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<document id="6F5ABDB78E2760E8E8B25FF8EB44318E" ID-ISSN="0024-4082" ID-ZooBank="74C07292-2BD6-4E3E-B68D-B144B81BBD83" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629585783401" checkinUser="felipe" docAuthor="Bonifácio, Paulo &amp; Menot, Lénaïck" docDate="2019" docId="0B1B8791FFE60636F8D5E8DE7FBB5933" docLanguage="en" docName="ZoolJLinnSoc.185.555–635.pdf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Bathyeliasona mariaae Bonifácio &amp; Menot 2019, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="585" masterDocId="F722FFE9FFFA0628F820E81A7D025E58" masterDocTitle="New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)" masterLastPageNumber="635" masterPageNumber="555" pageNumber="583" updateTime="1699081480826" updateUser="plazi">
<mods:mods id="4740FBED3A2EA1B5FA56C67CA5EE1CC7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D18A5CF5A01B11BFD6A06FF30CEA2F49">
<mods:title id="31468E31691CA25B5CDBF13C5A33C502">New genera and species from the Equatorial Pacific provide phylogenetic insights into deep-sea Polynoidae (Annelida)</mods:title>
</mods:titleInfo>
<mods:name id="64867D2C5D68F36DBE77903D17E8AD32" type="personal">
<mods:role id="13FD6DFE6BF66FD753D4CD8A880B4AD6">
<mods:roleTerm id="D66C898BD7EB843ED3F075A619B17C66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98C3A5B3F289970DA258DA0CA6244CB4">Bonifácio, Paulo</mods:namePart>
</mods:name>
<mods:name id="6AEE87A109D1B0AE2A01F44B45ABFB3B" type="personal">
<mods:role id="2898E6CF5D1AD35DA6FE5FF508FBBE3E">
<mods:roleTerm id="547C0569FA43DA2754384A2AC989BD73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0400C92E5F5C56DBAD8FAFEE90282825">Menot, Lénaïck</mods:namePart>
</mods:name>
<mods:typeOfResource id="E823DDF59DDB4F095A1186A8C6CBF602">text</mods:typeOfResource>
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<mods:titleInfo id="C116C5C20FFC0CF061370FEEDE4E0B4A">
<mods:title id="05F28F5ED9D8D0484815FA58C75E7B5D">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="28FACFAE01F4E642B3BE8734FAE5FEE1">
<mods:date id="10ED65CC23B8CD11E16C913F660B184A">2019</mods:date>
<mods:detail id="59D95B927CF363FC07548CEC481891DC" type="volume">
<mods:number id="E990CC2F83C488A05F0B63BB7F0427A0">185</mods:number>
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<mods:extent id="6ED11F7F53DDFCCE2A5B25CF81EA4E9E" unit="page">
<mods:start id="09589BA46384475B4B5BE95BE5575EC3">555</mods:start>
<mods:end id="18743CA2E1E47A6A7C08C1D00AB4E125">635</mods:end>
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<mods:classification id="94DB3804F981CA07CCC652E9C934889E">journal article</mods:classification>
<mods:identifier id="69DB90C8F302A629D46425F6741C66E5" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="F915A617DD859B65E7A092129F44449E" type="ZooBank">74C07292-2BD6-4E3E-B68D-B144B81BBD83</mods:identifier>
</mods:mods>
<treatment id="0B1B8791FFE60636F8D5E8DE7FBB5933" LSID="urn:lsid:plazi:treatment:0B1B8791FFE60636F8D5E8DE7FBB5933" httpUri="http://treatment.plazi.org/id/0B1B8791FFE60636F8D5E8DE7FBB5933" lastPageId="30" lastPageNumber="585" pageId="28" pageNumber="583">
<subSubSection id="CBA8650CFFE60634F8D5E8DE7F965E84" box="[245,660,196,220]" pageId="28" pageNumber="583" type="nomenclature">
<paragraph id="830D3687FFE60634F8D5E8DE7F965E84" blockId="28.[245,660,196,220]" box="[245,660,196,220]" pageId="28" pageNumber="583">
<heading id="D84581EBFFE60634F8D5E8DE7F965E84" box="[245,660,196,220]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="583" reason="2">
<taxonomicName id="44B24D04FFE60634F8D5E8DE7F285E82" ID-CoL="KWZ3" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[245,554,196,219]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="28" pageNumber="627" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">BATHYELIASONA MARIAAE</taxonomicName>
<emphasis id="B1C6EA95FFE60634FA11E8DD7F965E84" bold="true" box="[561,660,196,220]" pageId="28" pageNumber="583">
<taxonomicNameLabel id="AAF557EEFFE60634FA11E8DD7F965E84" box="[561,660,196,220]" pageId="28" pageNumber="583" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CBA8650CFFE60636F919E8F17FBB5933" lastPageId="30" lastPageNumber="585" pageId="28" pageNumber="583" type="description">
<paragraph id="830D3687FFE60634F919E8F17F525F5C" blockId="28.[313,592,235,260]" box="[313,592,235,260]" pageId="28" pageNumber="583">
