593D729A9D2C69" pageId="2" pageNumber="569" type="description">
<paragraph id="E44127137B7CFFC7FF593D7298152EC1" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593D729B0228ED" bold="true" box="[199,526,621,648]" pageId="2" pageNumber="569">
Description.
<typeStatus id="3B4599B17B7CFFC7FEC03D7198C728ED" box="[350,459,622,648]" pageId="2" pageNumber="569" type="holotype">Holotype</typeStatus>
Male
</emphasis>
: length 9.5 mm; humeral width
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.
<emphasis id="D68AFB017B7CFFC7FC223D729AF128E2" box="[956,1021,621,647]" italics="true" pageId="2" pageNumber="569">Color</emphasis>
: integument of the body black glossy with yellow-green iridescence; antennae bicolored; clypeus, elytra, and legs reddish yellow (apex of tibiae and tarsi darker); head, pronotum, scutellum, and pygidium black with yellow-green iridescence.
<emphasis id="D68AFB017B7CFFC7FBE83DA99DC028B5" box="[1142,1228,694,720]" italics="true" pageId="2" pageNumber="569">Surface</emphasis>
: pronotum, elytra, and pygidium covered by pale yellow vestiture; venter with vestiture long and dense (Figs. 1, 3).
<emphasis id="D68AFB017B7CFFC7FB743DC69C2B2896" box="[1258,1319,729,755]" italics="true" pageId="2" pageNumber="569">Head</emphasis>
: elongate, with a longitudinal ridge and a lateral depression, coarsely punctate; clypeus trapezoidal, anterior border nearly straight.
<emphasis id="D68AFB017B7CFFC7FF643C3E9847295E" box="[250,331,801,827]" italics="true" pageId="2" pageNumber="569">Thorax</emphasis>
: pronotum hexagonal, convex, length 2.65 mm, width 2.60; prosternal process long, slender, subtriangular, curved, apex acute (easily to see in frontal view); scutellum elongate with rounded apex. Elytra reddish yellow, vestiture pale yellow. Metasternum with two longitudinal rows of long, yellow setae between mesocoxae and metacoxae.
<emphasis id="D68AFB017B7CFFC7FED53C91988C29C2" box="[331,384,910,935]" italics="true" pageId="2" pageNumber="569">Legs</emphasis>
: Profemur short, with a subtriangular projection near to the base, elongate and twisted near the apex, followed by an obvious depression. Mesotibia widened towards apex with two short, straight spurs and long spine-like setae around the apex. Tarsi with rings of long, white setae; all tarsal claws cleft.
<emphasis id="D68AFB017B7CFFC7FB163CCA9DFE298A" box="[1160,1266,981,1007]" italics="true" pageId="2" pageNumber="569">Abdomen</emphasis>
: Sternites II–V moderately concave at middle, with five to six long yellow setae on each side of midline; pygidial plate elongate, oval with vestiture equal to the ventral.
<emphasis id="D68AFB017B7CFFC7FDC33B029BC32E52" box="[605,719,1053,1079]" italics="true" pageId="2" pageNumber="569">Genitalia:</emphasis>
total length 2.38 mm (Fig. 7); parameres short, oval, glabrous, fused dorsobasally, with a distinct dorsobasal internal emargination; widest mid-basally, distal half bent medially, apices rounded, forming an oval in distal view, length of parameres 0.80 mm, width across fused structure 0.70 mm (Figs. 5–6).
</paragraph>
<paragraph id="E44127137B7CFFC7FF593BB299FE2F32" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593BB298862EAD" bold="true" box="[199,394,1197,1224]" pageId="2" pageNumber="569">
<typeStatus id="3B4599B17B7CFFC7FF593BB298202EAD" box="[199,300,1197,1224]" pageId="2" pageNumber="569" type="allotype">Allotype</typeStatus>
Female
</emphasis>
: length
<quantity id="23068AF67B7CFFC7FE793BB29B352EAD" box="[487,569,1197,1224]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="569" unit="mm" value="10.0">10 mm</quantity>
; humeral width 3.5 mm. Similar to male except: vestiture is longer and erect; pronotal length 2.25 mm, width 2.50 mm, with poorly defined, glabrous midline (longitudinal stripe) and the posterior angles acute and extended laterally; abdominal sternites II–IV convex; pygidium short and nearly triangular. Genital plates short oval, length 0.82 mm, width 0.28 mm, apex asymmetrical, and external border nearly straight (Fig. 8).
</paragraph>
<paragraph id="E44127137B7CFFC7FF593A7E98762FFA" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593A7E98332F19" bold="true" box="[199,319,1377,1404]" pageId="2" pageNumber="569">Variation.</emphasis>
Body length 9.5–10.0 mm; width across humeri 3.0–3.5 mm, apex of the tarsi and tibiae in some specimens are paler.
</paragraph>
<paragraph id="E44127137B7CFFC7FF593AB69D172FA6" blockId="2.[151,1437,261,1692]" box="[199,1051,1449,1476]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593AB69B092FA1" bold="true" box="[199,517,1449,1476]" pageId="2" pageNumber="569">Geographical distribution.</emphasis>
Known only from eastern
<collectingCountry id="9CE967837B7CFFC7FCAF3AB69AA62FA6" box="[817,938,1449,1475]" name="Guatemala" pageId="2" pageNumber="569">Guatemala</collectingCountry>
(Fig. 84).
</paragraph>
<paragraph id="E44127137B7CFFC7FF593AD19A9D2C69" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF593AD1988B2F8D" bold="true" box="[199,391,1485,1512]" pageId="2" pageNumber="569">Natural history.</emphasis>
Three males and two females were collected as pupae in mountain forest at an elevation of
<quantity id="23068AF67B7CFFC7FF093AEE99FF2C69" box="[151,243,1521,1548]" metricMagnitude="0" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="569" unit="m" value="2.1">2,100 m</quantity>
on
<date id="904001D37B7CFFC7FE833AEE98B02C6E" box="[285,444,1521,1547]" pageId="2" pageNumber="569" value="1994-04-10">10 April 1994</date>
, and the adults emerged in May and June.
</paragraph>
</subSubSection>
<subSubSection id="ACE474987B7CFFC7FF5939099D692CF9" pageId="2" pageNumber="569" type="etymology">
<paragraph id="E44127137B7CFFC7FF59390998172C36" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF59390998422C55" bold="true" box="[199,334,1558,1584]" pageId="2" pageNumber="569">Etymology.</emphasis>
Dedicated to Enio Cano, enthusiastic scarabaeidologist, who provided the original material for the description.
</paragraph>
<paragraph id="E44127137B7CFFC7FF5939429D692CF9" blockId="2.[151,1437,261,1692]" pageId="2" pageNumber="569">
<emphasis id="D68AFB017B7CFFC7FF59394298DD2C1D" bold="true" box="[199,465,1629,1656]" pageId="2" pageNumber="569">Taxonomic comments.</emphasis>
<taxonomicName id="23FE5C907B7CFFC7FE4939429BFE2C12" authorityName="Arce-Perez &amp; Moron" authorityYear="2020" box="[471,754,1629,1655]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="569" phylum="Arthropoda" rank="species" species="eniocanoi">
<emphasis id="D68AFB017B7CFFC7FE4939429BFE2C12" box="[471,754,1629,1655]" italics="true" pageId="2" pageNumber="569">Macrodactylus eniocanoi</emphasis>
</taxonomicName>
is placed in a new species group (that will be described later) characterized by an acute triangular projection of the profemora and falcate parameres.
</paragraph>
</subSubSection>
</treatment>
</document>
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