(
<figureCitation id="1B892A02FFE60634F961E8F17F155F5A" box="[321,535,235,260]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">FIG. 7A–J; TABLES</figureCitation>
1, 2)
</paragraph>
<paragraph id="830D3687FFE60634F8B1E90E7DF55CA2" blockId="28.[145,762,275,762]" pageId="28" pageNumber="583">
<emphasis id="B1C6EA95FFE60634F8B1E90E7C495F71" box="[145,331,276,297]" italics="true" pageId="28" pageNumber="583">Type material:</emphasis>
<materialsCitation id="33DA3CDAFFE60634F979E90E7F795C5A" collectingDate="2015-03-26" collectionCode="BGR" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="11.829267" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-117.4929" municipality="Eastern Pacific Ocean" pageId="28" pageNumber="583" specimenCode="IFR107" specimenCount="1" typeStatus="holotype">
<typeStatus id="5C098825FFE60634F979E90E7CCE5F72" box="[345,460,276,298]" pageId="28" pageNumber="583" type="holotype">Holotype</typeStatus>
, MNHN-IA-TYPE 1815 (
<specimenCode id="D3149EFCFFE60634F8B9E9297DF25F11" box="[153,240,307,329]" collectionCode="IFR" pageId="28" pageNumber="583">IFR107</specimenCode>
), complete, length
<quantity id="444A9B62FFE60634F99FE9287F205F10" box="[447,546,306,328]" metricMagnitude="-3" metricUnit="m" metricValue="9.86" pageId="28" pageNumber="583" unit="mm" value="9.86">9.86 mm</quantity>
, width
<quantity id="444A9B62FFE60634FA52E9297FD75F10" box="[626,725,307,328]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="28" pageNumber="583" unit="mm" value="1.88">1.88 mm</quantity>
, 17 segments (including tentacular segment),
<location id="866D605CFFE60634FA5CE94B7C995FDE" LSID="urn:lsid:plazi:treatment:0B1B8791FFE60636F8D5E8DE7FBB5933:866D605CFFE60634FA5CE94B7C995FDE" county="Clarion-Clipperton Fracture Zone" latitude="11.829267" longLatPrecision="1" longitude="-117.4929" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="28" pageNumber="583">
Equatorial
<collectingMunicipality id="6369ACFDFFE60634F8B1E96B7C995FDE" box="[145,411,368,390]" pageId="28" pageNumber="583">Eastern Pacific Ocean</collectingMunicipality>
</location>
,
<collectingCounty id="6A6C4E0BFFE60634F988E96A7DCB5FFD" pageId="28" pageNumber="583">Clarion-Clipperton Fracture Zone</collectingCounty>
,
<collectionCode id="E5A3AE42FFE60634F8F4E98A7C095FFD" box="[212,267,400,421]" country="Germany" httpUri="http://grbio.org/cool/r4mz-yr8h" name="Bundesanstalt fur Geowissenschaften und Rohstoffe" pageId="28" pageNumber="583">BGR</collectionCode>
license area, station 50, collected
<date id="F70C1047FFE60634FAADE9957DC95F9C" pageId="28" pageNumber="583" value="2015-03-26">
<collectingDate id="E748E9AFFFE60634FAADE9957DC95F9C" pageId="28" pageNumber="583" value="2015-03-26">26 March 2015</collectingDate>
</date>
, epibenthic sledge supra-net, start
<geoCoordinate id="E6865040FFE60634FA46E9B47FF45F9C" box="[614,758,429,452]" degrees="11" direction="north" minutes="49.592" orientation="latitude" pageId="28" pageNumber="583" precision="1" value="11.826533">11°49.592′N</geoCoordinate>
,
<geoCoordinate id="E6865040FFE60634F8B1E9D77C325FBB" box="[145,304,460,483]" degrees="117" direction="west" minutes="30.786" orientation="longitude" pageId="28" pageNumber="583" precision="1" value="-117.5131">117°30.786′W</geoCoordinate>
, end
<geoCoordinate id="E6865040FFE60634F94EE9D77CFE5FBB" box="[366,508,460,483]" degrees="11" direction="north" minutes="49.756" orientation="latitude" pageId="28" pageNumber="583" precision="1" value="11.829267">11°49.756′N</geoCoordinate>
,
<geoCoordinate id="E6865040FFE60634FA28E9D77FAA5FBB" box="[520,680,460,483]" degrees="117" direction="west" minutes="29.574" orientation="longitude" pageId="28" pageNumber="583" precision="1" value="-117.4929">117°29.574′W</geoCoordinate>
, 4360–
<elevation id="089FD1B4FFE60634F8B1E9F67C9F5C5A" box="[145,413,492,514]" isEnumeration="true" pageId="28" pageNumber="583">
<quantity id="444A9B62FFE60634F8B1E9F67DEB5C5A" box="[145,233,492,514]" metricMagnitude="3" metricUnit="m" metricValue="4.328" pageId="28" pageNumber="583" unit="m" value="4328.0">4328 m</quantity>
depth,
<quantity id="444A9B62FFE60634F965E9F67C9F5C5A" box="[325,413,492,514]" metricMagnitude="3" metricUnit="m" metricValue="2.469" pageId="28" pageNumber="583" unit="m" value="2469.0">2469 m</quantity>
</elevation>
trawling distance
</materialsCitation>
.
<materialsCitation id="33DA3CDAFFE60634FAA7E9F77DF15CA2" collectingDate="2015-04-22" collectionCode="IFR, APEI" county="Clarion-Clipperton Fracture Zone" elevation="0" latitude="18.818134" location="Equatorial Eastern Pacific Ocean" longLatPrecision="1" longitude="-128.35481" municipality="Eastern Pacific Ocean" pageId="28" pageNumber="583" specimenCode="IFR666-4, APEI#3" specimenCount="1" typeStatus="paratype">
<typeStatus id="5C098825FFE60634FAA7E9F77FF65C5A" box="[647,756,493,514]" pageId="28" pageNumber="583" type="paratype">Paratype</typeStatus>
, MNHN-IA-TYPE 1816 (
<specimenCode id="D3149EFCFFE60634F98DEA117F225C79" box="[429,544,523,545]" collectionCode="IFR" pageId="28" pageNumber="583">IFR666-4</specimenCode>
), complete, length
<quantity id="444A9B62FFE60634F8B1EA307DFC5C18" box="[145,254,554,576]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="28" pageNumber="583" unit="mm" value="2.9">2.90 mm</quantity>
, width
<quantity id="444A9B62FFE60634F97DEA307CC95C18" box="[349,459,554,576]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="28" pageNumber="583" unit="mm" value="0.44">0.44 mm</quantity>
, 15 segments (including tentacular segment),
<location id="866D605CFFE60634F95EEA537FF75C06" LSID="urn:lsid:plazi:treatment:0B1B8791FFE60636F8D5E8DE7FBB5933:866D605CFFE60634F95EEA537FF75C06" box="[382,757,585,607]" county="Clarion-Clipperton Fracture Zone" latitude="18.818134" longLatPrecision="1" longitude="-128.35481" municipality="Eastern Pacific Ocean" name="Equatorial Eastern Pacific Ocean" pageId="28" pageNumber="583">
Equatorial
<collectingMunicipality id="6369ACFDFFE60634F9DDEA507FF75C06" box="[509,757,585,607]" pageId="28" pageNumber="583">Eastern Pacific Ocean</collectingMunicipality>
</location>
,
<collectingCounty id="6A6C4E0BFFE60634F8B1EA727F295C26" box="[145,555,616,638]" pageId="28" pageNumber="583">Clarion-Clipperton Fracture Zone</collectingCounty>
,
<specimenCode id="D3149EFCFFE60634FA18EA727F955C25" box="[568,663,616,637]" collectionCode="APEI" country="Canada" httpUri="http://grbio.org/cool/pg9t-riai" name="Agriculture Canada Research Station" pageId="28" pageNumber="583">APEI#3</specimenCode>
, station 197, collected
<date id="F70C1047FFE60634F910EA9D7CCE5CC5" box="[304,460,647,669]" pageId="28" pageNumber="583" value="2015-04-22">
<collectingDate id="E748E9AFFFE60634F910EA9D7CCE5CC5" box="[304,460,647,669]" pageId="28" pageNumber="583" value="2015-04-22">22 April 2015</collectingDate>
</date>
, epibenthic sledge epi-net, start
<geoCoordinate id="E6865040FFE60634F8F5EABC7C6B5CE3" box="[213,361,677,700]" degrees="18" direction="north" minutes="48.659" orientation="latitude" pageId="28" pageNumber="583" precision="1" value="18.810984">18°48.659′N</geoCoordinate>
,
<geoCoordinate id="E6865040FFE60634F958EABC7F1F5CE3" box="[376,541,677,700]" degrees="128" direction="west" minutes="22.753" orientation="longitude" pageId="28" pageNumber="583" precision="1" value="-128.37921">128°22.753′W</geoCoordinate>
, end
<geoCoordinate id="E6865040FFE60634FA40EABC7FF75CE3" box="[608,757,677,700]" degrees="18" direction="north" minutes="49.088" orientation="latitude" pageId="28" pageNumber="583" precision="1" value="18.818134">18°49.088′N</geoCoordinate>
,
<geoCoordinate id="E6865040FFE60634F8B1EADF7C305C82" box="[145,306,708,731]" degrees="128" direction="west" minutes="21.289" orientation="longitude" pageId="28" pageNumber="583" precision="1" value="-128.35481">128°21.289′W</geoCoordinate>
,
<elevation id="089FD1B4FFE60634F91EEADF7F8E5C82" box="[318,652,709,731]" isEnumeration="true" pageId="28" pageNumber="583">
<quantity id="444A9B62FFE60634F91EEADF7CD95C82" box="[318,475,709,731]" metricMagnitude="3" metricUnit="m" metricValue="4.814" metricValueMax="4.823" metricValueMin="4.805" pageId="28" pageNumber="583" unit="m" value="4814.0" valueMax="4823.0" valueMin="4805.0">4805–4823 m</quantity>
depth,
<quantity id="444A9B62FFE60634FA16EADF7F8E5C82" box="[566,652,709,731]" metricMagnitude="3" metricUnit="m" metricValue="2.529" pageId="28" pageNumber="583" unit="m" value="2529.0">2529 m</quantity>
</elevation>
trawling distance
</materialsCitation>
.
</paragraph>
<paragraph id="830D3687FFE60634F8B1EB3E7CAB5A4A" blockId="28.[145,762,803,1877]" pageId="28" pageNumber="583">
<emphasis id="B1C6EA95FFE60634F8B1EB3E7F855D60" box="[145,647,803,825]" italics="true" pageId="28" pageNumber="583">
Description (based on
<typeStatus id="5C098825FFE60634F9A2EB397CE25D60" box="[386,480,803,824]" pageId="28" pageNumber="583" type="holotype">holotype</typeStatus>
and
<typeStatus id="5C098825FFE60634FA34EB397F7B5D60" box="[532,633,803,824]" pageId="28" pageNumber="583" type="paratype">paratype</typeStatus>
):
</emphasis>
<typeStatus id="5C098825FFE60634FAB2EB397FFA5D61" box="[658,760,803,825]" pageId="28" pageNumber="583" type="holotype">Holotype</typeStatus>
complete,
<quantity id="444A9B62FFE60634F936EB587C855D0F" box="[278,391,834,856]" metricMagnitude="-3" metricUnit="m" metricValue="9.86" pageId="28" pageNumber="583" unit="mm" value="9.86">9.86 mm</quantity>
long and
<quantity id="444A9B62FFE60634FA31EB587F835D0F" box="[529,641,834,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="28" pageNumber="583" unit="mm" value="1.88">1.88 mm</quantity>
wide for 17 segments (including tentacular segment), dorsoventrally slightly flattened, not tapering posteriorly; live specimen iridescent, purplish in colour dorso-anteriorly, prostomium whitish (
<figureCitation id="1B892A02FFE60634FA4EEBA47FEB5D8B" box="[622,745,958,980]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7A, B</figureCitation>
); ethanol-preserved specimen pale yellow, prostomium whitish; golden aciculae.
</paragraph>
<paragraph id="830D3687FFE60634F889EC017C915BDD" blockId="28.[145,762,803,1877]" pageId="28" pageNumber="583">
Prostomium bilobed, about as wide as long, lobes pronounced, anteriorly extending into slender, long frontal filaments; median notch between prostomial lobes moderately wide and deep; eyes absent; a pair of comma-shaped, pigmented nuchal organs in a slight depression present on mid anterior prostomial lobes (
<figureCitation id="1B892A02FFE60634F8B9ECCE7C045AB1" box="[153,262,1236,1258]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7B, C</figureCitation>
). Median antenna present, lateral antennae absent; ceratophore of median antenna cylindrical, long (not surpassing anterior end of frontal filaments), inserted in median notch, style missing. Palps smooth, tapering, very long (reaching to segment 8;
<figureCitation id="1B892A02FFE60634FAB5ED4A7FE85B3D" box="[661,746,1360,1382]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7C</figureCitation>
). Facial tubercle absent.
</paragraph>
<paragraph id="830D3687FFE60634F889ED947C20590D" blockId="28.[145,762,803,1877]" pageId="28" pageNumber="583">
<materialsCitation id="33DA3CDAFFE60634F889ED947C1C590D" collectionCode="TYPE" pageId="28" pageNumber="583" specimenCode="TYPE 1816" specimenCount="3" typeStatus="holotype">
Tentacular segment with elongate acicular lobe, inserted laterally and slightly ventral to prostomium; with acicula not penetrating epidermis, with chaetae; tentaculophores small, cylindrical, equal sized, insertion arranged horizontally, inserted distally; tentacular styles papillated, tapering, long (reaching segment 5), thin; dorsal tentacular style (innermost, now lost in
<typeStatus id="5C098825FFE60634F904EE7D7C8F5825" box="[292,397,1639,1661]" pageId="28" pageNumber="583" type="holotype">holotype</typeStatus>
) slightly longer than ventral (outermost) tentacular style (
<figureCitation id="1B892A02FFE60634F9D5EE9C7F4958C3" box="[501,587,1670,1692]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7C</figureCitation>
). Pharynx not everted in
<typeStatus id="5C098825FFE60634F92AEEBF7C6F58E3" box="[266,365,1701,1723]" pageId="28" pageNumber="583" type="holotype">holotype</typeStatus>
; dissected in
<typeStatus id="5C098825FFE60634FA24EEBF7F6858E2" box="[516,618,1701,1722]" pageId="28" pageNumber="583" type="paratype">paratype</typeStatus>
(MNHN-IA-
<specimenCode id="D3149EFCFFE60634F8B1EEDE7C155882" box="[145,279,1732,1754]" collectionCode="TYPE" pageId="28" pageNumber="583">TYPE 1816</specimenCode>
) with nine pairs of subtriangular, equalsized distal papillae; two pairs of jaws, each with main fang, margin smooth (
<figureCitation id="1B892A02FFE60634F9B3EF187CEA594F" box="[403,488,1794,1816]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7D</figureCitation>
). Second segment with elytrophores, subbiramous parapodia, chaetae and ventral cirri
</materialsCitation>
.
</paragraph>
<paragraph id="830D3687FFE60634FB61E8DF7E935F40" blockId="28.[809,1426,197,1414]" pageId="28" pageNumber="583">Eight pairs of distinct, knob-like elytrophores present on segments 2, 4, 5, 7, 9, 11, 13 and 15 (all elytra missing).</paragraph>
<paragraph id="830D3687FFE60634FB61E93B78605F2C" blockId="28.[809,1426,197,1414]" pageId="28" pageNumber="583">
Cirrigerous segments with distinct, cylindrical dorsal cirrophores (
<figureCitation id="1B892A02FFE60634FC33E92579655F0D" box="[1043,1127,319,341]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7F</figureCitation>
), inserted subdistally on notopodia; styles missing; dorsal tubercles absent.
</paragraph>
<paragraph id="830D3687FFE60634FB61E967783C5C12" blockId="28.[809,1426,197,1414]" pageId="28" pageNumber="583">
Ventral cirri smooth, tapering, present from segment 2 to last segment; inserted basally on neuropodia of segment 2, style short (much shorter than tip of neuroacicular lobe); in subsequent segments inserted subdistally on neuropodia (
<figureCitation id="1B892A02FFE60634FCA7E9ED79E25C55" box="[1159,1248,503,525]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7F</figureCitation>
), styles short (approaching tip of neuroacicular lobe); in last segment, style longer than in preceding segments.
</paragraph>
<paragraph id="830D3687FFE60634FB61EA49780B5B72" blockId="28.[809,1426,197,1414]" pageId="28" pageNumber="583">
Parapodia subbiramous; notopodia shorter than neuropodia (
<figureCitation id="1B892A02FFE60634FBAFEA687EE25CDF" box="[911,992,626,648]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7F</figureCitation>
). Notopodia arising from the dorsum, as two thickened ridges; notopodia subtriangular, tapering into long acicular lobe, tip of notoacicula not penetrating epidermis. Neuropodia large, rectangular to subtriangular, tapering into long acicular lobe, tip of neuroacicula not penetrating epidermis. The last pair of parapodia similar to preceding ones. Notochaetae of
<specimenCount id="95B4FD0EFFE60634FB65EB537EB75D06" box="[837,949,841,862]" pageId="28" pageNumber="583" type="generic" typeStatus="types">two types</specimenCount>
: (1) very few (four observed), short, stout, slightly curved with distinct spinous rows on convex side, with blunt tips (
<figureCitation id="1B892A02FFE60634FC10EB9C798A5DC3" box="[1072,1160,902,924]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7G</figureCitation>
); and (2) moderate in number (12 observed), long to very long, slender, slightly curved with distinct, well-developed, clear spinous rows, occasionally one of those notochaetae can be stouter and straight (
<figureCitation id="1B892A02FFE60634FC53EC1A79CA5A4E" box="[1139,1224,1024,1046]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7H</figureCitation>
), with blunt tips; notochaetae stouter and narrower than neurochaetae. Neurochaetae of
<specimenCount id="95B4FD0EFFE60634FBD0EC2479605A0B" box="[1008,1122,1086,1107]" pageId="28" pageNumber="583" type="generic" typeStatus="types">two types</specimenCount>
: (1) upper group, variable in number (three to 19 observed), long, distally very wide, flattened, serrated along both margins, with pointed tips (
<figureCitation id="1B892A02FFE60634FBE1EC80790E5AF7" box="[961,1036,1178,1200]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7I</figureCitation>
); and (2) middle and lower group, numerous (12–35 observed), long to short, distally wide, flattened, serrated along both margins, with pointed tips (
<figureCitation id="1B892A02FFE60634FBE8ECEC791A5B53" box="[968,1048,1270,1292]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7J</figureCitation>
), with a kind of central rib distally present on some specimens (
<figureCitation id="1B892A02FFE60634FC8CED0E79F85B72" box="[1196,1274,1300,1322]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7J</figureCitation>
).
</paragraph>
<paragraph id="830D3687FFE60634FB61ED2978905BDE" blockId="28.[809,1426,197,1414]" pageId="28" pageNumber="583">
Nephridial papillae absent. Pygidium ventrally bilobed, lobes subconical; enclosed by last segment; with dorsal anus (
<figureCitation id="1B892A02FFE60634FB9CED6A792E5BDE" box="[956,1068,1392,1414]" captionStart="Figure 7" captionStartId="29.[163,241,1682,1704]" captionTargetBox="[163,1443,158,1640]" captionTargetId="figure-1212@29.[711,895,747,932]" captionTargetPageId="29" captionText="Figure 7. Bathyeliasona mariaae sp. nov., holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ." pageId="28" pageNumber="583">Fig. 7A, E</figureCitation>
). Anal cirri lost, scars not seen.
</paragraph>
<paragraph id="830D3687FFE60634FB09EDB579E55881" blockId="28.[809,1425,1455,1753]" pageId="28" pageNumber="583">
<emphasis id="B1C6EA95FFE60634FB09EDB579445B9D" box="[809,1094,1455,1477]" italics="true" pageId="28" pageNumber="583">Morphological variation:</emphasis>
The specimens found in the CCFZ are morphologically very similar, having 15 and 17 segments. The common morphological characters included: shape of prostomium and form of prostomial appendages, shape of parapodia and
<typeStatus id="5C098825FFE60634FCF1EE30780C5867" box="[1233,1294,1578,1599]" pageId="28" pageNumber="583">types</typeStatus>
of chaetae. However, the specimen with 15 segments showed some differences probably related to growth: seven pairs of elytrophores, median notch less prominent, longer ventral cirri, longer palps, pygidium not clearly bilobed and not enclosed by the last parapodia.
</paragraph>
<paragraph id="830D3687FFE60636FB09EF187C195BFF" blockId="28.[809,1425,1794,1877]" lastBlockId="30.[145,762,197,1447]" lastPageId="30" lastPageNumber="585" pageId="28" pageNumber="583">
<emphasis id="B1C6EA95FFE60634FB09EF187EA8594F" box="[809,938,1794,1815]" italics="true" pageId="28" pageNumber="583">R e m a r k s:</emphasis>
O n l y t h r e e s p e c i e s b e l o n g i n g t o
<taxonomicName id="44B24D04FFE60634FB09EF3A7ECC596D" authorityName="Pettibone" authorityYear="1976" box="[809,974,1824,1845]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="28" pageNumber="583" phylum="Annelida" rank="genus">
<emphasis id="B1C6EA95FFE60634FB09EF3A7ECC596D" box="[809,974,1824,1845]" italics="true" pageId="28" pageNumber="583">Bathyeliasona</emphasis>
</taxonomicName>
are known:
<taxonomicName id="44B24D04FFE60634FC7DEF3A7EF1590D" authority="(Uschakov, 1971)" baseAuthorityName="Uschakov" baseAuthorityYear="1971" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="28" pageNumber="583" phylum="Annelida" rank="species" species="kirkegaardi">
<emphasis id="B1C6EA95FFE60634FC7DEF3A7893596D" box="[1117,1425,1824,1845]" italics="true" pageId="28" pageNumber="583">Bathyeliasona kirkegaardi</emphasis>
(
<bibRefCitation id="E7234B76FFE60634FB10EF257EE9590D" author="Uschakov PV" box="[816,1003,1855,1877]" pageId="28" pageNumber="583" pagination="36 - 40" refId="ref57956" refString="Uschakov PV. 1971. On a new abyssal species of Macellicephala McIntosh (Polychaeta, Errantia) from the Aleutian Trench. Trudy Instituta Okeanologii Akademiia Nauk SSSR 92: 36 - 40." type="journal article" year="1971">Uschakov, 1971</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="44B24D04FFE60636FC20EF257DD25E83" authority="(Fauvel, 1913)" baseAuthorityName="Fauvel" baseAuthorityYear="1913" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" lastPageId="30" lastPageNumber="585" order="Phyllodocida" pageId="28" pageNumber="583" phylum="Annelida" rank="species" species="abyssicola">
<emphasis id="B1C6EA95FFE60634FC20EF25782B590C" box="[1024,1321,1855,1876]" italics="true" pageId="28" pageNumber="583">Bathyeliasona abyssicola</emphasis>
(
<bibRefCitation id="E7234B76FFE60636FD1AEF257DCB5E83" author="Fauvel P" lastPageId="30" lastPageNumber="585" pageId="28" pageNumber="583" pagination="1 - 80" refId="ref52934" refString="Fauvel P. 1913. Quatrieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la Princesse- Alice, ou deposees dans le Musee Oceanographique de Monaco. Bulletin de l'Institut Oceanographique de Monaco 270: 1 - 80." type="journal article" year="1913">Fauvel, 1913</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="44B24D04FFE40636F92AE8DF7FA95E83" authority="Hartman, 1967" authorityName="Hartman" authorityYear="1967" box="[266,683,197,219]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="nigra">
<emphasis id="B1C6EA95FFE40636F92AE8DF7CF05E83" box="[266,498,197,219]" italics="true" pageId="30" pageNumber="585">Bathyeliasona nigra</emphasis>
<bibRefCitation id="E7234B76FFE40636F9DAE8DC7FA95E83" author="Hartman O" box="[506,683,197,219]" pageId="30" pageNumber="585" pagination="1 - 387" refId="ref53543" refString="Hartman O. 1967. Polychaetous annelids collected by the USNS Eltanin and Staten Island cruises, chiefly from Antarctic Seas. Allan Hancock Monographs in Marine Biology 2: 1 - 387." type="journal article" year="1967">Hartman, 1967</bibRefCitation>
</taxonomicName>
. These species were reviewed by
<bibRefCitation id="E7234B76FFE40636F9C2E8FF7FB65EA3" author="Pettibone MH" box="[482,692,229,251]" pageId="30" pageNumber="585" refId="ref56082" refString="Pettibone MH. 1976. Revision of the genus Macellicephala McIntosh and the subfamily Macellicephalinae Hartmann- Schroder (Polychaeta: Polynoidae). Washington: Smithsonian Institution Press." type="book" year="1976">Pettibone (1976)</bibRefCitation>
, who separated them using characters such as the number of segments,
<typeStatus id="5C098825FFE40636F909E93D7C595F64" box="[297,347,295,316]" pageId="30" pageNumber="585">type</typeStatus>
of notochaetae and development of notopodia/neuropodia in the last segments. Although not mentioned by
<bibRefCitation id="E7234B76FFE40636F943E97D7F235F25" author="Pettibone MH" box="[355,545,359,381]" pageId="30" pageNumber="585" refId="ref56082" refString="Pettibone MH. 1976. Revision of the genus Macellicephala McIntosh and the subfamily Macellicephalinae Hartmann- Schroder (Polychaeta: Polynoidae). Washington: Smithsonian Institution Press." type="book" year="1976">Pettibone (1976)</bibRefCitation>
, differences in the shape of pygidium can be highlighted from her drawings, which can also be used to separate species. The species
<taxonomicName id="44B24D04FFE40636F8B1E9D27CC35F85" baseAuthorityName="Fauvel" baseAuthorityYear="1913" box="[145,449,456,477]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="abyssicola">
<emphasis id="B1C6EA95FFE40636F8B1E9D27CC35F85" box="[145,449,456,477]" italics="true" pageId="30" pageNumber="585">Bathyeliasona abyssicola</emphasis>
</taxonomicName>
and
<taxonomicName id="44B24D04FFE40636FA22E9D27FFB5F86" authorityName="Hartman" authorityYear="1967" box="[514,761,456,478]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="nigra">
<emphasis id="B1C6EA95FFE40636FA22E9D27FFB5F86" box="[514,761,456,478]" italics="true" pageId="30" pageNumber="585">Bathyeliasona nigra</emphasis>
</taxonomicName>
have 18 segments (including tentacular segment), whereas
<taxonomicName id="44B24D04FFE40636F8D9EA137F2E5C46" baseAuthorityName="Uschakov" baseAuthorityYear="1971" box="[249,556,521,542]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="kirkegaardi">
<emphasis id="B1C6EA95FFE40636F8D9EA137F2E5C46" box="[249,556,521,542]" italics="true" pageId="30" pageNumber="585">Bathyeliasona kirkegaardi</emphasis>
</taxonomicName>
has 17 segments, like
<taxonomicName id="44B24D04FFE40636F8EDEA337CFB5C67" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[205,505,553,575]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE40636F8EDEA337CFB5C67" box="[205,505,553,575]" italics="true" pageId="30" pageNumber="585">Bathyeliasona mariaae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFE40636FA26EA337F645C66" box="[518,614,553,574]" pageId="30" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
Additional similarities between the latter two species are: shape of jaws, shape of pygidium and notopodial lobes shorter than neuropodia on the last segments. However, while
<taxonomicName id="44B24D04FFE40636F8B1EAB07CA35C98" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[145,417,682,704]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE40636F8B1EAB07CA35C98" box="[145,417,682,704]" italics="true" pageId="30" pageNumber="585">Bathyeliasona mariaae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFE40636F989EAB17CFD5C98" box="[425,511,683,704]" pageId="30" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
has prostomial lobes anteriorly extending into long frontal filaments that reach the distal end of the ceratophore of the median antenna, in
<taxonomicName id="44B24D04FFE40636F93BEB117F485D78" baseAuthorityName="Uschakov" baseAuthorityYear="1971" box="[283,586,779,800]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="kirkegaardi">
<emphasis id="B1C6EA95FFE40636F93BEB117F485D78" box="[283,586,779,800]" italics="true" pageId="30" pageNumber="585">Bathyeliasona kirkegaardi</emphasis>
</taxonomicName>
the prostomial lobes are anteriorly rounded and the frontal filaments are minute and filiform, shorter than the ceratophore of the median antenna. Furthermore, in
<taxonomicName id="44B24D04FFE40636F8B1EB977CC45DFA" baseAuthorityName="Uschakov" baseAuthorityYear="1971" box="[145,454,909,930]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="kirkegaardi">
<emphasis id="B1C6EA95FFE40636F8B1EB977CC45DFA" box="[145,454,909,930]" italics="true" pageId="30" pageNumber="585">Bathyeliasona kirkegaardi</emphasis>
</taxonomicName>
,
<taxonomicName id="44B24D04FFE40636F9F4EB977FFB5DFA" baseAuthorityName="Fauvel" baseAuthorityYear="1913" box="[468,761,909,930]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="abyssicola">
<emphasis id="B1C6EA95FFE40636F9F4EB977FFB5DFA" box="[468,761,909,930]" italics="true" pageId="30" pageNumber="585">Bathyeliasona abyssicola</emphasis>
</taxonomicName>
and
<taxonomicName id="44B24D04FFE40636F8E2EBB77CAA5D9B" authorityName="Hartman" authorityYear="1967" box="[194,424,941,963]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="nigra">
<emphasis id="B1C6EA95FFE40636F8E2EBB77CAA5D9B" box="[194,424,941,963]" italics="true" pageId="30" pageNumber="585">Bathyeliasona nigra</emphasis>
</taxonomicName>
the length of the ventral cirri is much shorter than the neuroacicular tip and the notochaetae are of
<specimenCount id="95B4FD0EFFE40636F951EBF47CD65A5B" box="[369,468,1006,1027]" pageId="30" pageNumber="585" type="generic" typeStatus="type">one type</specimenCount>
only (stout with distinct spinous rows), whereas in
<taxonomicName id="44B24D04FFE40636F9E5EC147FD05A7C" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[453,722,1038,1060]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE40636F9E5EC147FD05A7C" box="[453,722,1038,1060]" italics="true" pageId="30" pageNumber="585">Bathyeliasona mariaae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFE40636FAF9EC147DBC5A1C" pageId="30" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
the length of the ventral cirri is approaching the neuroacicula tip and the notochaetae are of
<specimenCount id="95B4FD0EFFE40636FAABEC557FFB5A3C" box="[651,761,1103,1124]" pageId="30" pageNumber="585" type="generic" typeStatus="types">two types</specimenCount>
(stouter with distinct spinous rows and slender with developed spinous rows).
<bibRefCitation id="E7234B76FFE40636F99FEC957F835AFD" author="Pettibone MH" box="[447,641,1167,1189]" pageId="30" pageNumber="585" refId="ref56082" refString="Pettibone MH. 1976. Revision of the genus Macellicephala McIntosh and the subfamily Macellicephalinae Hartmann- Schroder (Polychaeta: Polynoidae). Washington: Smithsonian Institution Press." type="book" year="1976">Pettibone (1976)</bibRefCitation>
described all the
<taxonomicName id="44B24D04FFE40636F8C5ECAA7CE75A9E" box="[229,485,1200,1222]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="B1C6EA95FFE40636F8C5ECAA7C895A9D" box="[229,395,1200,1221]" italics="true" pageId="30" pageNumber="585">Bathyeliasona</emphasis>
species
</taxonomicName>
with notochaetae more slender than neurochaetae, but in
<taxonomicName id="44B24D04FFE40636FA69ECCA7DF25B5E" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE40636FA69ECCA7DF25B5E" italics="true" pageId="30" pageNumber="585">Bathyeliasona mariaae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFE40636F8D5ECEB7C445B5E" box="[245,326,1265,1286]" pageId="30" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
one notochaeta per notopodium can be stouter (and always narrower) than the neurochaetae. Furthermore, in
<taxonomicName id="44B24D04FFE40636F979ED2B7F6E5B1F" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[345,620,1329,1351]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="30" pageNumber="585" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE40636F979ED2B7F6E5B1F" box="[345,620,1329,1351]" italics="true" pageId="30" pageNumber="585">Bathyeliasona mariaae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AAF557EEFFE40636FA54ED2B7FC95B1E" box="[628,715,1329,1350]" pageId="30" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
the middle and lower neurochaetae show a kind of central rib distally that gives the neurochaetae a bilimbate appearance.
</paragraph>
<caption id="D7CD660FFFE70635F883EE887CEF5963" pageId="29" pageNumber="584" startId="29.[163,241,1682,1704]" targetBox="[163,1443,158,1640]" targetPageId="29">
<paragraph id="830D3687FFE70635F883EE887CEF5963" blockId="29.[163,1443,1682,1851]" pageId="29" pageNumber="584">
<emphasis id="B1C6EA95FFE70635F883EE887F2558FF" bold="true" box="[163,551,1682,1704]" pageId="29" pageNumber="584">
Figure 7.
<taxonomicName id="44B24D04FFE70635F936EE887F2558FF" authority="Bonifácio &amp; Menot, 2019" authorityName="Bonifácio &amp; Menot" authorityYear="2019" box="[278,551,1682,1704]" class="Polychaeta" family="Polynoidae" genus="Bathyeliasona" kingdom="Animalia" order="Phyllodocida" pageId="29" pageNumber="584" phylum="Annelida" rank="species" species="mariaae" status="sp. nov.">
<emphasis id="B1C6EA95FFE70635F936EE887F2558FF" bold="true" box="[278,551,1682,1704]" italics="true" pageId="29" pageNumber="584">Bathyeliasona mariaae</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="AAF557EEFFE70635FA0EEE897F8758F1" box="[558,645,1683,1705]" pageId="29" pageNumber="584" rank="species">
<emphasis id="B1C6EA95FFE70635FA0EEE897F8258F0" bold="true" box="[558,640,1683,1704]" pageId="29" pageNumber="584">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype MNHN-IA-TYPE 1815 (A–C, E–J) and paratype MNHN-IA-TYPE 1816 (D). A, dorsal view of a live complete specimen. B, anterior end, dorsal view of a live specimen. C, anterior end, dorsal view, chaetae omitted. D, inner view of half side of dissected pharynx with papillae. E, posterior end, dorsal view, chaetae omitted. F, left parapodia, posterior view (ventral cirri lost), segment 8. G, notochaeta with distinct spinous rows, segment 8. H, notochaeta with developed spinous, segment 8. I, upper neurochaeta, segment 8. J, lower neurochaeta, segment 8. Abbreviation: nu, nuchal organ.
</paragraph>
</caption>
<paragraph id="830D3687FFE40636F8B1EDC87CC45851" blockId="30.[145,760,1490,1545]" pageId="30" pageNumber="585">
<emphasis id="B1C6EA95FFE40636F8B1EDC87C105BBF" box="[145,274,1490,1511]" italics="true" pageId="30" pageNumber="585">Etymology:</emphasis>
This species is dedicated to Maria Silva, mother of P.B., for her love.
</paragraph>
<paragraph id="830D3687FFE40636F8B1EE297F6B5892" blockId="30.[145,761,1587,1738]" pageId="30" pageNumber="585">
<emphasis id="B1C6EA95FFE40636F8B1EE297C3F5810" box="[145,317,1587,1608]" italics="true" pageId="30" pageNumber="585">Genetic data:</emphasis>
DNA sequencing for this species was successful for
<emphasis id="B1C6EA95FFE40636F951EE4E7CA35831" box="[369,417,1620,1641]" italics="true" pageId="30" pageNumber="585">COI</emphasis>
, 16S and 18S, with all three genes sharing 100% of genetic material between the specimens. The average K2P distance for intraspecific variation was 0.0% for both
<emphasis id="B1C6EA95FFE40636F9F0EEAF7CFD5892" box="[464,511,1717,1738]" italics="true" pageId="30" pageNumber="585">COI</emphasis>
and 16S.
</paragraph>
<paragraph id="830D3687FFE40636F8B1EEEF7FBB5933" blockId="30.[145,760,1781,1900]" pageId="30" pageNumber="585">
<emphasis id="B1C6EA95FFE40636F8B1EEEF7C285952" box="[145,298,1781,1802]" italics="true" pageId="30" pageNumber="585">Distribution:</emphasis>
Based on the material examined (
<specimenCount id="95B4FD0EFFE40636FAEDEEEF7C0B5973" pageId="30" pageNumber="585" type="generic">two specimens</specimenCount>
), this species has a wide distribution within the Clarion-Clipperton Fracture Zone, being sampled in BGR license (
<typeStatus id="5C098825FFE40636F968EF4C7C785933" box="[328,378,1878,1899]" pageId="30" pageNumber="585">type</typeStatus>
locality) and APEI#3 areas.
</paragraph>
</subSubSection>
</treatment>
</document>
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